X0306 5 2016-12-31 0 0 0 0000875357 BOK FINANCIAL CORP ET AL BOKF 0001640868 Keesling Rebecca D C/O FREDERIC DORWART, LAWYERS 124 EAST FOURTH STREET TULSA OK 74103 0 1 0 0 EVP, Chief Auditor Common Stock 2016-12-31 5 J 0 9.42 0 A 353.78 I by IRA Common Stock 2016-12-31 5 J 0 17.4526 0 A 7271.4526 D These shares were acquired through a BOK Financial Corporation dividend reinvestment plan in which Ms. Keesling is a participant. These shares are related to the BOK Thrift Plan for which employees own investment units which hold BOKF common stock. 17.4526 shares are related to Ms. Keesling's BOK Thrift Plan holdings only. Ms. Keesling also owns 7,254 shares directly as of 12/31/2016. /s/ Frederic Dorwart, Power of Attorney 2017-02-14