The Securities and Exchange Commission has not necessarily reviewed the information in this filing and has not determined if it is accurate and complete.
The reader should not assume that the information is accurate and complete.

UNITED STATES SECURITIES AND EXCHANGE COMMISSION
Washington, D.C. 20549
FORM 13F

FORM 13F INFORMATION TABLE
OMB APPROVAL
OMB Number: 3235-0006
Estimated average burden
hours per response: 23.8
COLUMN 1 COLUMN 2 COLUMN 3 COLUMN 4 COLUMN 5 COLUMN 6 COLUMN 7 COLUMN 8
VALUE SHRS OR SH/ PUT/ INVESTMENT OTHER VOTING AUTHORITY
NAME OF ISSUER TITLE OF CLASS CUSIP (x$1000) PRN AMT PRN CALL DISCRETION MANAGER SOLE SHARED NONE
10X CAPITAL VENTURE ACQU COR COM CL A 88025V107 7,028 700,000 SH   DFND 5 700,000 0 0
10X GENOMICS INC CL A COM 88025U109 7,460 37,008 SH   DFND 3 37,008 0 0
10X GENOMICS INC CL A COM 88025U109 57 282 SH   DFND 5 282 0 0
10X GENOMICS INC CL A COM 88025U109 103 509 SH   DFND 6 0 0 509
10X GENOMICS INC CL A COM 88025U109 8 40 SH   DFND   40 0 0
1LIFE HEALTHCARE INC COM 68269G107 16,241 472,404 SH   DFND 5 472,404 0 0
1LIFE HEALTHCARE INC COM 68269G107 0 12 SH   DFND 6 0 0 12
1LIFE HEALTHCARE INC COM 68269G107 33 960 SH   DFND   960 0 0
1ST SOURCE CORP COM 336901103 5,225 112,584 SH   DFND 2 110,979 0 1,605
1ST SOURCE CORP COM 336901103 305 6,566 SH   DFND 4 6,566 0 0
1ST SOURCE CORP COM 336901103 0 1 SH   DFND 5 1 0 0
1ST SOURCE CORP COM 336901103 6 133 SH   DFND 6 0 0 133
1ST SOURCE CORP COM 336901103 1,141 24,594 SH   DFND 7 0 0 24,594
22ND CENTY GROUP INC COM 90137F103 0 5 SH   DFND 5 5 0 0
22ND CENTY GROUP INC COM 90137F103 83 17,000 SH   DFND   17,000 0 0
2U INC COM 90214J101 1,664 39,162 SH   DFND 2 39,162 0 0
2U INC COM 90214J101 11 250 SH   DFND 3 250 0 0
2U INC COM 90214J101 0 2 SH   DFND 5 2 0 0
2U INC COM 90214J101 146 3,432 SH   DFND 6 0 0 3,432
2U INC COM 90214J101 43 1,001 SH   DFND 7 1,001 0 0
2U INC COM 90214J101 103 2,421 SH   DFND   2,421 0 0
3-D SYS CORP DEL COM NEW 88554D205 86 2,198 SH   DFND 2 2,198 0 0
3-D SYS CORP DEL COM NEW 88554D205 1,507 38,487 SH   DFND 3 38,487 0 0
3-D SYS CORP DEL COM NEW 88554D205 2,353 60,104 SH   DFND 5 60,104 0 0
3-D SYS CORP DEL COM NEW 88554D205 26 660 SH   DFND 6 0 0 660
3-D SYS CORP DEL COM NEW 88554D205 2 40 SH   DFND 11 40 0 0
3-D SYS CORP DEL COM NEW 88554D205 68 1,749 SH   DFND   1,749 0 0
3M CO COM 88579Y101 29,919 152,491 SH   DFND 2 152,491 0 0
3M CO COM 88579Y101 68,881 351,073 SH   DFND 3 351,073 0 0
3M CO COM 88579Y101 20,609 105,041 SH   DFND 4 22,248 0 82,793
3M CO COM 88579Y101 40,789 207,896 SH   DFND 5 207,896 0 0
3M CO COM 88579Y101 2,957 15,073 SH   DFND 6 0 0 15,073
3M CO COM 88579Y101 65,071 331,655 SH   DFND 7 323,971 0 7,684
3M CO COM 88579Y101 3,108 15,842 SH   DFND 9 15,842 0 0
3M CO COM 88579Y101 391 1,995 SH   DFND 11 1,995 0 0
3M CO COM 88579Y101 29,048 148,055 SH   DFND   147,555 0 500
890 5TH AVE PARTNERS INC COM CL A 28250A105 1,491 149,999 SH   DFND 5 149,999 0 0
8X8 INC NEW COM 282914100 107 3,794 SH   DFND 2 3,794 0 0
8X8 INC NEW COM 282914100 948 33,465 SH   DFND 3 33,465 0 0
8X8 INC NEW COM 282914100 47,408 1,674,003 SH   DFND 5 1,674,003 0 0
8X8 INC NEW COM 282914100 26,904 950,000 SH Put DFND 5 950,000 0 0
8X8 INC NEW COM 282914100 4 124 SH   DFND 6 0 0 124
8X8 INC NEW NOTE 0.500% 2/0 282914AB6 2,024 1,600,000 PRN   DFND 8 1,600,000 0 0
9 METERS BIOPHARMA INC COM 654405109 0 7 SH   DFND 5 7 0 0
9 METERS BIOPHARMA INC COM 654405109 13 11,646 SH   DFND   11,646 0 0
A10 NETWORKS INC COM 002121101 1,945 169,717 SH   DFND 2 169,717 0 0
A10 NETWORKS INC COM 002121101 0 2 SH   DFND 5 2 0 0
AAON INC COM PAR $0.004 000360206 143 2,244 SH   DFND 2 2,244 0 0
AAON INC COM PAR $0.004 000360206 796 12,465 SH   DFND 3 12,465 0 0
AAON INC COM PAR $0.004 000360206 0 1 SH   DFND 5 1 0 0
AAON INC COM PAR $0.004 000360206 121 1,899 SH   DFND 7 1,899 0 0
AAR CORP COM 000361105 29 755 SH   DFND 2 755 0 0
AAR CORP COM 000361105 396 10,248 SH   DFND 3 10,248 0 0
AAR CORP COM 000361105 515 13,344 SH   DFND 5 13,344 0 0
AAR CORP COM 000361105 282 7,310 SH   DFND 6 0 0 7,310
ABB LTD SPONSORED ADR 000375204 93 2,680 SH   DFND 3 2,680 0 0
ABB LTD SPONSORED ADR 000375204 261 7,551 SH   DFND 6 0 0 7,551
ABB LTD SPONSORED ADR 000375204 24 696 SH   DFND 7 0 0 696
ABB LTD SPONSORED ADR 000375204 29,601 855,766 SH   DFND   855,766 0 0
ABBOTT LABS COM 002824100 6,409 59,946 SH   DFND 2 59,946 0 0
ABBOTT LABS COM 002824100 59,699 558,395 SH   DFND 3 558,395 0 0
ABBOTT LABS COM 002824100 22,642 211,777 SH   DFND 4 211,777 0 0
ABBOTT LABS COM 002824100 94,102 880,176 SH   DFND 5 880,176 0 0
ABBOTT LABS COM 002824100 4,212 39,401 SH   DFND 6 0 0 39,401
ABBOTT LABS COM 002824100 70,016 654,891 SH   DFND 7 578,074 54,971 21,846
ABBOTT LABS COM 002824100 2,763 25,847 SH   DFND 9 25,847 0 0
ABBOTT LABS COM 002824100 342 3,202 SH   DFND 11 3,202 0 0
ABBOTT LABS COM 002824100 97,534 912,277 SH   DFND   912,277 0 0
ABBVIE INC COM 00287Y109 23,126 205,362 SH   DFND 2 205,362 0 0
ABBVIE INC COM 00287Y109 99,518 883,742 SH   DFND 3 883,742 0 0
ABBVIE INC COM 00287Y109 13,768 122,262 SH   DFND 4 122,262 0 0
ABBVIE INC COM 00287Y109 82,838 735,615 SH   DFND 5 735,615 0 0
ABBVIE INC COM 00287Y109 5,067 45,000 SH Call DFND 5 45,000 0 0
ABBVIE INC COM 00287Y109 7,285 64,696 SH   DFND 6 0 0 64,696
ABBVIE INC COM 00287Y109 58,097 515,911 SH   DFND 7 487,003 5,725 23,183
ABBVIE INC COM 00287Y109 44,400 394,281 SH   DFND 8 394,281 0 0
ABBVIE INC COM 00287Y109 1,424 12,645 SH   DFND 9 12,645 0 0
ABBVIE INC COM 00287Y109 658 5,844 SH   DFND 11 5,844 0 0
ABBVIE INC COM 00287Y109 84,483 750,223 SH   DFND   750,223 0 0
ABCAM PLC ADS 000380204 188 7,155 SH   DFND 6 0 0 7,155
ABCAM PLC ADS 000380204 3,652 138,660 SH   DFND   138,660 0 0
ABCELLERA BIOLOGICS INC COM 00288U106 1,545 72,490 SH   DFND   72,490 0 0
ABERCROMBIE & FITCH CO CL A 002896207 4,497 98,126 SH   DFND 2 98,126 0 0
ABERCROMBIE & FITCH CO CL A 002896207 862 18,807 SH   DFND 3 18,807 0 0
ABERCROMBIE & FITCH CO CL A 002896207 0 2 SH   DFND 5 2 0 0
ABERCROMBIE & FITCH CO CL A 002896207 4 88 SH   DFND 6 0 0 88
ABERCROMBIE & FITCH CO CL A 002896207 952 20,772 SH   DFND 7 0 0 20,772
ABERDEEN STD PLATINUM ETF TR PHYSCL PLATM SHS 003260106 719 7,170 SH   DFND 1 7,170 0 0
ABERDEEN STD PLATINUM ETF TR PHYSCL PLATM SHS 003260106 121 1,210 SH   DFND   1,210 0 0
ABIOMED INC COM 003654100 171 544 SH   DFND 2 544 0 0
ABIOMED INC COM 003654100 11,213 35,749 SH   DFND 3 35,749 0 0
ABIOMED INC COM 003654100 7,102 22,644 SH   DFND 5 22,644 0 0
ABIOMED INC COM 003654100 283 903 SH   DFND 6 0 0 903
ABIOMED INC COM 003654100 10,445 33,300 SH   DFND 8 33,300 0 0
ABIOMED INC COM 003654100 69 220 SH   DFND 9 220 0 0
ABIOMED INC COM 003654100 11 35 SH   DFND   35 0 0
ABM INDS INC COM 000957100 0 1 SH   DFND 1 1 0 0
ABM INDS INC COM 000957100 45 990 SH   DFND 2 990 0 0
ABM INDS INC COM 000957100 920 20,464 SH   DFND 3 20,464 0 0
ABM INDS INC COM 000957100 0 2 SH   DFND 5 2 0 0
ABM INDS INC COM 000957100 158 3,517 SH   DFND 6 0 0 3,517
ABSOLUTE SOFTWARE CORP COM 00386B109 70 4,877 SH   DFND 8 4,877 0 0
ABSOLUTE SOFTWARE CORP COM 00386B109 2,269 158,151 SH   DFND   158,151 0 0
ACADIA HEALTHCARE COMPANY IN COM 00404A109 0 1 SH   DFND 1 1 0 0
ACADIA HEALTHCARE COMPANY IN COM 00404A109 6,288 97,369 SH   DFND 2 97,369 0 0
ACADIA HEALTHCARE COMPANY IN COM 00404A109 334 5,172 SH   DFND 3 5,172 0 0
ACADIA HEALTHCARE COMPANY IN COM 00404A109 5,444 84,306 SH   DFND 4 0 0 84,306
ACADIA HEALTHCARE COMPANY IN COM 00404A109 65 1,007 SH   DFND 6 0 0 1,007
ACADIA HEALTHCARE COMPANY IN COM 00404A109 64 990 SH   DFND 7 990 0 0
ACADIA HEALTHCARE COMPANY IN COM 00404A109 310 4,800 SH   DFND 8 4,800 0 0
ACADIA PHARMACEUTICALS INC COM 004225108 1,337 52,834 SH   DFND 2 47,587 0 5,247
ACADIA PHARMACEUTICALS INC COM 004225108 535 21,138 SH   DFND 4 21,138 0 0
ACADIA PHARMACEUTICALS INC COM 004225108 0 4 SH   DFND 5 4 0 0
ACADIA PHARMACEUTICALS INC COM 004225108 158 6,262 SH   DFND 6 0 0 6,262
ACADIA PHARMACEUTICALS INC COM 004225108 38 1,509 SH   DFND 7 1,509 0 0
ACADIA PHARMACEUTICALS INC COM 004225108 5 200 SH   DFND   200 0 0
ACADIA RLTY TR COM SH BEN INT 004239109 9,043 407,180 SH   DFND 2 407,180 0 0
ACADIA RLTY TR COM SH BEN INT 004239109 591 26,593 SH   DFND 3 26,593 0 0
ACADIA RLTY TR COM SH BEN INT 004239109 0 3 SH   DFND 5 3 0 0
ACADIA RLTY TR COM SH BEN INT 004239109 6 280 SH   DFND 6 0 0 280
ACADIA RLTY TR COM SH BEN INT 004239109 1,397 62,915 SH   DFND 7 406 0 62,509
ACASTI PHARMA INC CL A NEW 00430K402 6 10,900 SH   DFND   10,900 0 0
ACCELERON PHARMA INC COM 00434H108 144 1,133 SH   DFND 2 339 0 794
ACCELERON PHARMA INC COM 00434H108 408 3,209 SH   DFND 4 3,209 0 0
ACCELERON PHARMA INC COM 00434H108 161 1,271 SH   DFND 5 1,271 0 0
ACCELERON PHARMA INC COM 00434H108 139 1,096 SH   DFND 6 0 0 1,096
ACCELERON PHARMA INC COM 00434H108 24,274 191,100 SH   DFND 8 191,100 0 0
ACCENTURE PLC IRELAND SHS CLASS A G1151C101 5,978 20,116 SH   DFND 2 20,116 0 0
ACCENTURE PLC IRELAND SHS CLASS A G1151C101 80,976 272,481 SH   DFND 3 272,481 0 0
ACCENTURE PLC IRELAND SHS CLASS A G1151C101 171,524 577,172 SH   DFND 4 577,172 0 0
ACCENTURE PLC IRELAND SHS CLASS A G1151C101 86,143 289,868 SH   DFND 5 289,868 0 0
ACCENTURE PLC IRELAND SHS CLASS A G1151C101 5,008 16,852 SH   DFND 6 0 0 16,852
ACCENTURE PLC IRELAND SHS CLASS A G1151C101 39,962 134,471 SH   DFND 7 70,233 57,475 6,763
ACCENTURE PLC IRELAND SHS CLASS A G1151C101 892 3,000 SH   DFND 8 3,000 0 0
ACCENTURE PLC IRELAND SHS CLASS A G1151C101 9,183 30,900 SH   DFND 9 30,900 0 0
ACCENTURE PLC IRELAND SHS CLASS A G1151C101 359 1,209 SH   DFND 11 1,209 0 0
ACCENTURE PLC IRELAND SHS CLASS A G1151C101 230,060 774,143 SH   DFND   773,578 0 565
ACI WORLDWIDE INC COM 004498101 98 2,604 SH   DFND 2 2,604 0 0
ACI WORLDWIDE INC COM 004498101 276 7,314 SH   DFND 3 7,314 0 0
ACI WORLDWIDE INC COM 004498101 26 701 SH   DFND 6 0 0 701
ACLARIS THERAPEUTICS INC COM 00461U105 1,573 91,479 SH   DFND 2 91,479 0 0
ACLARIS THERAPEUTICS INC COM 00461U105 0 1 SH   DFND 5 1 0 0
ACON S2 ACQUISITION CORP CL SHS G00748106 11,987 1,202,337 SH   DFND 5 1,202,337 0 0
ACON S2 ACQUISITION CORP CL SHS G00748106 4,776 479,000 SH   DFND 8 479,000 0 0
ACTIVISION BLIZZARD INC COM 00507V109 3,531 36,925 SH   DFND 2 36,925 0 0
ACTIVISION BLIZZARD INC COM 00507V109 32,918 344,255 SH   DFND 3 344,255 0 0
ACTIVISION BLIZZARD INC COM 00507V109 2,819 29,485 SH   DFND 4 29,485 0 0
ACTIVISION BLIZZARD INC COM 00507V109 27,071 283,107 SH   DFND 5 283,107 0 0
ACTIVISION BLIZZARD INC COM 00507V109 1,630 17,049 SH   DFND 6 0 0 17,049
ACTIVISION BLIZZARD INC COM 00507V109 33,388 349,178 SH   DFND 7 336,609 0 12,569
ACTIVISION BLIZZARD INC COM 00507V109 3,366 35,200 SH   DFND 8 35,200 0 0
ACTIVISION BLIZZARD INC COM 00507V109 6,291 65,796 SH   DFND 9 65,796 0 0
ACTIVISION BLIZZARD INC COM 00507V109 43 447 SH   DFND 11 447 0 0
ACTIVISION BLIZZARD INC COM 00507V109 70,362 735,852 SH   DFND   735,012 0 840
ACUITY BRANDS INC COM 00508Y102 1,877 9,826 SH   DFND 2 9,826 0 0
ACUITY BRANDS INC COM 00508Y102 398 2,083 SH   DFND 3 2,083 0 0
ACUITY BRANDS INC COM 00508Y102 194,120 1,016,068 SH   DFND 4 1,016,068 0 0
ACUITY BRANDS INC COM 00508Y102 392 2,053 SH   DFND 6 0 0 2,053
ACUITY BRANDS INC COM 00508Y102 1,848 9,675 SH   DFND 7 8,600 0 1,075
ACUITY BRANDS INC COM 00508Y102 267 1,400 SH   DFND 8 1,400 0 0
ACUITYADS HLDGS INC COM 00510L106 8 764 SH   DFND 9 764 0 0
ACUITYADS HLDGS INC COM 00510L106 334 32,800 SH   DFND   32,800 0 0
ACUTUS MED INC COM 005111109 0 1 SH   DFND 5 1 0 0
ACUTUS MED INC COM 005111109 5 310 SH   DFND 6 0 0 310
ACUTUS MED INC COM 005111109 8,498 483,100 SH   DFND 8 483,100 0 0
ADAMAS PHARMACEUTICALS INC COM 00548A106 1,336 242,390 SH   DFND 2 242,390 0 0
ADAMS DIVERSIFIED EQUITY FD COM 006212104 1,594 69,712 SH   DFND 5 69,712 0 0
ADDUS HOMECARE CORP COM 006739106 39 439 SH   DFND 2 439 0 0
ADDUS HOMECARE CORP COM 006739106 403 4,588 SH   DFND 3 4,588 0 0
ADDUS HOMECARE CORP COM 006739106 0 1 SH   DFND 5 1 0 0
ADIENT PLC ORD SHS G0084W101 109 2,465 SH   DFND 2 2,465 0 0
ADIENT PLC ORD SHS G0084W101 241 5,446 SH   DFND 3 5,446 0 0
ADIENT PLC ORD SHS G0084W101 30 682 SH   DFND 6 0 0 682
ADIENT PLC ORD SHS G0084W101 190 4,301 SH   DFND   4,301 0 0
ADIT EDTECH ACQUISITION CORP COMMON STOCK 007024102 2,393 246,747 SH   DFND 5 246,747 0 0
ADOBE SYSTEMS INCORPORATED COM 00724F101 84,603 143,200 SH   DFND 2 143,200 0 0
ADOBE SYSTEMS INCORPORATED COM 00724F101 168,467 285,150 SH   DFND 3 285,150 0 0
ADOBE SYSTEMS INCORPORATED COM 00724F101 55,584 94,083 SH   DFND 4 43,389 0 50,694
ADOBE SYSTEMS INCORPORATED COM 00724F101 145,768 246,730 SH   DFND 5 246,730 0 0
ADOBE SYSTEMS INCORPORATED COM 00724F101 23,632 40,000 SH Put DFND 5 40,000 0 0
ADOBE SYSTEMS INCORPORATED COM 00724F101 5,639 9,545 SH   DFND 6 0 0 9,545
ADOBE SYSTEMS INCORPORATED COM 00724F101 122,248 206,920 SH   DFND 7 201,723 8 5,189
ADOBE SYSTEMS INCORPORATED COM 00724F101 16,779 28,400 SH   DFND 8 28,400 0 0
ADOBE SYSTEMS INCORPORATED COM 00724F101 21,899 37,067 SH   DFND 9 37,067 0 0
ADOBE SYSTEMS INCORPORATED COM 00724F101 1,650 2,792 SH   DFND 11 2,792 0 0
ADOBE SYSTEMS INCORPORATED COM 00724F101 195,881 331,552 SH   DFND   330,267 0 1,285
ADTALEM GLOBAL ED INC COM 00737L103 23 633 SH   DFND 2 633 0 0
ADTALEM GLOBAL ED INC COM 00737L103 102 2,862 SH   DFND 3 2,862 0 0
ADTALEM GLOBAL ED INC COM 00737L103 3,699 103,400 SH   DFND 5 103,400 0 0
ADTALEM GLOBAL ED INC COM 00737L103 14 393 SH   DFND 6 0 0 393
ADTRAN INC COM 00738A106 322 14,911 SH   DFND 3 14,911 0 0
ADTRAN INC COM 00738A106 0 2 SH   DFND 5 2 0 0
ADVANCE AUTO PARTS INC COM 00751Y106 3,824 18,562 SH   DFND 2 18,562 0 0
ADVANCE AUTO PARTS INC COM 00751Y106 4,630 22,479 SH   DFND 3 22,479 0 0
ADVANCE AUTO PARTS INC COM 00751Y106 39,144 190,030 SH   DFND 4 190,030 0 0
ADVANCE AUTO PARTS INC COM 00751Y106 4,969 24,124 SH   DFND 5 24,124 0 0
ADVANCE AUTO PARTS INC COM 00751Y106 1,275 6,188 SH   DFND 6 0 0 6,188
ADVANCE AUTO PARTS INC COM 00751Y106 2,631 12,772 SH   DFND 7 7,450 137 5,185
ADVANCE AUTO PARTS INC COM 00751Y106 21 100 SH   DFND 11 100 0 0
ADVANCE AUTO PARTS INC COM 00751Y106 452 2,192 SH   DFND   2,192 0 0
ADVANCED DRAIN SYS INC DEL COM 00790R104 3 22 SH   DFND 5 22 0 0
ADVANCED DRAIN SYS INC DEL COM 00790R104 116 993 SH   DFND 7 446 0 547
ADVANCED DRAIN SYS INC DEL COM 00790R104 2,771 23,700 SH   DFND 8 23,700 0 0
ADVANCED ENERGY INDS COM 007973100 3,957 34,772 SH   DFND 2 33,830 0 942
ADVANCED ENERGY INDS COM 007973100 1,340 11,774 SH   DFND 3 11,774 0 0
ADVANCED ENERGY INDS COM 007973100 426 3,741 SH   DFND 4 3,741 0 0
ADVANCED ENERGY INDS COM 007973100 0 1 SH   DFND 5 1 0 0
ADVANCED ENERGY INDS COM 007973100 630 5,534 SH   DFND 6 0 0 5,534
ADVANCED ENERGY INDS COM 007973100 181 1,592 SH   DFND 7 1,592 0 0
ADVANCED MICRO DEVICES INC COM 007903107 672 7,493 SH   DFND 1 7,493 0 0
ADVANCED MICRO DEVICES INC COM 007903107 1,042 11,613 SH   DFND 2 11,613 0 0
ADVANCED MICRO DEVICES INC COM 007903107 40,655 453,056 SH   DFND 3 453,056 0 0
ADVANCED MICRO DEVICES INC COM 007903107 11,730 130,716 SH   DFND 4 130,716 0 0
ADVANCED MICRO DEVICES INC COM 007903107 403,801 4,499,924 SH   DFND 5 4,499,924 0 0
ADVANCED MICRO DEVICES INC COM 007903107 338,750 3,775,000 SH Put DFND 5 3,775,000 0 0
ADVANCED MICRO DEVICES INC COM 007903107 47,560 530,000 SH Call DFND 5 530,000 0 0
ADVANCED MICRO DEVICES INC COM 007903107 2,004 22,330 SH   DFND 6 0 0 22,330
ADVANCED MICRO DEVICES INC COM 007903107 8,974 100,000 SH   DFND 8 100,000 0 0
ADVANCED MICRO DEVICES INC COM 007903107 261 2,908 SH   DFND 9 2,908 0 0
ADVANCED MICRO DEVICES INC COM 007903107 204 2,278 SH   DFND 11 2,278 0 0
ADVANCED MICRO DEVICES INC COM 007903107 24,283 270,612 SH   DFND   270,612 0 0
ADVANCED MICRO DEVICES INC COM 007903107 3,589 40,000 SH Call DFND   40,000 0 0
ADVANSIX INC COM 00773T101 6 210 SH   DFND 2 210 0 0
ADVANSIX INC COM 00773T101 255 8,549 SH   DFND 3 8,549 0 0
ADVANSIX INC COM 00773T101 0 1 SH   DFND 5 1 0 0
ADVANSIX INC COM 00773T101 5 179 SH   DFND 6 0 0 179
ADVANSIX INC COM 00773T101 0 5 SH   DFND   5 0 0
ADVISORSHARES TR PURE US CANNABIS 00768Y453 1,591 39,707 SH   DFND   39,707 0 0
ADVISORSHARES TR PURE CANNABIS 00768Y495 18 867 SH   DFND   867 0 0
AECOM COM 00766T100 173 2,717 SH   DFND 2 2,717 0 0
AECOM COM 00766T100 536 8,394 SH   DFND 3 8,394 0 0
AECOM COM 00766T100 31,377 491,492 SH   DFND 4 491,492 0 0
AECOM COM 00766T100 209 3,272 SH   DFND 6 0 0 3,272
AECOM COM 00766T100 2,190 34,300 SH   DFND 8 34,300 0 0
AECOM COM 00766T100 125 1,953 SH   DFND   1,953 0 0
AEGON N V NY REGISTRY SHS 007924103 245 59,523 SH   DFND 6 0 0 59,523
AEGON N V NY REGISTRY SHS 007924103 1 200 SH   DFND 7 0 0 200
AEGON N V NY REGISTRY SHS 007924103 1 133 SH   DFND 11 133 0 0
AEGON N V NY REGISTRY SHS 007924103 111 26,957 SH   DFND   26,957 0 0
AEQUI ACQUISITION CORP COM CL A 00775W102 485 49,526 SH   DFND   49,526 0 0
AERCAP HOLDINGS NV SHS N00985106 14,505 275,511 SH   DFND 2 275,511 0 0
AERCAP HOLDINGS NV SHS N00985106 297 5,632 SH   DFND 6 0 0 5,632
AERIE PHARMACEUTICALS INC COM 00771V108 1,315 72,719 SH   DFND 2 72,719 0 0
AERIE PHARMACEUTICALS INC COM 00771V108 0 1 SH   DFND 5 1 0 0
AERIE PHARMACEUTICALS INC COM 00771V108 43 2,389 SH   DFND 7 2,389 0 0
AEROJET ROCKETDYNE HLDGS INC COM 007800105 90 1,866 SH   DFND 2 1,866 0 0
AEROJET ROCKETDYNE HLDGS INC COM 007800105 1,082 22,462 SH   DFND 3 22,462 0 0
AEROJET ROCKETDYNE HLDGS INC COM 007800105 902 18,732 SH   DFND 5 18,732 0 0
AEROJET ROCKETDYNE HLDGS INC COM 007800105 125 2,602 SH   DFND 6 0 0 2,602
AEROVIRONMENT INC COM 008073108 56 507 SH   DFND 2 507 0 0
AEROVIRONMENT INC COM 008073108 759 6,880 SH   DFND 3 6,880 0 0
AEROVIRONMENT INC COM 008073108 891 8,082 SH   DFND 5 8,082 0 0
AEROVIRONMENT INC COM 008073108 56 507 SH   DFND 7 507 0 0
AES CORP COM 00130H105 80,675 3,055,858 SH   DFND 2 2,826,812 0 229,046
AES CORP COM 00130H105 4,519 171,188 SH   DFND 3 171,188 0 0
AES CORP COM 00130H105 2,439 92,374 SH   DFND 4 92,374 0 0
AES CORP COM 00130H105 4,784 181,219 SH   DFND 5 181,219 0 0
AES CORP COM 00130H105 274 10,371 SH   DFND 6 0 0 10,371
AES CORP COM 00130H105 35,616 1,349,080 SH   DFND 7 1,331,074 3,358 14,648
AES CORP COM 00130H105 2,751 104,200 SH   DFND 8 104,200 0 0
AES CORP COM 00130H105 3,527 133,586 SH   DFND 11 133,586 0 0
AES CORP COM 00130H105 8,264 313,042 SH   DFND   309,812 0 3,230
AETERNA ZENTARIS INC COM 007975402 15 16,936 SH   DFND   16,936 0 0
AFFILIATED MANAGERS GROUP IN COM 008252108 1 6 SH   DFND 1 6 0 0
AFFILIATED MANAGERS GROUP IN COM 008252108 61 392 SH   DFND 2 392 0 0
AFFILIATED MANAGERS GROUP IN COM 008252108 396 2,525 SH   DFND 3 2,525 0 0
AFFILIATED MANAGERS GROUP IN COM 008252108 110 703 SH   DFND 6 0 0 703
AFFILIATED MANAGERS GROUP IN COM 008252108 763 4,867 SH   DFND 7 3,051 0 1,816
AFFILIATED MANAGERS GROUP IN COM 008252108 31 200 SH   DFND 9 200 0 0
AFFILIATED MANAGERS GROUP IN COM 008252108 12 75 SH   DFND   75 0 0
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ARCH CAP GROUP LTD ORD G0450A105 437 11,242 SH   DFND 4 11,242 0 0
ARCH CAP GROUP LTD ORD G0450A105 1,118 28,743 SH   DFND 5 28,743 0 0
ARCH CAP GROUP LTD ORD G0450A105 212 5,447 SH   DFND 6 0 0 5,447
ARCH CAP GROUP LTD ORD G0450A105 6,547 168,265 SH   DFND 9 168,265 0 0
ARCH RESOURCES INC CL A 03940R107 19,359 334,407 SH   DFND 5 334,407 0 0
ARCHER DANIELS MIDLAND CO COM 039483102 117 1,778 SH   DFND 2 1,778 0 0
ARCHER DANIELS MIDLAND CO COM 039483102 10,076 153,501 SH   DFND 3 153,501 0 0
ARCHER DANIELS MIDLAND CO COM 039483102 19,331 294,499 SH   DFND 5 294,499 0 0
ARCHER DANIELS MIDLAND CO COM 039483102 836 12,729 SH   DFND 6 0 0 12,729
ARCHER DANIELS MIDLAND CO COM 039483102 1,003 15,285 SH   DFND 7 10,735 0 4,550
ARCHER DANIELS MIDLAND CO COM 039483102 20 304 SH   DFND 11 304 0 0
ARCHER DANIELS MIDLAND CO COM 039483102 2,452 37,359 SH   DFND   37,359 0 0
ARCHROCK INC COM 03957W106 11 1,206 SH   DFND 2 1,206 0 0
ARCHROCK INC COM 03957W106 352 40,102 SH   DFND 3 40,102 0 0
ARCHROCK INC COM 03957W106 0 4 SH   DFND 5 4 0 0
ARCHROCK INC COM 03957W106 194 22,132 SH   DFND 6 0 0 22,132
ARCONIC CORPORATION COM 03966V107 96 2,665 SH   DFND 2 2,665 0 0
ARCONIC CORPORATION COM 03966V107 4,057 112,800 SH   DFND 3 112,800 0 0
ARCONIC CORPORATION COM 03966V107 0 4 SH   DFND 5 4 0 0
ARCONIC CORPORATION COM 03966V107 106 2,935 SH   DFND 6 0 0 2,935
ARCONIC CORPORATION COM 03966V107 814 22,623 SH   DFND   22,623 0 0
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ARES CAPITAL CORP COM 04010L103 87 4,457 SH   DFND 6 0 0 4,457
ARES CAPITAL CORP COM 04010L103 694 35,450 SH   DFND 7 35,450 0 0
ARES CAPITAL CORP COM 04010L103 8 418 SH   DFND 11 418 0 0
ARES CAPITAL CORP COM 04010L103 85 4,315 SH   DFND   4,315 0 0
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ARGO GROUP INTL HLDGS LTD COM G0464B107 79 1,520 SH   DFND 6 0 0 1,520
ARGO GROUP INTL HLDGS LTD COM G0464B107 16 300 SH   DFND   300 0 0
ARISTA NETWORKS INC COM 040413106 22 60 SH   DFND 2 60 0 0
ARISTA NETWORKS INC COM 040413106 6,359 17,471 SH   DFND 3 17,471 0 0
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ARISTA NETWORKS INC COM 040413106 358 983 SH   DFND 6 0 0 983
ARISTA NETWORKS INC COM 040413106 189 519 SH   DFND 7 245 0 274
ARISTA NETWORKS INC COM 040413106 2 5 SH   DFND 9 5 0 0
ARISTA NETWORKS INC COM 040413106 3,202 8,797 SH   DFND   8,797 0 0
ARK ETF TR INNOVATION ETF 00214Q104 78,528 600,000 SH Call DFND 5 600,000 0 0
ARK ETF TR INNOVATION ETF 00214Q104 1,045 7,981 SH   DFND 7 7,981 0 0
ARK ETF TR INNOVATION ETF 00214Q104 29 225 SH   DFND 9 225 0 0
ARK ETF TR INNOVATION ETF 00214Q104 31 240 SH   DFND 11 240 0 0
ARK ETF TR INNOVATION ETF 00214Q104 18,093 138,242 SH   DFND   138,242 0 0
ARK ETF TR INNOVATION ETF 00214Q104 7,984 61,000 SH Put DFND   61,000 0 0
ARK ETF TR AUTNMUS TECHNLGY 00214Q203 5 60 SH   DFND 7 60 0 0
ARK ETF TR AUTNMUS TECHNLGY 00214Q203 14 162 SH   DFND 9 162 0 0
ARK ETF TR AUTNMUS TECHNLGY 00214Q203 9 100 SH   DFND 11 100 0 0
ARK ETF TR AUTNMUS TECHNLGY 00214Q203 211 2,420 SH   DFND   2,420 0 0
ARK ETF TR GENOMIC REV ETF 00214Q302 67 707 SH   DFND 7 707 0 0
ARK ETF TR GENOMIC REV ETF 00214Q302 36 382 SH   DFND 9 382 0 0
ARK ETF TR GENOMIC REV ETF 00214Q302 28 300 SH   DFND 11 300 0 0
ARK ETF TR GENOMIC REV ETF 00214Q302 596 6,326 SH   DFND   6,326 0 0
ARK ETF TR NEXT GNRTN INTER 00214Q401 258 1,655 SH   DFND 7 1,655 0 0
ARK ETF TR NEXT GNRTN INTER 00214Q401 27 174 SH   DFND 11 174 0 0
ARK ETF TR NEXT GNRTN INTER 00214Q401 259 1,659 SH   DFND   1,659 0 0
ARK ETF TR 3D PRINTING ETF 00214Q500 18 437 SH   DFND 9 437 0 0
ARK ETF TR 3D PRINTING ETF 00214Q500 120 2,945 SH   DFND   2,945 0 0
ARK ETF TR ISRAEL INOVATE 00214Q609 78 2,395 SH   DFND 7 2,395 0 0
ARK ETF TR ISRAEL INOVATE 00214Q609 6 185 SH   DFND   185 0 0
ARK ETF TR FINTECH INNOVA 00214Q708 27 500 SH   DFND 11 500 0 0
ARK ETF TR FINTECH INNOVA 00214Q708 260 4,740 SH   DFND   4,740 0 0
ARK ETF TR ARK SPACE EXPL 00214Q807 45 2,100 SH   DFND 11 2,100 0 0
ARK ETF TR ARK SPACE EXPL 00214Q807 43 2,011 SH   DFND   2,011 0 0
ARLINGTON ASSET INVST CORP CL A NEW 041356205 1,540 366,615 SH   DFND 1 366,615 0 0
ARLINGTON ASSET INVST CORP CL A NEW 041356205 9 2,050 SH   DFND 6 0 0 2,050
ARLO TECHNOLOGIES INC COM 04206A101 22 3,128 SH   DFND 2 3,128 0 0
ARLO TECHNOLOGIES INC COM 04206A101 173 24,574 SH   DFND 3 24,574 0 0
ARLO TECHNOLOGIES INC COM 04206A101 0 3 SH   DFND 5 3 0 0
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ARMADA HOFFLER PPTYS INC COM 04208T108 8 620 SH   DFND 2 620 0 0
ARMADA HOFFLER PPTYS INC COM 04208T108 714 52,520 SH   DFND 3 52,520 0 0
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ARMADA HOFFLER PPTYS INC COM 04208T108 70 5,141 SH   DFND 6 0 0 5,141
ARMADA HOFFLER PPTYS INC COM 04208T108 141 10,351 SH   DFND   10,351 0 0
ARMOUR RESIDENTIAL REIT INC COM NEW 042315507 240 19,948 SH   DFND 3 19,948 0 0
ARMOUR RESIDENTIAL REIT INC COM NEW 042315507 0 2 SH   DFND 5 2 0 0
ARMOUR RESIDENTIAL REIT INC COM NEW 042315507 5 438 SH   DFND 6 0 0 438
ARMSTRONG WORLD INDS INC NEW COM 04247X102 4 36 SH   DFND 5 36 0 0
ARMSTRONG WORLD INDS INC NEW COM 04247X102 21 199 SH   DFND 6 0 0 199
ARMSTRONG WORLD INDS INC NEW COM 04247X102 7 67 SH   DFND 7 67 0 0
ARMSTRONG WORLD INDS INC NEW COM 04247X102 7,853 72,900 SH   DFND 8 72,900 0 0
ARMSTRONG WORLD INDS INC NEW COM 04247X102 2 23 SH   DFND 9 23 0 0
ARRAY TECHNOLOGIES INC COM SHS 04271T100 195 11,900 SH   DFND 3 11,900 0 0
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ARRAY TECHNOLOGIES INC COM SHS 04271T100 194 11,836 SH   DFND   11,836 0 0
ARROW ELECTRS INC COM 042735100 174 1,525 SH   DFND 2 1,525 0 0
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ARROW ELECTRS INC COM 042735100 144 1,264 SH   DFND 6 0 0 1,264
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ARROW FINL CORP COM 042744102 72 1,950 SH   DFND 6 0 0 1,950
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ASANA INC CL A 04342Y104 1,320 20,625 SH   DFND 7 6,945 0 13,680
ASANA INC CL A 04342Y104 107 1,680 SH   DFND   1,680 0 0
ASBURY AUTOMOTIVE GROUP INC COM 043436104 46 277 SH   DFND 2 277 0 0
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ASBURY AUTOMOTIVE GROUP INC COM 043436104 64 383 SH   DFND 6 0 0 383
ASCENDANT DIGITAL ACQUISITIO COM CL A G05155109 23,537 2,358,439 SH   DFND 5 2,358,439 0 0
ASE TECHNOLOGY HLDG CO LTD SPONSORED ADS 00215W100 7 813 SH   DFND 6 0 0 813
ASE TECHNOLOGY HLDG CO LTD SPONSORED ADS 00215W100 116 14,432 SH   DFND   14,432 0 0
ASGN INC COM 00191U102 424 4,317 SH   DFND 2 2,000 0 2,317
ASGN INC COM 00191U102 296 3,014 SH   DFND 3 3,014 0 0
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ASHLAND GLOBAL HLDGS INC COM 044186104 78 884 SH   DFND 2 884 0 0
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ASHLAND GLOBAL HLDGS INC COM 044186104 3 35 SH   DFND   35 0 0
ASML HOLDING N V N Y REGISTRY SHS N07059210 5,258 7,465 SH   DFND 3 7,465 0 0
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ASML HOLDING N V N Y REGISTRY SHS N07059210 46,789 66,432 SH   DFND   66,017 0 415
ASPEN AEROGELS INC COM 04523Y105 1,111 45,001 SH   DFND 5 45,001 0 0
ASPEN TECHNOLOGY INC COM 045327103 168 1,193 SH   DFND 2 1,193 0 0
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ASPEN TECHNOLOGY INC COM 045327103 313 2,218 SH   DFND   2,218 0 0
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ASPIRATIONAL CONSUMER LIFEST COM G05436103 2 200 SH   DFND   200 0 0
ASSOCIATED BANC CORP COM 045487105 14,161 688,418 SH   DFND 2 688,418 0 0
ASSOCIATED BANC CORP COM 045487105 182 8,845 SH   DFND 3 8,845 0 0
ASSOCIATED BANC CORP COM 045487105 42 2,025 SH   DFND 6 0 0 2,025
ASSOCIATED BANC CORP COM 045487105 2,894 140,711 SH   DFND 7 12,010 0 128,701
ASSURANT INC COM 04621X108 991 6,250 SH   DFND 2 6,250 0 0
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ASSURANT INC COM 04621X108 259 1,633 SH   DFND 6 0 0 1,633
ASSURANT INC COM 04621X108 196 1,236 SH   DFND 7 536 0 700
ASSURED GUARANTY LTD COM G0585R106 55 1,155 SH   DFND 2 1,155 0 0
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ASSURED GUARANTY LTD COM G0585R106 4 79 SH   DFND 5 79 0 0
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ASTEC INDS INC COM 046224101 9,731 154,110 SH   DFND 2 151,874 0 2,236
ASTEC INDS INC COM 046224101 442 6,995 SH   DFND 3 6,995 0 0
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ASTEC INDS INC COM 046224101 1,889 29,914 SH   DFND 7 629 0 29,285
ASTRAZENECA PLC SPONSORED ADR 046353108 606 10,067 SH   DFND 6 0 0 10,067
ASTRAZENECA PLC SPONSORED ADR 046353108 2,175 36,110 SH   DFND 7 35,607 0 503
ASTRAZENECA PLC SPONSORED ADR 046353108 30 491 SH   DFND 9 491 0 0
ASTRAZENECA PLC SPONSORED ADR 046353108 82 1,355 SH   DFND 11 1,355 0 0
ASTRAZENECA PLC SPONSORED ADR 046353108 329,544 5,470,522 SH   DFND   5,470,522 0 0
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ATHENE HOLDING LTD CL A G0684D107 339 5,000 SH   DFND 8 5,000 0 0
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ATLANTIC AVE ACQUISITION COR COM CL A 04906P101 2,438 250,000 SH   DFND 8 250,000 0 0
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ATLANTICA SUSTAINABLE INFR P SHS G0751N103 984 26,031 SH   DFND   26,031 0 0
ATLANTICUS HOLDINGS CORP COM 04914Y102 2,756 62,128 SH   DFND 2 60,187 0 1,941
ATLANTICUS HOLDINGS CORP COM 04914Y102 348 7,843 SH   DFND 4 7,843 0 0
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ATLAS AIR WORLDWIDE HLDGS IN COM NEW 049164205 269 3,927 SH   DFND 6 0 0 3,927
ATLAS AIR WORLDWIDE HLDGS IN COM NEW 049164205 7 100 SH   DFND   100 0 0
ATLAS AIR WORLDWIDE HLDGS IN NOTE 2.250% 6/0 049164BH8 4,357 4,022,000 PRN   DFND 8 4,022,000 0 0
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AUTHENTIC EQUITY ACQUISTN CO CL A SHS G06607108 305 31,500 SH   DFND 5 31,500 0 0
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CHANNELADVISOR CORP COM 159179100 65 2,624 SH   DFND 7 2,624 0 0
CHARLES RIV LABS INTL INC COM 159864107 2,485 6,679 SH   DFND 2 6,679 0 0
CHARLES RIV LABS INTL INC COM 159864107 4,837 13,000 SH   DFND 3 13,000 0 0
CHARLES RIV LABS INTL INC COM 159864107 7,969 21,417 SH   DFND 5 21,417 0 0
CHARLES RIV LABS INTL INC COM 159864107 170 456 SH   DFND 6 0 0 456
CHARLES RIV LABS INTL INC COM 159864107 47 125 SH   DFND 7 125 0 0
CHARLES RIV LABS INTL INC COM 159864107 1,674 4,500 SH   DFND 8 4,500 0 0
CHARLES RIV LABS INTL INC COM 159864107 56 150 SH   DFND 9 150 0 0
CHARLES RIV LABS INTL INC COM 159864107 54 146 SH   DFND   146 0 0
CHART INDS INC COM 16115Q308 186 1,268 SH   DFND 2 1,268 0 0
CHART INDS INC COM 16115Q308 1,595 10,861 SH   DFND 3 10,861 0 0
CHART INDS INC COM 16115Q308 0 1 SH   DFND 5 1 0 0
CHART INDS INC COM 16115Q308 316 2,153 SH   DFND 6 0 0 2,153
CHART INDS INC COM 16115Q308 23 160 SH   DFND   160 0 0
CHARTER COMMUNICATIONS INC N CL A 16119P108 30,934 42,946 SH   DFND 2 42,946 0 0<