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Collaboration and License Agreements - Revenue (Details) - USD ($)
$ in Thousands
3 Months Ended
Dec. 31, 2024
Dec. 31, 2023
Disaggregation of Revenue [Line Items]    
Revenue $ 2,500 $ 3,551
GSK    
Disaggregation of Revenue [Line Items]    
Revenue 2,500 2,685
Takeda    
Disaggregation of Revenue [Line Items]    
Revenue $ 0 $ 866