The Securities and Exchange Commission has not necessarily reviewed the information in this filing and has not determined if it is accurate and complete.
The reader should not assume that the information is accurate and complete.

UNITED STATES SECURITIES AND EXCHANGE COMMISSION
Washington, D.C. 20549
FORM 13F

FORM 13F INFORMATION TABLE
OMB APPROVAL
OMB Number: 3235-0006
Estimated average burden
hours per response: 23.8
COLUMN 1 COLUMN 2 COLUMN 3 COLUMN 4 COLUMN 5 COLUMN 6 COLUMN 7 COLUMN 8
VALUE SHRS OR SH/ PUT/ INVESTMENT OTHER VOTING AUTHORITY
NAME OF ISSUER TITLE OF CLASS CUSIP (x$1000) PRN AMT PRN CALL DISCRETION MANAGER SOLE SHARED NONE
AXON ENTERPRISE INC COM 05464C101 34,724 382,854 SH   DFND 2 0 380,804 2,050
AXON ENTERPRISE INC COM 05464C101 895 9,869 SH   DFND 16 0 0 9,869
AXOS FINANCIAL INC COM 05465C100 35,584 1,526,577 SH   DFND 2 0 1,469,777 56,800
AXOS FINANCIAL INC COM 05465C100 474 20,320 SH   DFND 2,3 0 20,320 0
AXOS FINANCIAL INC COM 05465C100 442 18,961 SH   DFND 16 0 16,439 2,522
AXONICS MODULATION TECHNOLOG COM 05465P101 200 3,915 SH   DFND 2 0 3,915 0
AXONICS MODULATION TECHNOLOG COM 05465P101 713 13,960 SH   DFND 16 0 0 13,960
AZEK CO INC/THE COM 05478C105 1,275 36,641 SH   DFND 2 0 36,641 0
AZEK CO INC/THE COM 05478C105 204 5,848 SH   DFND 16 0 5,848 0
B&G FOODS INC COM 05508R106 8,792 316,268 SH   DFND 2 0 308,917 7,351
B&G FOODS INC COM 05508R106 659 23,748 SH   DFND 16 0 0 23,748
BCE INC COM 05534B760 9 203 SH   DFND 2 0 203 0
BCE INC COM 05534B760 53,882 1,288,215 SH   DFND 16 0 589,902 698,313
BCE INC COM 05534B760 42 1,019 SH   DFND 16,18 0 1,019 0
BCE INC COM 05534B760 283 6,824 SH   DFND 18 0 6,344 480
BG STAFFING INC COM 05544A109 193 22,800 SH   DFND 2 0 22,800 0
BG STAFFING INC COM 05544A109 3 300 SH   DFND 16 0 0 300
BHP GROUP PLC-ADR ADR 05545E209 5,625 131,837 SH   DFND 2 0 0 131,837
BHP GROUP PLC-ADR ADR 05545E209 993 23,269 SH   DFND 16 0 0 23,269
BJ'S WHOLESALE CLUB HOLDINGS COM 05550J101 33,134 797,445 SH   DFND 2 0 750,445 47,000
BJ'S WHOLESALE CLUB HOLDINGS COM 05550J101 719 17,300 SH   DFND 2,3 0 17,300 0
BJ'S WHOLESALE CLUB HOLDINGS COM 05550J101 769 18,620 SH   DFND 16 0 15,659 2,961
BJ'S WHOLESALE CLUB HOLDINGS COM 05550J101 47,972 1,154,559 SH   DFND 17 0 1,154,559 0
BOK FINANCIAL CORPORATION COM 05561Q201 607 11,784 SH   DFND 2 0 11,784 0
BOK FINANCIAL CORPORATION COM 05561Q201 3 57 SH   DFND 16 0 0 57
BOK FINANCIAL CORPORATION COM 05561Q201 3 63 SH   DFND 18 0 0 63
BP PLC-SPONS ADR ADR 055622104 19,027 1,089,730 SH   DFND 2 0 0 1,089,730
BP PLC-SPONS ADR ADR 055622104 12,195 698,258 SH   DFND 16 0 0 698,258
BP PLC-SPONS ADR ADR 055622104 9 500 SH   DFND 16,18 0 0 500
BP PLC-SPONS ADR ADR 055622104 103 5,926 SH   DFND 18 0 0 5,926
BNYM STRAT MUNI COM 05588W108 790 95,364 SH   DFND 16 0 0 95,364
BNYM STRAT MUNI COM 05588W108 50 5,992 SH   DFND 18 0 5,992 0
BRP GROUP INC-A COM 05589G102 2,040 81,903 SH   DFND 2 0 81,903 0
BNYM MUNICIPAL INCOME COM 05589T104 162 18,721 SH   DFND 16 0 0 18,721
BMC STOCK HOLDINGS INC COM 05591B109 384 8,980 SH   DFND 2 0 8,980 0
BMC STOCK HOLDINGS INC COM 05591B109 13 307 SH   DFND 16 0 0 307
BWX TECHNOLOGIES INC COM 05605H100 25,190 447,339 SH   DFND 2 0 447,339 0
BWX TECHNOLOGIES INC COM 05605H100 6,403 113,677 SH   DFND 16 0 112,373 1,304
BWX TECHNOLOGIES INC COM 05605H100 29 520 SH   DFND 16,18 0 520 0
BWX TECHNOLOGIES INC COM 05605H100 52,941 940,174 SH   DFND 17 0 940,174 0
AVERY DENNISON CORP COM 053611109 55,766 436,221 SH   DFND 17 0 436,221 0
AVERY DENNISON CORP COM 053611109 0 1 SH   DFND 18 0 1 0
AVIENT CORP COM 05368V106 4,541 171,501 SH   DFND 2 0 169,401 2,100
AVIENT CORP COM 05368V106 28 1,048 SH   DFND 16 0 0 1,048
AVIENT CORP COM 05368V106 36,249 1,369,960 SH   DFND 17 0 1,369,960 0
AVIENT CORP COM 05368V106 3 114 SH   DFND 18 0 0 114
AVIS BUDGET GROUP INC COM 053774105 2,550 96,883 SH   DFND 2 0 94,565 2,318
AVIS BUDGET GROUP INC COM 053774105 101 3,842 SH   DFND 16 0 0 3,842
AVISTA CORP COM 05379B107 12,050 353,164 SH   DFND 2 0 347,914 5,250
AVISTA CORP COM 05379B107 18,502 542,259 SH   DFND 10 0 542,259 0
AVISTA CORP COM 05379B107 16 464 SH   DFND 16 0 0 464
AVISTA CORP COM 05379B107 4 110 SH   DFND 18 0 0 110
AVNET INC COM 053807103 11,192 433,111 SH   DFND 2 0 428,811 4,300
AVNET INC COM 053807103 30 1,171 SH   DFND 16 0 0 1,171
AVNET INC COM 053807103 2 70 SH   DFND 18 0 70 0
AXCELIS TECHNOLOGIES INC COM 054540208 3,269 148,569 SH   DFND 2 0 145,769 2,800
AXCELIS TECHNOLOGIES INC COM 054540208 2 88 SH   DFND 16 0 0 88
AVROBIO INC COM 05455M100 1,178 90,460 SH   DFND 2 0 90,460 0
AVROBIO INC COM 05455M100 2 170 SH   DFND 16 0 0 170
AXOGEN INC COM 05463X106 1,726 148,397 SH   DFND 2 0 148,397 0
AXOGEN INC COM 05463X106 467 40,135 SH   DFND 16 0 39,735 400
BIG LOTS INC COM 089302103 19,984 448,079 SH   DFND 2 0 444,379 3,700
BIG LOTS INC COM 089302103 2,780 62,333 SH   DFND 16 0 0 62,333
BIGCOMMERCE HOLDINGS-SER 1 COM 08975P108 2,089 25,085 SH   DFND 2 0 25,085 0
BIGCOMMERCE HOLDINGS-SER 1 COM 08975P108 84 1,012 SH   DFND 16 0 688 324
BILIBILI INC-SPONSORED ADR ADR 090040106 108 2,601 SH   DFND 2 0 2,601 0
BILIBILI INC-SPONSORED ADR ADR 090040106 651 15,642 SH   DFND 16 0 0 15,642
BILL.COM HOLDINGS INC COM 090043100 74,186 739,568 SH   DFND 2 0 739,568 0
BILL.COM HOLDINGS INC COM 090043100 132 1,312 SH   DFND 2,3 0 1,312 0
BILL.COM HOLDINGS INC COM 090043100 43,786 436,504 SH   DFND 10 0 320,100 116,404
BILL.COM HOLDINGS INC COM 090043100 446 4,446 SH   DFND 16 0 4,105 341
BIO-RAD LABORATORIES-A COM 090572207 30,763 59,682 SH   DFND 2 0 59,682 0
BIO-RAD LABORATORIES-A COM 090572207 36 70 SH   DFND 2,3 0 70 0
BIO-RAD LABORATORIES-A COM 090572207 13,740 26,663 SH   DFND 16 0 24,196 2,467
BIO-RAD LABORATORIES-A COM 090572207 42,268 82,000 SH   DFND 17 0 82,000 0
BIOCRYST PHARMACEUTICALS INC COM 09058V103 0 35 SH   DFND 2 0 35 0
BIOCRYST PHARMACEUTICALS INC COM 09058V103 174 50,589 SH   DFND 16 0 50,049 540
BIOMARIN PHARMACEUTICAL INC COM 09061G101 105,201 1,382,773 SH   DFND 2 0 1,232,378 150,395
BIOMARIN PHARMACEUTICAL INC COM 09061G101 96,387 1,268,877 SH   DFND 10 0 1,268,877 0
BIOMARIN PHARMACEUTICAL INC COM 09061G101 1,100 14,454 SH   DFND 16 0 5,443 9,011
BIOMARIN PHARMACEUTICAL INC COM 09061G101 25,867 340,000 SH   DFND 17 0 340,000 0
BIOMARIN PHARMACEUTICAL INC COM 09061G101 4 56 SH   DFND 18 0 0 56
BIOLIFE SOLUTIONS INC COM 09062W204 26,283 908,196 SH   DFND 2 0 908,196 0
BIOGEN INC COM 09062X103 28,875 101,793 SH   DFND 2 0 101,781 12
BIOGEN INC COM 09062X103 155 545 SH   DFND 2,3 0 545 0
BIOGEN INC COM 09062X103 6,910 24,349 SH   DFND 16 0 2,398 21,951
BIOGEN INC COM 09062X103 18 63 SH   DFND 16,18 0 63 0
BIOGEN INC COM 09062X103 56 199 SH   DFND 18 0 199 0
BIOTELEMETRY INC COM 090672106 10,055 220,596 SH   DFND 2 0 217,896 2,700
BIOTELEMETRY INC COM 090672106 477 10,471 SH   DFND 16 0 0 10,471
BIOCEPT INC COM 09072V501 70 15,805 SH   DFND 16 0 0 15,805
BIO-TECHNE CORP COM 09073M104 111,435 449,827 SH   DFND 2 0 448,597 1,230
BIO-TECHNE CORP COM 09073M104 199 802 SH   DFND 2,3 0 802 0
BIO-TECHNE CORP COM 09073M104 398 1,607 SH   DFND 16 0 1,345 262
BIO-TECHNE CORP COM 09073M104 8 34 SH   DFND 16,18 0 34 0
BIO-TECHNE CORP COM 09073M104 57,350 231,502 SH   DFND 17 0 231,502 0
BIOSIG TECHNOLOGIES INC COM 09073N201 205 41,508 SH   DFND 16 0 0 41,508
BIONTECH SE-ADR ADR 09075V102 44,523 643,112 SH   DFND 2 0 643,112 0
BIOSPECIFICS TECHNOLOGIES COM 090931106 954 18,049 SH   DFND 2 0 18,049 0
BIOSPECIFICS TECHNOLOGIES COM 090931106 145 2,746 SH   DFND 16 0 0 2,746
BJ'S RESTAURANTS INC COM 09180C106 2,986 101,406 SH   DFND 2 0 99,401 2,005
BJ'S RESTAURANTS INC COM 09180C106 3 100 SH   DFND 16 0 0 100
BLACKROCK FLT RT INC COM 091941104 1,519 135,261 SH   DFND 16 0 0 135,261
BLACK HILLS CORP COM 092113109 11,093 207,393 SH   DFND 2 0 204,893 2,500
BLACK HILLS CORP COM 092113109 481 8,919 SH   DFND 16 0 8,364 555
BLACK KNIGHT INC COM 09215C105 981 11,264 SH   DFND 2 0 11,264 0
BLACK KNIGHT INC COM 09215C105 2,971 34,099 SH   DFND 16 0 12,167 21,932
BLACKBAUD INC COM 09227Q100 4,989 89,354 SH   DFND 2 0 87,604 1,750
BLACKBAUD INC COM 09227Q100 41 738 SH   DFND 16 0 0 738
BLACKLINE INC COM 09239B109 57,030 636,275 SH   DFND 2 0 636,275 0
BLACKLINE INC COM 09239B109 55 615 SH   DFND 2,3 0 615 0
BLACKLINE INC COM 09239B109 47,545 530,457 SH   DFND 10 0 388,997 141,460
BLACKLINE INC COM 09239B109 3,181 35,492 SH   DFND 16 0 17,150 18,342
BLACKLINE INC COM 09239B109 51,447 573,992 SH   DFND 17 0 573,992 0
BLACKLINE INC COM 09239B109 1 15 SH   DFND 18 0 0 15
BLACKROCK MUNICIPAL INCOME Q COM 092479104 280 19,675 SH   DFND 16 0 0 19,675
BLACKROCK INVT QUALITY MUNI COM 09247D105 1,670 106,769 SH   DFND 16 0 0 106,769
BLACKROCK INCOME TRUST COM 09247F100 2,165 355,412 SH   DFND 16 0 0 355,412
BLACKROCK INC COM 09247X101 899,882 1,596,807 SH   DFND 2 0 1,442,908 153,899
BLACKROCK INC COM 09247X101 13,232 23,478 SH   DFND 2,3 0 23,478 0
BLACKROCK INC COM 09247X101 45,861 81,943 SH   DFND 16 0 6,854 75,089
BLACKROCK INC COM 09247X101 165 292 SH   DFND 16,18 0 292 0
BLACKROCK INC COM 09247X101 355 630 SH   DFND 18 0 630 0
BLCKRCK UTIL INFRA & PWR OPP COM 09248D104 6,459 296,679 SH   DFND 16 0 0 296,679
BLACKROCK CALIFOR MUNI IN TR COM 09248E102 395 29,935 SH   DFND 16 0 0 29,935
BLACKROCK MUNICIPAL INC TRST COM 09248F109 458 33,436 SH   DFND 16 0 0 33,436
BLACKROCK MUNICIPAL INC TRST COM 09248F109 233 16,984 SH   DFND 16,18 0 16,984 0
BLACKROCK STRATEGIC MUNICIPL COM 09248T109 191 14,295 SH   DFND 16 0 0 14,295
BLACKROCK TAXABLE MUNICIPAL COM 09248X100 21,753 839,215 SH   DFND 16 0 0 839,215
BLACKROCK TAXABLE MUNICIPAL COM 09248X100 152 5,868 SH   DFND 18 0 5,868 0
BLACKROCK CORE BOND TRUST COM 09249E101 1,830 116,832 SH   DFND 16 0 0 116,832
BLACKROCK MUN INC TRUST II COM 09249N101 171 11,507 SH   DFND 16 0 0 11,507
BLACKROCK LTD DURATION INC COM 09249W101 944 64,174 SH   DFND 16 0 0 64,174
BLACKROCK MUNI 2020 TERM TR COM 09249X109 481 31,971 SH   DFND 16 0 0 31,971
BLACKROCK ENHANCED GLOBAL DI COM 092501105 1,929 201,688 SH   DFND 16 0 0 201,688
BLACKROCK CREDIT ALLOCATION COM 092508100 543 38,179 SH   DFND 16 0 0 38,179
BLACKROCK LNG-TM MUN ADV TST COM 09250B103 253 21,436 SH   DFND 16 0 0 21,436
BLACKROCK ENRGY & RES COM 09250U101 709 118,276 SH   DFND 16 0 0 118,276
BLACKROCK HEALTH SCIENCES COM 09250W107 474 11,070 SH   DFND 16 0 0 11,070
BLACKROCK ENHANCED EQTY DVD COM 09251A104 2,998 411,316 SH   DFND 16 0 0 411,316
BLACKROCK ENHANCED EQTY DVD COM 09251A104 14 1,930 SH   DFND 18 0 1,930 0
BLACKROCK ENHANCED INTERNATI COM 092524107 337 63,971 SH   DFND 16 0 0 63,971
BLACKROCK CAPITAL INVESTMENT COM 092533108 0 132 SH   DFND 2 0 132 0
BLACKROCK CAPITAL INVESTMENT COM 092533108 66 27,114 SH   DFND 16 0 0 27,114
BLACKROCK MUNIHOLDINGS FUND COM 09253N104 471 30,998 SH   DFND 16 0 0 30,998
BLACKROCK MUNIHOLDINGS FD II COM 09253P109 308 21,016 SH   DFND 16 0 0 21,016
BLACKROCK MUNIVEST FUND COM 09253R105 495 57,070 SH   DFND 16 0 0 57,070
BLACKROCK MUNIVEST FUND II COM 09253T101 1,243 89,044 SH   DFND 16 0 0 89,044
BLACKROCK MUNIVEST FUND II COM 09253T101 37 2,644 SH   DFND 18 0 2,644 0
BLACKROCK MUNIYIELD FUND COM 09253W104 1,155 85,173 SH   DFND 16 0 0 85,173
BLACKROCK MUNIYIELD FUND COM 09253W104 69 5,117 SH   DFND 18 0 5,117 0
BLACKROCK MUNIENHANCED FUND COM 09253Y100 434 38,411 SH   DFND 16 0 0 38,411
BLACKROCK MUNIHOLDINGS QUALI COM 09254A101 538 41,815 SH   DFND 16 0 0 41,815
BLACKROCK MUNIHOLDINGS QU II COM 09254C107 1,166 91,508 SH   DFND 16 0 0 91,508
BLACKROCK MUNIYIELD QUALITY COM 09254E103 1,052 78,298 SH   DFND 16 0 0 78,298
BLACKROCK MUNIYIELD QUALITY COM 09254E103 20 1,515 SH   DFND 18 0 1,515 0
BLACKROCK MUNIYIELD QUALITY COM 09254F100 544 33,984 SH   DFND 16 0 0 33,984
BLACKROCK MUNIYIELD QUAL II COM 09254G108 415 31,742 SH   DFND 16 0 0 31,742
BLACKROCK MUNIASSETS FUND COM 09254J102 766 55,667 SH   DFND 16 0 0 55,667
BLACKROCK MUNIHOLDINGS CA QU COM 09254L107 381 27,229 SH   DFND 16 0 0 27,229
BLACKROCK MUNIYIELD CALI FD COM 09254M105 248 17,890 SH   DFND 16 0 0 17,890
BLACKROCK MUNIHOLDINGS QUALI COM 09254P108 193 14,416 SH   DFND 16 0 0 14,416
BLACKROCK MUNIYIELD INVESTME COM 09254R104 685 51,234 SH   DFND 16 0 0 51,234
BLACKROCK MUNIHOLDINGS NY QU COM 09255C106 162 12,022 SH   DFND 16 0 0 12,022
BLACKROCK MUNIYIELD NY QUALI COM 09255E102 1,337 105,245 SH   DFND 16 0 0 105,245
BLACKROCK CORP HI YLD COM 09255P107 255 23,877 SH   DFND 16 0 0 23,877
BLACKROCK DEBT STRATEGIES FD COM 09255R202 448 45,111 SH   DFND 16 0 0 45,111
BLACKROCK DEBT STRATEGIES FD COM 09255R202 20 2,000 SH   DFND 18 0 2,000 0
BLACKROCK FLOAT RT INCOME ST COM 09255X100 347 30,195 SH   DFND 16 0 0 30,195
BLACKROCK ENH CAP AND INC COM 09256A109 877 56,727 SH   DFND 16 0 0 56,727
BLACKROCK ENH CAP AND INC COM 09256A109 6 400 SH   DFND 18 0 400 0
BLACKROCK RESOURCES & COMMOD COM 09257A108 8,497 1,430,762 SH   DFND 16 0 0 1,430,762
BLACKROCK MUNICIPAL 2030 TAR COM 09257P105 2,085 83,444 SH   DFND 16 0 0 83,444
BLACKROCK MUNICIPAL 2030 TAR COM 09257P105 37 1,500 SH   DFND 18 0 1,500 0
BLACKSTONE MORTGAGE TRU-CL A COM 09257W100 19,015 865,390 SH   DFND 2 0 833,090 32,300
BLACKSTONE MORTGAGE TRU-CL A COM 09257W100 279 12,708 SH   DFND 2,3 0 12,708 0
BLACKSTONE MORTGAGE TRU-CL A COM 09257W100 3,610 162,771 SH   DFND 16 0 0 162,771
BLACKSTONE MORTGAGE TRU-CL A COM 09257W100 11 512 SH   DFND 16,18 0 512 0
BLACKSTONE MORTGAGE TRU-CL A COM 09257W100 55 2,500 SH   DFND 18 0 2,500 0
BLACKSTONE MORTGAGE TRUST INC CONV BND 09257WAB6 19,947 20,750,000 PRN   DFND 2 0 0 20,750,000
BLACKSTONE MORTGAGE TRUST INC CONV BND 09257WAB6 9 9,000 PRN   DFND 16 0 0 9,000
BLACKROCK MULTI-SECTOR INCOM COM 09258A107 3,709 234,916 SH   DFND 16 0 0 234,916
BLACKROCK SCIENCE & TECH TR COM 09258G104 6,396 148,534 SH   DFND 16 0 0 148,534
BLACKROCK TCP CAPITAL CORP COM 09259E108 355 36,177 SH   DFND 16 0 0 36,177
BLACKSTONE GROUP INC/THE-A LP 09260D107 41 795 SH   DFND 2 0 0 795
BLACKSTONE GROUP INC/THE-A LP 09260D107 23,764 455,257 SH   DFND 16 0 0 455,257
BLACKSTONE GROUP INC/THE-A LP 09260D107 230 4,410 SH   DFND 16,18 0 0 4,410
BLACKSTONE GROUP INC/THE-A LP 09260D107 269 5,147 SH   DFND 18 0 0 5,147
BLACKROCK HEALTH SCI TR II COM 09260E105 3,320 139,204 SH   DFND 16 0 0 139,204
BLACKROCK SCIENCE & TECH II COM 09260K101 4,386 170,964 SH   DFND 16 0 0 170,964
BLINK CHARGING CO COM 09354A100 239 23,224 SH   DFND 16 0 0 23,224
H&R BLOCK INC COM 093671105 16,703 1,025,100 SH   DFND 2 0 983,100 42,000
H&R BLOCK INC COM 093671105 73 4,464 SH   DFND 16 0 168 4,296
BLOOM ENERGY CORP- A COM 093712107 223,311 12,426,729 SH   DFND 2 0 12,077,223 349,506
BLOOM ENERGY CORP- A COM 093712107 679 37,800 SH   DFND 2,3 0 37,800 0
BLOOM ENERGY CORP- A COM 093712107 177 9,844 SH   DFND 16 0 0 9,844
BLOOMIN' BRANDS INC COM 094235108 5,523 361,675 SH   DFND 2 0 354,475 7,200
BLOOMIN' BRANDS INC COM 094235108 159 10,399 SH   DFND 16 0 5,146 5,253
BLOOMIN' BRANDS INC COM 094235108 20 1,290 SH   DFND 18 0 1,290 0
BLUCORA INC COM 095229100 3,017 320,281 SH   DFND 2 0 320,281 0
BLUCORA INC COM 095229100 15 1,588 SH   DFND 16 0 0 1,588
BLUE BIRD CORP COM 095306106 655 53,940 SH   DFND 2 0 53,940 0
BLUEBIRD BIO INC COM 09609G100 1,296 24,020 SH   DFND 2 0 24,020 0
BLUEBIRD BIO INC COM 09609G100 1,196 22,199 SH   DFND 16 0 11,213 10,986
BLUEBIRD BIO INC COM 09609G100 7 124 SH   DFND 16,18 0 124 0
BLUEBIRD BIO INC COM 09609G100 1 21 SH   DFND 18 0 0 21
BLUEPRINT MEDICINES CORP COM 09627Y109 4,741 51,141 SH   DFND 2 0 51,141 0
BLUEPRINT MEDICINES CORP COM 09627Y109 35 381 SH   DFND 16 0 0 381
BLUEPRINT MEDICINES CORP COM 09627Y109 4 46 SH   DFND 18 0 0 46
BNYM HIGH YIELD STRAT COM 09660L105 526 194,007 SH   DFND 16 0 0 194,007
BNYM STRAT MUNI BND COM 09662E109 1,965 269,785 SH   DFND 16 0 0 269,785
BNYM STRAT MUNI BND COM 09662E109 47 6,400 SH   DFND 18 0 6,400 0
BNYM MUNI BND INFRA COM 09662W109 1,607 119,465 SH   DFND 16 0 0 119,465
BOEING CO/THE COM 097023105 144,017 871,458 SH   DFND 2 0 864,738 6,720
BOEING CO/THE COM 097023105 301 1,824 SH   DFND 2,3 0 1,824 0
BOEING CO/THE COM 097023105 33,917 205,414 SH   DFND 16 0 11,449 193,965
BOEING CO/THE COM 097023105 112 677 SH   DFND 16,18 0 677 0
BOEING CO/THE COM 097023105 228 1,379 SH   DFND 18 0 1,379 0
BADGER METER INC COM 056525108 8,531 130,497 SH   DFND 2 0 127,147 3,350
BADGER METER INC COM 056525108 2,377 36,323 SH   DFND 16 0 35,330 993
BADGER METER INC COM 056525108 9,018 137,953 SH   DFND 17 0 137,953 0
BADGER METER INC COM 056525108 6 95 SH   DFND 18 0 95 0
BAIDU INC - SPON ADR ADR 056752108 20,463 161,644 SH   DFND 2 0 161,644 0
BAIDU INC - SPON ADR ADR 056752108 9,375 73,487 SH   DFND 16 0 980 72,507
BAIDU INC - SPON ADR ADR 056752108 4 32 SH   DFND 18 0 32 0
BAKER HUGHES CO COM 05722G100 3,712 279,305 SH   DFND 2 0 279,167 138
BAKER HUGHES CO COM 05722G100 30 2,256 SH   DFND 2,3 0 2,256 0
BAKER HUGHES CO COM 05722G100 1,165 87,651 SH   DFND 16 0 67,563 20,088
BALCHEM CORP COM 057665200 43,286 443,370 SH   DFND 2 0 440,345 3,025
BALCHEM CORP COM 057665200 197 2,013 SH   DFND 16 0 1,751 262
BALL CORP COM 058498106 25,922 311,864 SH   DFND 2 0 311,864 0
BALL CORP COM 058498106 94 1,129 SH   DFND 2,3 0 1,129 0
BALL CORP COM 058498106 13,916 167,524 SH   DFND 16 0 145,853 21,671
BALL CORP COM 058498106 115 1,381 SH   DFND 18 0 1,381 0
BANCFIRST CORP COM 05945F103 13,093 320,554 SH   DFND 2 0 319,604 950
BANCFIRST CORP COM 05945F103 25 620 SH   DFND 16 0 589 31
BANCO BRADESCO-ADR ADR 059460303 27 7,600 SH   DFND 2 0 0 7,600
BANCO BRADESCO-ADR ADR 059460303 122 35,910 SH   DFND 16 0 0 35,910
BANCO BILBAO VIZCAYA-SP ADR ADR 05946K101 372 135,726 SH   DFND 2 0 0 135,726
BANCO BILBAO VIZCAYA-SP ADR ADR 05946K101 9 3,365 SH   DFND 16 0 0 3,365
BANCO SANTANDER SA-SPON ADR ADR 05964H105 9 4,656 SH   DFND 2 0 0 4,656
BANCO SANTANDER SA-SPON ADR ADR 05964H105 1,133 612,343 SH   DFND 16 0 0 612,343
BANCO SANTANDER SA-SPON ADR ADR 05964H105 1 344 SH   DFND 18 0 0 344
BANCOLOMBIA S.A.-SPONS ADR ADR 05968L102 81 3,131 SH   DFND 2 0 0 3,131
BANCOLOMBIA S.A.-SPONS ADR ADR 05968L102 7,808 305,715 SH   DFND 10 0 0 305,715
BANCOLOMBIA S.A.-SPONS ADR ADR 05968L102 2 73 SH   DFND 16 0 0 73
BANCROFT FUND LTD COM 059695106 316 12,713 SH   DFND 16 0 0 12,713
BANCORP INC/THE COM 05969A105 12,692 1,468,886 SH   DFND 2 0 1,366,886 102,000
BANCORP INC/THE COM 05969A105 350 40,500 SH   DFND 2,3 0 40,500 0
BANCORP INC/THE COM 05969A105 3 366 SH   DFND 16 0 0 366
BANCORPSOUTH BANK COM 05971J102 3,352 172,943 SH   DFND 2 0 169,443 3,500
BANDWIDTH INC-CLASS A COM 05988J103 3,771 21,601 SH   DFND 2 0 21,601 0
BANDWIDTH INC-CLASS A COM 05988J103 421 2,411 SH   DFND 16 0 0 2,411
BANC OF CALIFORNIA INC COM 05990K106 1,991 196,685 SH   DFND 2 0 196,685 0
BANK OF AMERICA CORP COM 060505104 1,119,489 46,471,163 SH   DFND 2 0 40,977,453 5,493,710
BANK OF AMERICA CORP COM 060505104 10,127 420,425 SH   DFND 2,3 0 420,425 0
BANK OF AMERICA CORP COM 060505104 203,378 8,449,284 SH   DFND 10 0 8,449,284 0
BANK OF AMERICA CORP COM 060505104 57,394 2,379,697 SH   DFND 16 0 634,455 1,745,241
BANK OF AMERICA CORP COM 060505104 123 5,097 SH   DFND 16,18 0 5,097 0
BANK OF AMERICA CORP COM 060505104 564 23,404 SH   DFND 18 0 22,489 915
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BANK OF HAWAII CORP COM 062540109 134 2,652 SH   DFND 16 0 0 2,652
BANK OF HAWAII CORP COM 062540109 3 66 SH   DFND 18 0 66 0
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BANK OF MONTREAL COM 063671101 511 8,748 SH   DFND 2 0 8,748 0
BANK OF MONTREAL COM 063671101 1,461 25,018 SH   DFND 16 0 65 24,953
BANK OF MONTREAL COM 063671101 12 200 SH   DFND 18 0 200 0
BANK OF NEW YORK MELLON CORP COM 064058100 122,409 3,564,600 SH   DFND 2 0 2,590,650 973,950
BANK OF NEW YORK MELLON CORP COM 064058100 96 2,810 SH   DFND 2,3 0 2,810 0
BANK OF NEW YORK MELLON CORP COM 064058100 2,522 73,433 SH   DFND 16 0 37,857 35,576
BANK OF NEW YORK MELLON CORP COM 064058100 53 1,543 SH   DFND 18 0 1,543 0
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BANK OZK COM 06417N103 4,685 219,678 SH   DFND 16 0 90,826 128,852
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BANKUNITED INC COM 06652K103 105 4,814 SH   DFND 16 0 0 4,814
BANNER CORPORATION COM 06652V208 17,221 533,856 SH   DFND 2 0 532,056 1,800
BANNER CORPORATION COM 06652V208 4 114 SH   DFND 16 0 0 114
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IPATH BLOOMBERG COMMODITY IN COM 06738C778 97 4,970 SH   DFND 18 0 0 4,970
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BARCLAYS PLC-SPONS ADR ADR 06738E204 607 118,887 SH   DFND 16 0 0 118,887
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IPATH WOMEN IN LEADERSHIP ET COM 06742W430 260 3,642 SH   DFND 16 0 0 3,642
BARNES & NOBLE EDUCATION INC COM 06777U101 337 130,551 SH   DFND 2 0 130,551 0
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BARRICK GOLD CORP COM 067901108 10,357 352,197 SH   DFND 16 0 97,976 254,221
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BAXTER INTERNATIONAL INC COM 071813109 540,744 6,723,811 SH   DFND 2 0 5,533,244 1,190,567
BAXTER INTERNATIONAL INC COM 071813109 1,352 16,807 SH   DFND 2,3 0 16,807 0
BAXTER INTERNATIONAL INC COM 071813109 19,178 238,469 SH   DFND 2,10 0 238,469 0
BAXTER INTERNATIONAL INC COM 071813109 121,544 1,511,543 SH   DFND 10 0 1,511,543 0
BAXTER INTERNATIONAL INC COM 071813109 12,475 155,213 SH   DFND 16 0 12,931 142,282
BAXTER INTERNATIONAL INC COM 071813109 8 100 SH   DFND 16,18 0 100 0
BAXTER INTERNATIONAL INC COM 071813109 122 1,507 SH   DFND 18 0 1,401 106
BEAZER HOMES USA INC COM 07556Q881 528 39,945 SH   DFND 2 0 39,945 0
BECTON DICKINSON AND CO COM 075887109 259,050 1,113,331 SH   DFND 2 0 1,071,646 41,685
BECTON DICKINSON AND CO COM 075887109 7,184 30,873 SH   DFND 2,3 0 30,873 0
BECTON DICKINSON AND CO COM 075887109 29 140 SH   DFND 10 0 140 0
BECTON DICKINSON AND CO COM 075887109 21,345 91,716 SH   DFND 16 0 39,384 52,333
BECTON DICKINSON AND CO COM 075887109 83 355 SH   DFND 16,18 0 355 0
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BED BATH & BEYOND INC COM 075896100 501 33,444 SH   DFND 16 0 0 33,444
BED BATH & BEYOND INC COM 075896100 1 50 SH   DFND 18 0 50 0
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BEIGENE LTD-ADR ADR 07725L102 112 392 SH   DFND 2,3 0 392 0
BEIGENE LTD-ADR ADR 07725L102 2,001 6,987 SH   DFND 2,10 0 6,987 0
BEIGENE LTD-ADR ADR 07725L102 6,412 22,421 SH   DFND 10 0 22,421 0
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BEIGENE LTD-ADR ADR 07725L102 3 9 SH   DFND 18 0 0 9
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BEL FUSE INC-CL B COM 077347300 26 2,475 SH   DFND 16 0 0 2,475
BELDEN INC COM 077454106 4,351 139,821 SH   DFND 2 0 139,821 0
BELDEN INC COM 077454106 6 192 SH   DFND 16 0 0 192
BELLRING BRANDS INC-CLASS A COM 079823100 13,270 639,818 SH   DFND 2 0 585,818 54,000
BELLRING BRANDS INC-CLASS A COM 079823100 436 21,000 SH   DFND 2,3 0 21,000 0
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BELLUS HEALTH INC COM 07987C204 296 130,300 SH   DFND 2 0 130,300 0
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BENCHMARK ELECTRONICS INC COM 08160H101 118 5,871 SH   DFND 16 0 5,871 0
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BENEFITFOCUS INC COM 08180D106 17 1,500 SH   DFND 16 0 0 1,500
BERKELEY LIGHTS INC COM 084310101 19,314 252,936 SH   DFND 2 0 252,936 0
BERKELEY LIGHTS INC COM 084310101 36 475 SH   DFND 2,3 0 475 0
WR BERKLEY CORP COM 084423102 12,706 207,782 SH   DFND 2 0 207,755 27
WR BERKLEY CORP COM 084423102 29 480 SH   DFND 2,3 0 480 0
WR BERKLEY CORP COM 084423102 2,572 42,076 SH   DFND 16 0 37,609 4,467
WR BERKLEY CORP COM 084423102 2 37 SH   DFND 18 0 37 0
BERKSHIRE HATHAWAY INC-CL A COM 084670108 3,840 12 SH   DFND 2 0 12 0
BERKSHIRE HATHAWAY INC-CL A COM 084670108 1,920 6 SH   DFND 16 0 0 6
BERKSHIRE HATHAWAY INC-CL A COM 084670108 1,920 6 SH   DFND 18 0 6 0
BERKSHIRE HATHAWAY INC-CL B COM 084670702 962,180 4,518,548 SH   DFND 2 0 4,339,341 179,207
BERKSHIRE HATHAWAY INC-CL B COM 084670702 26,906 126,351 SH   DFND 2,3 0 126,351 0
BERKSHIRE HATHAWAY INC-CL B COM 084670702 144,359 678,085 SH   DFND 10 0 678,085 0
BERKSHIRE HATHAWAY INC-CL B COM 084670702 98,054 460,222 SH   DFND 16 0 61,822 398,401
BERKSHIRE HATHAWAY INC-CL B COM 084670702 38 180 SH   DFND 16,18 0 180 0
BERKSHIRE HATHAWAY INC-CL B COM 084670702 1,092 5,127 SH   DFND 18 0 5,106 21
BERKSHIRE HILLS BANCORP INC COM 084680107 3,313 327,619 SH   DFND 2 0 327,619 0
BERRY GLOBAL GROUP INC COM 08579W103 154 3,193 SH   DFND 2 0 3,193 0
BERRY GLOBAL GROUP INC COM 08579W103 212 4,396 SH   DFND 16 0 0 4,396
BERRY GLOBAL GROUP INC COM 08579W103 3 56 SH   DFND 18 0 0 56
BERRY CORP COM 08579X101 423 133,265 SH   DFND 2 0 133,265 0
BEST BUY CO INC COM 086516101 262,090 2,354,837 SH   DFND 2 0 1,822,027 532,810
BEST BUY CO INC COM 086516101 88 791 SH   DFND 2,3 0 791 0
BEST BUY CO INC COM 086516101 122,075 1,078,823 SH   DFND 16 0 33,091 1,045,732
BEST BUY CO INC COM 086516101 213 1,912 SH   DFND 16,18 0 1,912 0
BEST BUY CO INC COM 086516101 415 3,722 SH   DFND 18 0 3,660 62
BHP GROUP LTD-SPON ADR ADR 088606108 20,957 405,265 SH   DFND 2 0 0 405,265
BHP GROUP LTD-SPON ADR ADR 088606108 2,841 54,923 SH   DFND 16 0 0 54,923
BHP GROUP LTD-SPON ADR ADR 088606108 6 125 SH   DFND 18 0 0 125
BEYOND MEAT INC COM 08862E109 21,402 128,881 SH   DFND 2 0 128,881 0
BEYOND MEAT INC COM 08862E109 784 4,726 SH   DFND 16 0 328 4,398
BEYOND MEAT INC COM 08862E109 42,088 253,450 SH   DFND 17 0 253,450 0
BEYOND MEAT INC COM 08862E109 58 350 SH   DFND 18 0 350 0
BEYOND AIR INC COM 08862L103 832 160,321 SH   DFND 16 0 0 160,321
AAON INC COM 000360206 17,299 287,112 SH   DFND 2 0 284,162 2,950
AAON INC COM 000360206 22,358 371,145 SH   DFND 10 0 272,169 98,976
AAON INC COM 000360206 47 783 SH   DFND 16 0 0 783
AAR CORP COM 000361105 4,762 253,247 SH   DFND 2 0 253,247 0
AAR CORP COM 000361105 9 500 SH   DFND 16 0 0 500
AAR CORP COM 000361105 1 49 SH   DFND 18 0 49 0
ABB LTD-SPON ADR ADR 000375204 418 16,412 SH   DFND 2 0 0 16,412
ABB LTD-SPON ADR ADR 000375204 19,151 751,605 SH   DFND 16 0 0 751,605
ABB LTD-SPON ADR ADR 000375204 27 1,077 SH   DFND 18 0 0 1,077
ACCO BRANDS CORP COM 00081T108 735 126,882 SH   DFND 2 0 126,882 0
ACCO BRANDS CORP COM 00081T108 6 1,002 SH   DFND 16 0 0 1,002
ADMA BIOLOGICS INC COM 000899104 1 480 SH   DFND 2 0 480 0
ADMA BIOLOGICS INC COM 000899104 115 48,200 SH   DFND 16 0 0 48,200
ADT INC COM 00090Q103 40 4,824 SH   DFND 2 0 4,824 0
ADT INC COM 00090Q103 322 39,400 SH   DFND 16 0 28,308 11,092
ABM INDUSTRIES INC COM 000957100 11,155 304,275 SH   DFND 2 0 297,275 7,000
ABM INDUSTRIES INC COM 000957100 1,808 49,274 SH   DFND 16 0 48,819 455
ABM INDUSTRIES INC COM 000957100 1 40 SH   DFND 18 0 40 0
AFLAC INC COM 001055102 20,161 554,630 SH   DFND 2 0 554,630 0
AFLAC INC COM 001055102 83 2,279 SH   DFND 2,3 0 2,279 0
AFLAC INC COM 001055102 8,385 230,575 SH   DFND 16 0 40,856 189,719
AFLAC INC COM 001055102 71 1,965 SH   DFND 18 0 1,965 0
AGCO CORP COM 001084102 31,584 425,238 SH   DFND 2 0 422,888 2,350
AGCO CORP COM 001084102 67 897 SH   DFND 16 0 0 897
AGNC INVESTMENT CORP COM 00123Q104 167 11,905 SH   DFND 2 0 11,905 0
AGNC INVESTMENT CORP COM 00123Q104 1,125 80,766 SH   DFND 16 0 0 80,766
AGNC INVESTMENT CORP COM 00123Q104 101 7,263 SH   DFND 18 0 7,263 0
AES CORP COM 00130H105 433,124 23,916,300 SH   DFND 2 0 18,828,849 5,087,451
AES CORP COM 00130H105 5,652 312,052 SH   DFND 2,3 0 312,052 0
AES CORP COM 00130H105 4,836 256,905 SH   DFND 16 0 216,979 39,926
AES CORP COM 00130H105 41 2,250 SH   DFND 16,18 0 2,250 0
AES CORP COM 00130H105 8 418 SH   DFND 18 0 0 418
ALERIAN MLP ETF COM 00162Q452 4 212 SH   DFND 2 0 105 107
ALERIAN MLP ETF COM 00162Q452 4,193 209,855 SH   DFND 16 0 0 209,855
ALERIAN MLP ETF COM 00162Q452 57 2,843 SH   DFND 18 0 2,843 0
ALERIAN ENERGY INFRASTRUCTUR COM 00162Q676 600 48,675 SH   DFND 16 0 0 48,675
ALERIAN ENERGY INFRASTRUCTUR COM 00162Q676 12 1,000 SH   DFND 18 0 1,000 0
ALPS INTERNATIONAL SECTOR DI COM 00162Q718 303 14,047 SH   DFND 16 0 0 14,047
ALPS INTERNATIONAL SECTOR DI COM 00162Q718 43 2,003 SH   DFND 18 0 2,003 0
ALPS SECTOR DIVIDEND DOGS COM 00162Q858 2,120 56,070 SH   DFND 16 0 0 56,070
ALPS SECTOR DIVIDEND DOGS COM 00162Q858 34 902 SH   DFND 18 0 902 0
AMAG PHARMACEUTICALS INC COM 00163U106 1,999 212,667 SH   DFND 2 0 208,767 3,900
AMAG PHARMACEUTICALS INC CONV BND 00163UAD8 7,182 8,000,000 PRN   DFND 2 0 0 8,000,000
AMC NETWORKS INC-A COM 00164V103 1,900 76,905 SH   DFND 2 0 74,700 2,205
AMC NETWORKS INC-A COM 00164V103 635 25,720 SH   DFND 16 0 0 25,720
AMC NETWORKS INC-A COM 00164V103 0 12 SH   DFND 18 0 12 0
AMC ENTERTAINMENT HLDS-CL A COM 00165C104 77 16,445 SH   DFND 2 0 16,445 0
AMC ENTERTAINMENT HLDS-CL A COM 00165C104 169 35,847 SH   DFND 16 0 0 35,847
AMN HEALTHCARE SERVICES INC COM 001744101 18,633 318,742 SH   DFND 2 0 314,492 4,250
AMN HEALTHCARE SERVICES INC COM 001744101 581 9,925 SH   DFND 16 0 4,748 5,177
ANI PHARMACEUTICALS INC COM 00182C103 5,890 208,773 SH   DFND 2 0 207,373 1,400
API GROUP CORP COM 00187Y100 7,781 546,799 SH   DFND 2 0 499,899 46,900
API GROUP CORP COM 00187Y100 277 19,500 SH   DFND 2,3 0 19,500 0
API GROUP CORP COM 00187Y100 1 40 SH   DFND 16 0 0 40
ASGN INC COM 00191U102 13,386 210,612 SH   DFND 2 0 208,812 1,800
ASGN INC COM 00191U102 141 2,219 SH   DFND 16 0 672 1,547
AT&T INC COM 00206R102 461,185 16,176,242 SH   DFND 2 0 15,593,700 582,542
AT&T INC COM 00206R102 1,954 68,543 SH   DFND 2,3 0 68,543 0
AT&T INC COM 00206R102 1,906 67,001 SH   DFND 10 0 67,001 0
AT&T INC COM 00206R102 164,512 5,776,922 SH   DFND 16 0 1,469,354 4,307,568
AT&T INC COM 00206R102 461 16,170 SH   DFND 16,18 0 16,170 0
AT&T INC COM 00206R102 1,928 67,633 SH   DFND 18 0 66,502 1,131
ATN INTERNATIONAL INC COM 00215F107 2,501 49,859 SH   DFND 2 0 48,609 1,250
AZZ INC COM 002474104 13,680 400,955 SH   DFND 2 0 398,055 2,900
AZZ INC COM 002474104 14 400 SH   DFND 16 0 0 400
AARON'S INC COM 002535300 26,175 462,025 SH   DFND 2 0 458,725 3,300
AARON'S INC COM 002535300 3,382 59,641 SH   DFND 16 0 57,143 2,498
AARON'S INC COM 002535300 11 200 SH   DFND 16,18 0 200 0
ABBOTT LABORATORIES COM 002824100 920,133 8,454,746 SH   DFND 2 0 7,738,890 715,856
ABBOTT LABORATORIES COM 002824100 10,832 99,530 SH   DFND 2,3 0 99,530 0
ABBOTT LABORATORIES COM 002824100 21,178 194,598 SH   DFND 2,10 0 194,598 0
ABBOTT LABORATORIES COM 002824100 288,806 2,654,689 SH   DFND 10 0 2,654,689 0
ABBOTT LABORATORIES COM 002824100 213,730 1,950,813 SH   DFND 16 0 40,102 1,910,711
ABBOTT LABORATORIES COM 002824100 597 5,478 SH   DFND 16,18 0 5,478 0
ABBOTT LABORATORIES COM 002824100 1,299 11,926 SH   DFND 18 0 11,873 53
ABBVIE INC COM 00287Y109 521,897 5,958,386 SH   DFND 2 0 5,596,966 361,420
ABBVIE INC COM 00287Y109 532 6,075 SH   DFND 2,3 0 6,075 0
ABBVIE INC COM 00287Y109 172,472 1,982,860 SH   DFND 16 0 510,398 1,472,462
ABBVIE INC COM 00287Y109 886 10,209 SH   DFND 16,18 0 10,209 0
ABBVIE INC COM 00287Y109 944 10,781 SH   DFND 18 0 10,356 425
ABERCROMBIE & FITCH CO-CL A COM 002896207 3,911 280,764 SH   DFND 2 0 274,364 6,400
ABERDEEN ASIA-PAC INCOME FD COM 003009107 1,185 296,877 SH   DFND 16 0 900 295,977
ABERDEEN ASIA-PAC INCOME FD COM 003009107 16 3,900 SH   DFND 18 0 3,900 0
ABERDEEN GLBL PREM PROP COM 00302L108 1,377 302,474 SH   DFND 16 0 0 302,474
ABERDEEN GLBL PREM PROP COM 00302L108 0 15 SH   DFND 18 0 15 0
ABERDEEN TTL DYN DIVID COM 00326L100 1,012 127,620 SH   DFND 16 0 0 127,620
ABERDEEN TTL DYN DIVID COM 00326L100 37 4,684 SH   DFND 18 0 4,684 0
ABIOMED INC COM 003654100 141,389 510,311 SH   DFND 2 0 429,796 80,515
ABIOMED INC COM 003654100 9,676 34,923 SH   DFND 2,3 0 34,923 0
ABIOMED INC COM 003654100 5,380 19,420 SH   DFND 16 0 6,261 13,159
ABRAXAS PETROLEUM CORP COM 003830106 2 16,565 SH   DFND 2 0 16,565 0
ABRAXAS PETROLEUM CORP COM 003830106 4 26,225 SH   DFND 16 0 0 26,225
ACACIA COMMUNICATIONS INC COM 00401C108 5,939 88,119 SH   DFND 2 0 88,119 0
ACACIA COMMUNICATIONS INC COM 00401C108 144 2,140 SH   DFND 16 0 0 2,140
ACADIA HEALTHCARE CO INC COM 00404A109 6,308 213,963 SH   DFND 2 0 210,458 3,505
ACADIA HEALTHCARE CO INC COM 00404A109 18 587 SH   DFND 16 0 26 561
ACADIA PHARMACEUTICALS INC COM 004225108 4,504 109,173 SH   DFND 2 0 109,173 0
ACADIA PHARMACEUTICALS INC COM 004225108 2,985 72,364 SH   DFND 2,10 0 72,364 0
ACADIA PHARMACEUTICALS INC COM 004225108 2,729 66,190 SH   DFND 16 0 65,431 759
ACADIA PHARMACEUTICALS INC COM 004225108 39,942 968,280 SH   DFND 17 0 968,280 0
ACADIA REALTY TRUST COM 004239109 4,106 391,035 SH   DFND 2 0 385,635 5,400
ACADIA REALTY TRUST COM 004239109 12 1,114 SH   DFND 16 0 0 1,114
ACADIA REALTY TRUST COM 004239109 8 800 SH   DFND 18 0 800 0
ACAMAR PARTNERS ACQUISITI- A COM 004285102 517 50,539 SH   DFND 16 0 0 50,539
ACCELERON PHARMA INC COM 00434H108 1,489 13,238 SH   DFND 2 0 13,238 0
ACCELERON PHARMA INC COM 00434H108 279 2,481 SH   DFND 16 0 2,454 27
ACCELERON PHARMA INC COM 00434H108 91,281 811,172 SH   DFND 17 0 811,172 0
ACCELERON PHARMA INC COM 00434H108 3 23 SH   DFND 18 0 0 23
ACCOLADE INC COM 00437E102 23,505 604,679 SH   DFND 2 0 604,679 0
ACCOLADE INC COM 00437E102 133 3,434 SH   DFND 16 0 3,434 0
ACI WORLDWIDE INC COM 004498101 10,034 384,000 SH   DFND 2 0 379,300 4,700
ACI WORLDWIDE INC COM 004498101 501 19,192 SH   DFND 16 0 18,840 352
ACI WORLDWIDE INC COM 004498101 1 50 SH   DFND 18 0 0 50
ACORDA THERAPEUTICS INC COM 00484M106 27 52,538 SH   DFND 2 0 52,538 0
ACORDA THERAPEUTICS INC COM 00484M106 0 150 SH   DFND 16 0 0 150
ACORDA THERAPEUTICS INC CONV BND 00484MAA4 2,998 3,500,000 PRN   DFND 2 0 0 3,500,000
ACTIVISION BLIZZARD INC COM 00507V109 667,874 8,250,503 SH   DFND 2 0 7,524,284 726,219
ACTIVISION BLIZZARD INC COM 00507V109 21,012 259,566 SH   DFND 2,3 0 259,566 0
ACTIVISION BLIZZARD INC COM 00507V109 8,839 109,190 SH   DFND 2,10 0 109,190 0
ACTIVISION BLIZZARD INC COM 00507V109 125,624 1,552,827 SH   DFND 10 0 1,552,827 0
ACTIVISION BLIZZARD INC COM 00507V109 10,242 126,753 SH   DFND 16 0 36,554 90,199
ACTIVISION BLIZZARD INC COM 00507V109 19 239 SH   DFND 16,18 0 239 0
ACUITY BRANDS INC COM 00508Y102 15,921 155,564 SH   DFND 2 0 153,739 1,825
ACUITY BRANDS INC COM 00508Y102 23 221 SH   DFND 16 0 0 221
ACUSHNET HOLDINGS CORP COM 005098108 223 6,641 SH   DFND 2 0 6,641 0
ACUSHNET HOLDINGS CORP COM 005098108 5,859 174,320 SH   DFND 10 0 174,320 0
ACUSHNET HOLDINGS CORP COM 005098108 148 4,402 SH   DFND 16 0 0 4,402
ACUTUS MEDICAL INC COM 005111109 5,815 195,145 SH   DFND 2 0 195,145 0
ADAMS DIVERSIFIED EQUITY COM 006212104 285 17,581 SH   DFND 16 0 0 17,581
ADAPTIVE BIOTECHNOLOGIES COM 00650F109 28,444 584,893 SH   DFND 2 0 584,893 0
ADAPTIVE BIOTECHNOLOGIES COM 00650F109 72 1,486 SH   DFND 2,3 0 1,486 0
ADAPTIVE BIOTECHNOLOGIES COM 00650F109 244 5,029 SH   DFND 16 0 5,001 28
ADAPTHEALTH CORP COM 00653Q102 3,103 142,300 SH   DFND 2 0 142,300 0
ADAPTHEALTH CORP COM 00653Q102 325 14,888 SH   DFND 16 0 14,721 167
ADDUS HOMECARE CORP COM 006739106 31,225 330,386 SH   DFND 2 0 329,111 1,275
ADDUS HOMECARE CORP COM 006739106 390 4,120 SH   DFND 16 0 3,786 334
ADOBE INC COM 00724F101 1,338,666 2,729,583 SH   DFND 2 0 2,438,288 291,295
ADOBE INC COM 00724F101 34,846 71,050 SH   DFND 2,3 0 71,050 0
ADOBE INC COM 00724F101 28,411 57,930 SH   DFND 2,10 0 57,930 0
ADOBE INC COM 00724F101 466,081 951,880 SH   DFND 10 0 951,880 0
ADOBE INC COM 00724F101 113,406 230,905 SH   DFND 16 0 79,563 151,342
ADOBE INC COM 00724F101 52 107 SH   DFND 16,18 0 107 0
ADOBE INC COM 00724F101 173 353 SH   DFND 18 0 321 32
ADTALEM GLOBAL EDUCATION INC COM 00737L103 2,964 120,774 SH   DFND 2 0 118,474 2,300
ADTALEM GLOBAL EDUCATION INC COM 00737L103 0 8 SH   DFND 16 0 0 8
ADTRAN INC COM 00738A106 2,807 273,707 SH   DFND 2 0 270,407 3,300
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ADURO BIOTECH INC COM 00739L101 55 22,761 SH   DFND 2 0 22,761 0
ADURO BIOTECH INC COM 00739L101 1 400 SH   DFND 16 0 0 400
ADVANCE AUTO PARTS INC COM 00751Y106 6,815 44,390 SH   DFND 2 0 44,390 0
ADVANCE AUTO PARTS INC COM 00751Y106 37 242 SH   DFND 2,3 0 242 0
ADVANCE AUTO PARTS INC COM 00751Y106 345 2,245 SH   DFND 16 0 0 2,245
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AECOM COM 00766T100 14,093 336,827 SH   DFND 2 0 332,012 4,815
AECOM COM 00766T100 1,069 25,550 SH   DFND 16 0 0 25,550
AEGERION PHARMACEUTICALS INC CONV BND 00767EAC6 5,026 4,205,425 PRN   DFND 2 0 0 4,205,425
AEGION CORP COM 00770F104 4,154 294,002 SH   DFND 2 0 287,502 6,500
AEGION CORP COM 00770F104 7 500 SH   DFND 16 0 0 500
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AERIE PHARMACEUTICALS INC COM 00771V108 3,978 337,978 SH   DFND 2,10 0 337,978 0
AERIE PHARMACEUTICALS INC COM 00771V108 755 64,136 SH   DFND 16 0 64,136 0
AERIE PHARMACEUTICALS INC CONV BND 00771VAB4 10,066 12,500,000 PRN   DFND 2 0 0 12,500,000
ADVANSIX INC COM 00773T101 2,276 176,684 SH   DFND 2 0 176,684 0
ADVANSIX INC COM 00773T101 8 610 SH   DFND 16 0 0 610
ADVANSIX INC COM 00773T101 0 26 SH   DFND 18 0 26 0
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AEROJET ROCKETDYNE HOLDINGS COM 007800105 242 6,081 SH   DFND 16 0 5,422 659
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ADVANCED MICRO DEVICES COM 007903107 332 4,048 SH   DFND 2,3 0 4,048 0
ADVANCED MICRO DEVICES COM 007903107 35,921 438,303 SH   DFND 16 0 131,633 306,670
ADVANCED MICRO DEVICES COM 007903107 703 8,580 SH   DFND 18 0 8,580 0
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ADVANCED DRAINAGE SYSTEMS IN COM 00790R104 54,630 875,049 SH   DFND 10 0 634,474 240,575
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ADVANCED DRAINAGE SYSTEMS IN COM 00790R104 4 62 SH   DFND 18 0 0 62
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AEGON N.V.-NY REG SHR COM 007924103 259 100,948 SH   DFND 16 0 0 100,948
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ADVANCED ENERGY INDUSTRIES COM 007973100 64 1,012 SH   DFND 16 0 964 48
ADVANCED ENERGY INDUSTRIES COM 007973100 9,586 152,308 SH   DFND 17 0 152,308 0
ADVANCED ENERGY INDUSTRIES COM 007973100 1 17 SH   DFND 18 0 0 17
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AEROVIRONMENT INC COM 008073108 45 750 SH   DFND 16 0 0 750
AERPIO PHARMACEUTICALS INC COM 00810B105 296 197,114 SH   DFND 16 0 0 197,114
AFFILIATED MANAGERS GROUP COM 008252108 6,439 94,164 SH   DFND 2 0 92,764 1,400
AFFILIATED MANAGERS GROUP COM 008252108 149 2,174 SH   DFND 16 0 0 2,174
AGILENT TECHNOLOGIES INC COM 00846U101 65,778 651,654 SH   DFND 2 0 648,574 3,080
AGILENT TECHNOLOGIES INC COM 00846U101 108 1,065 SH   DFND 2,3 0 1,065 0
AGILENT TECHNOLOGIES INC COM 00846U101 4,450 44,099 SH   DFND 16 0 30,015 14,084
AGILENT TECHNOLOGIES INC COM 00846U101 52 518 SH   DFND 18 0 518 0
AGNICO EAGLE MINES LTD COM 008474108 5 58 SH   DFND 2 0 58 0
AGNICO EAGLE MINES LTD COM 008474108 955 11,998 SH   DFND 16 0 0 11,998
AGENUS INC COM 00847G705 0 5 SH   DFND 2 0 5 0
AGENUS INC COM 00847G705 49 12,248 SH   DFND 16 0 0 12,248
AGILYSYS INC COM 00847J105 2,142 88,680 SH   DFND 2 0 88,680 0
AGILE THERAPEUTICS INC COM 00847L100 191 62,690 SH   DFND 16 0 0 62,690
AGIOS PHARMACEUTICALS INC COM 00847X104 2,324 66,398 SH   DFND 2 0 66,398 0
AGIOS PHARMACEUTICALS INC COM 00847X104 1 31 SH   DFND 16 0 0 31
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AGREE REALTY CORP COM 008492100 951 14,946 SH   DFND 16 0 13,588 1,358
AGORA INC-ADR ADR 00851L103 717 16,736 SH   DFND 10 0 16,736 0
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AIMMUNE THERAPEUTICS INC COM 00900T107 23 667 SH   DFND 16 0 0 667
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AIR LEASE CORP COM 00912X302 964 32,765 SH   DFND 16 0 29,580 3,185
AIR PRODUCTS & CHEMICALS INC COM 009158106 161,461 541,837 SH   DFND 2 0 521,068 20,769
AIR PRODUCTS & CHEMICALS INC COM 009158106 4,364 14,650 SH   DFND 2,3 0 14,650 0
AIR PRODUCTS & CHEMICALS INC COM 009158106 21,354 71,734 SH   DFND 16 0 14,274 57,460
AIR PRODUCTS & CHEMICALS INC COM 009158106 59 197 SH   DFND 16,18 0 197 0
AIR PRODUCTS & CHEMICALS INC COM 009158106 77 257 SH   DFND 18 0 257 0
AIR TRANSPORT SERVICES GROUP COM 00922R105 15,141 604,203 SH   DFND 2 0 563,703 40,500
AIR TRANSPORT SERVICES GROUP COM 00922R105 426 17,000 SH   DFND 2,3 0 17,000 0
AIR TRANSPORT SERVICES GROUP COM 00922R105 43 1,703 SH   DFND 16 0 0 1,703
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AIRGAIN INC COM 00938A104 16 1,200 SH   DFND 16 0 0 1,200
AKAMAI TECHNOLOGIES INC COM 00971T101 8,776 79,399 SH   DFND 2 0 79,399 0
AKAMAI TECHNOLOGIES INC COM 00971T101 62 565 SH   DFND 2,3 0 565 0
AKAMAI TECHNOLOGIES INC COM 00971T101 2,576 23,305 SH   DFND 16 0 1,567 21,738
AKAMAI TECHNOLOGIES INC COM 00971T101 20 179 SH   DFND 16,18 0 179 0
AKAMAI TECHNOLOGIES INC COM 00971T101 10 87 SH   DFND 18 0 50 37
AKAMAI TECHNOLOGIES INC CONV BND 00971TAJ0 15 12,000 PRN   DFND 16 0 0 12,000
AKAMAI TECHNOLOGIES INC CONV BND 00971TAL5 24,295 21,000,000 PRN   DFND 2 0 0 21,000,000
AKEBIA THERAPEUTICS INC COM 00972D105 43 17,058 SH   DFND 16 0 0 17,058
ALAMO GROUP INC COM 011311107 4,724 43,734 SH   DFND 2 0 42,984 750
ALAMO GROUP INC COM 011311107 35 323 SH   DFND 16 0 0 323
ALARM.COM HOLDINGS INC COM 011642105 28,607 517,764 SH   DFND 2 0 513,664 4,100
ALARM.COM HOLDINGS INC COM 011642105 101 1,818 SH   DFND 16 0 717 1,101
ALASKA AIR GROUP INC COM 011659109 22,113 603,685 SH   DFND 2 0 603,685 0
ALASKA AIR GROUP INC COM 011659109 16 430 SH   DFND 2,3 0 430 0
ALASKA AIR GROUP INC COM 011659109 5,766 158,090 SH   DFND 16 0 0 158,090
ALASKA AIR GROUP INC COM 011659109 1 21 SH   DFND 18 0 21 0
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ALBANY INTL CORP-CL A COM 012348108 151 3,056 SH   DFND 16 0 1,620 1,436
ALBEMARLE CORP COM 012653101 30,827 345,277 SH   DFND 2 0 345,272 5
ALBEMARLE CORP COM 012653101 33 365 SH   DFND 2,3 0 365 0
ALBEMARLE CORP COM 012653101 5,045 56,584 SH   DFND 16 0 7,355 49,229
ALBEMARLE CORP COM 012653101 71 800 SH   DFND 18 0 800 0
ALBERTSONS COS INC - CLASS A COM 013091103 168 12,136 SH   DFND 16 0 0 12,136
ALCOA CORP COM 013872106 458 39,412 SH   DFND 2 0 39,412 0
ALCOA CORP COM 013872106 1,785 153,501 SH   DFND 16 0 11,933 141,568
ALCOA CORP COM 013872106 0 2 SH   DFND 18 0 2 0
ALDEYRA THERAPEUTICS INC COM 01438T106 1,754 236,674 SH   DFND 2 0 236,674 0
ALECTOR INC COM 014442107 396 37,574 SH   DFND 2 0 37,574 0
ALEXANDER & BALDWIN INC COM 014491104 5,243 467,763 SH   DFND 2 0 458,263 9,500
ALEXANDER & BALDWIN INC COM 014491104 2 137 SH   DFND 16 0 0 137
ALEXANDER'S INC COM 014752109 594 2,420 SH   DFND 2 0 2,420 0
ALEXANDRIA REAL ESTATE EQUIT COM 015271109 142,707 891,808 SH   DFND 2 0 889,560 2,248
ALEXANDRIA REAL ESTATE EQUIT COM 015271109 65 408 SH   DFND 2,3 0 408 0
ALEXANDRIA REAL ESTATE EQUIT COM 015271109 87 547 SH   DFND 10 0 547 0
ALEXANDRIA REAL ESTATE EQUIT COM 015271109 6,157 38,470 SH   DFND 16 0 19,252 19,218
ALEXION PHARMACEUTICALS INC COM 015351109 447,349 3,909,376 SH   DFND 2 0 3,378,746 530,630
ALEXION PHARMACEUTICALS INC COM 015351109 17,723 154,882 SH   DFND 2,3 0 154,882 0
ALEXION PHARMACEUTICALS INC COM 015351109 8,441 73,766 SH   DFND 2,10 0 73,766 0
ALEXION PHARMACEUTICALS INC COM 015351109 99,332 868,967 SH   DFND 10 0 868,967 0
ALEXION PHARMACEUTICALS INC COM 015351109 10,748 93,759 SH   DFND 16 0 21,556 72,203
ALEXION PHARMACEUTICALS INC COM 015351109 15 136 SH   DFND 18 0 109 27
ALGONQUIN POWER & UTILITIES COM 015857105 2,685 184,527 SH   DFND 16 0 182,112 2,415
ALIBABA GROUP HOLDING-SP ADR ADR 01609W102 1,086,098 3,694,463 SH   DFND 2 0 3,395,470 298,993
ALIBABA GROUP HOLDING-SP ADR ADR 01609W102 28,004 95,256 SH   DFND 2,3 0 95,256 0
ALIBABA GROUP HOLDING-SP ADR ADR 01609W102 37,649 128,066 SH   DFND 2,10 0 128,066 0
ALIBABA GROUP HOLDING-SP ADR ADR 01609W102 438,620 1,492,663 SH   DFND 10 0 1,492,663 0
ALIBABA GROUP HOLDING-SP ADR ADR 01609W102 171,026 575,588 SH   DFND 16 0 96,462 479,126
ALIBABA GROUP HOLDING-SP ADR ADR 01609W102 61 209 SH   DFND 16,18 0 209 0
ALIBABA GROUP HOLDING-SP ADR ADR 01609W102 247 840 SH   DFND 18 0 821 19
ALIGN TECHNOLOGY INC COM 016255101 258,704 790,270 SH   DFND 2 0 690,625 99,645
ALIGN TECHNOLOGY INC COM 016255101 15,328 46,822 SH   DFND 2,3 0 46,822 0
ALIGN TECHNOLOGY INC COM 016255101 6,681 20,427 SH   DFND 16 0 12,306 8,121
ALIGN TECHNOLOGY INC COM 016255101 12 38 SH   DFND 16,18 0 38 0
ALIGN TECHNOLOGY INC COM 016255101 18 56 SH   DFND 18 0 47 9
ALLEGHANY CORP COM 017175100 27,711 53,245 SH   DFND 2 0 52,878 367
ALLEGHANY CORP COM 017175100 1,297 2,493 SH   DFND 16 0 2,332 161
ALLEGHANY CORP COM 017175100 39 74 SH   DFND 18 0 74 0
ALLEGHENY TECHNOLOGIES INC COM 01741R102 5,060 580,281 SH   DFND 2 0 562,181 18,100
ALLEGHENY TECHNOLOGIES INC COM 01741R102 129 14,828 SH   DFND 16 0 0 14,828
ALLEGIANCE BANCSHARES INC COM 01748H107 2,141 91,610 SH   DFND 2 0 87,110 4,500
ALLEGIANT TRAVEL CO COM 01748X102 7,011 58,518 SH   DFND 2 0 57,333 1,185
ALLEGIANT TRAVEL CO COM 01748X102 4 34 SH   DFND 16 0 0 34
ALLETE INC COM 018522300 7,741 149,633 SH   DFND 2 0 149,633 0
ALLETE INC COM 018522300 269 5,204 SH   DFND 16 0 0 5,204
ALLETE INC COM 018522300 2 31 SH   DFND 18 0 31 0
ALLIANCE DATA SYSTEMS CORP COM 018581108 5,228 124,584 SH   DFND 2 0 122,834 1,750
ALLIANCE DATA SYSTEMS CORP COM 018581108 52 1,292 SH   DFND 16 0 0 1,292
ALLIANCE NATIONAL MUNI INC COM 01864U106 1,259 89,398 SH   DFND 16 0 0 89,398
ALLIANCEBERNSTEIN GL HI INC COM 01879R106 443 42,165 SH   DFND 16 0 0 42,165
ALLIANCEBERNSTEIN GL HI INC COM 01879R106 15 1,456 SH   DFND 18 0 1,456 0
ALLIANT ENERGY CORP COM 018802108 12,798 247,749 SH   DFND 2 0 247,739 10
ALLIANT ENERGY CORP COM 018802108 44 859 SH   DFND 2,3 0 859 0
ALLIANT ENERGY CORP COM 018802108 647 12,522 SH   DFND 16 0 0 12,522
ALLIANT ENERGY CORP COM 018802108 30 578 SH   DFND 18 0 578 0
ALLIANCEBERNSTEIN HOLDING LP LP 01881G106 1,181 43,660 SH   DFND 16 0 0 43,660
ALLIANZGI CONV & INCOME II COM 018825109 12,666 3,081,831 SH   DFND 16 0 0 3,081,831
ALLIANZGI CONVERTIBLE & INCO COM 018828103 2,249 490,984 SH   DFND 16 0 0 490,984
ALLIANZGI EQUITY & CONVERTIB COM 018829101 832 34,283 SH   DFND 16 0 0 34,283
ALLIANZGI CONVERTIBLE & INCO COM 01883H102 739 83,970 SH   DFND 16 0 0 83,970
ALLIANZGI ARTIFICIAL INTEL COM 01883M101 1,516 70,267 SH   DFND 16 0 0 70,267
ALLOGENE THERAPEUTICS INC COM 019770106 1,137 30,161 SH   DFND 2 0 30,161 0
ALLOGENE THERAPEUTICS INC COM 019770106 22 582 SH   DFND 16 0 515 67
ALLSCRIPTS HEALTHCARE SOLUTI COM 01988P108 8,116 997,052 SH   DFND 2 0 982,052 15,000
ALLSCRIPTS HEALTHCARE SOLUTI COM 01988P108 28 3,482 SH   DFND 16 0 0 3,482
ALLSTATE CORP COM 020002101 799,063 8,487,585 SH   DFND 2 0 7,511,254 976,331
ALLSTATE CORP COM 020002101 2,767 29,386 SH   DFND 2,3 0 29,386 0
ALLSTATE CORP COM 020002101 109,406 1,163,126 SH   DFND 10 0 1,163,126 0
ALLSTATE CORP COM 020002101 46,345 495,787 SH   DFND 16 0 27,022 468,765
ALLSTATE CORP COM 020002101 80 848 SH   DFND 16,18 0 848 0
ALLSTATE CORP COM 020002101 37 393 SH   DFND 18 0 348 45
ALLY FINANCIAL INC COM 02005N100 2,867 114,377 SH   DFND 2 0 114,377 0
ALLY FINANCIAL INC COM 02005N100 1,263 50,398 SH   DFND 16 0 42,282 8,116
ALNYLAM PHARMACEUTICALS INC COM 02043Q107 740 5,087 SH   DFND 2 0 5,078 9
ALNYLAM PHARMACEUTICALS INC COM 02043Q107 3,276 22,482 SH   DFND 16 0 20,403 2,079
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ALPHABET INC-CL C COM 02079K107 15,105 10,278 SH   DFND 2,3 0 0 10,278
ALPHABET INC-CL C COM 02079K107 5,252 3,578 SH   DFND 10 0 0 3,578
ALPHABET INC-CL C COM 02079K107 103,593 70,519 SH   DFND 16 0 0 70,519
ALPHABET INC-CL C COM 02079K107 917 624 SH   DFND 16,18 0 0 624
ALPHABET INC-CL C COM 02079K107 736 501 SH   DFND 18 0 0 501
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ALPHABET INC-CL A COM 02079K305 15,738 10,739 SH   DFND 2,3 0 10,739 0
ALPHABET INC-CL A COM 02079K305 26,966 18,399 SH   DFND 2,10 0 18,399 0
ALPHABET INC-CL A COM 02079K305 798,975 545,510 SH   DFND 10 0 545,510 0
ALPHABET INC-CL A COM 02079K305 327,934 224,070 SH   DFND 16 0 36,385 187,685
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ALPHABET INC-CL A COM 02079K305 1,307 892 SH   DFND 18 0 883 9
ALTABANCORP COM 021347109 5,390 267,890 SH   DFND 2 0 267,890 0
ALTAIR ENGINEERING INC - A COM 021369103 5,409 128,847 SH   DFND 2 0 128,847 0
ALTAIR ENGINEERING INC - A COM 021369103 54,519 1,298,723 SH   DFND 10 0 981,880 316,843
ALTAIR ENGINEERING INC - A COM 021369103 219 5,213 SH   DFND 16 0 5,142 71
ALTERYX INC - CLASS A COM 02156B103 61,773 544,012 SH   DFND 2 0 544,012 0
ALTERYX INC - CLASS A COM 02156B103 637 5,610 SH   DFND 16 0 2,014 3,595
ALTERYX INC - CLASS A COM 02156B103 43,791 385,655 SH   DFND 17 0 385,655 0
ALTERYX INC CONV BND 02156BAD5 14,021 14,000,000 PRN   DFND 2 0 0 14,000,000
ALTICE USA INC- A COM 02156K103 5,373 206,653 SH   DFND 2 0 206,653 0
ALTICE USA INC- A COM 02156K103 565 21,752 SH   DFND 16 0 19,314 2,438
ALTRA INDUSTRIAL MOTION CORP COM 02208R106 13,202 357,115 SH   DFND 2 0 327,615 29,500
ALTRA INDUSTRIAL MOTION CORP COM 02208R106 444 12,000 SH   DFND 2,3 0 12,000 0
ALTRA INDUSTRIAL MOTION CORP COM 02208R106 353 9,541 SH   DFND 16 0 5,917 3,624
ALTRIA GROUP INC COM 02209S103 347,386 8,984,999 SH   DFND 2 0 7,287,727 1,697,272
ALTRIA GROUP INC COM 02209S103 247 6,400 SH   DFND 2,3 0 6,400 0
ALTRIA GROUP INC COM 02209S103 76,654 1,994,927 SH   DFND 16 0 59,686 1,935,241
ALTRIA GROUP INC COM 02209S103 324 8,408 SH   DFND 16,18 0 8,408 0
ALTRIA GROUP INC COM 02209S103 482 12,255 SH   DFND 18 0 12,034 221
AMALGAMATED BK OF NEW YORK-A COM 022663108 619 58,567 SH   DFND 2 0 58,567 0
AMARIN CORP PLC -ADR ADR 023111206 1,599 379,335 SH   DFND 2 0 379,335 0
AMARIN CORP PLC -ADR ADR 023111206 1,349 320,343 SH   DFND 2,10 0 320,343 0
AMARIN CORP PLC -ADR ADR 023111206 3,143 749,743 SH   DFND 16 0 0 749,743
AMAZON.COM INC COM 023135106 4,207,782 1,336,342 SH   DFND 2 0 1,251,524 84,818
AMAZON.COM INC COM 023135106 72,391 22,991 SH   DFND 2,3 0 22,991 0
AMAZON.COM INC COM 023135106 45,383 14,413 SH   DFND 2,10 0 14,413 0
AMAZON.COM INC COM 023135106 1,146,307 364,318 SH   DFND 10 0 364,318 0
AMAZON.COM INC COM 023135106 566,388 179,815 SH   DFND 16 0 35,310 144,506
AMAZON.COM INC COM 023135106 1,105 351 SH   DFND 16,18 0 351 0
AMAZON.COM INC COM 023135106 3,876 1,231 SH   DFND 18 0 1,222 9
AMBAC FINANCIAL GROUP INC COM 023139884 3,184 249,280 SH   DFND 2 0 249,280 0
AMBEV SA-ADR ADR 02319V103 1 614 SH   DFND 2 0 614 0
AMBEV SA-ADR ADR 02319V103 768 339,894 SH   DFND 16 0 0 339,894
AMEDISYS INC COM 023436108 62,186 263,023 SH   DFND 2 0 261,813 1,210
AMEDISYS INC COM 023436108 61 256 SH   DFND 2,3 0 256 0
AMEDISYS INC COM 023436108 2,562 10,836 SH   DFND 16 0 6,759 4,077
AMEDISYS INC COM 023436108 59,401 251,241 SH   DFND 17 0 251,241 0
AMERCO COM 023586100 1,792 5,034 SH   DFND 2 0 5,034 0
AMERCO COM 023586100 241 678 SH   DFND 16 0 0 678
AMERCO COM 023586100 6 16 SH   DFND 18 0 0 16
AMEREN CORPORATION COM 023608102 341,632 4,320,039 SH   DFND 2 0 4,139,051 180,988
AMEREN CORPORATION COM 023608102 1,476 18,662 SH   DFND 2,3 0 18,662 0
AMEREN CORPORATION COM 023608102 2,729 34,045 SH   DFND 16 0 19,807 14,238
AMEREN CORPORATION COM 023608102 11 134 SH   DFND 18 0 134 0
AMERICA MOVIL-SPN ADR CL L ADR 02364W105 61 4,850 SH   DFND 2 0 0 4,850
AMERICA MOVIL-SPN ADR CL L ADR 02364W105 9,126 731,230 SH   DFND 10 0 0 731,230
AMERICA MOVIL-SPN ADR CL L ADR 02364W105 196 15,765 SH   DFND 16 0 0 15,765
AMERICAN AIRLINES GROUP INC COM 02376R102 1,564 127,213 SH   DFND 2 0 127,213 0
AMERICAN AIRLINES GROUP INC COM 02376R102 21 1,749 SH   DFND 2,3 0 1,749 0
AMERICAN AIRLINES GROUP INC COM 02376R102 1,249 101,849 SH   DFND 16 0 32 101,817
AMERICAN AIRLINES GROUP INC COM 02376R102 1 89 SH   DFND 18 0 89 0
AMERICAN AIRLINES GROUP INC CONV BND 02376RAF9 171 185,000 PRN   DFND 2 0 0 185,000
AMERICAN ASSETS TRUST INC COM 024013104 25,366 1,052,994 SH   DFND 2 0 1,011,594 41,400
AMERICAN ASSETS TRUST INC COM 024013104 349 14,500 SH   DFND 2,3 0 14,500 0
AMERICAN ASSETS TRUST INC COM 024013104 204 8,461 SH   DFND 16 0 8,289 172
AMERICAN ASSETS TRUST INC COM 024013104 0 8 SH   DFND 18 0 8 0
AMERICAN AXLE & MFG HOLDINGS COM 024061103 5,378 931,934 SH   DFND 2 0 921,934 10,000
AMERICAN AXLE & MFG HOLDINGS COM 024061103 27 4,600 SH   DFND 16 0 0 4,600
AMERICAN CAMPUS COMMUNITIES COM 024835100 9,643 276,151 SH   DFND 2 0 271,651 4,500
AMERICAN CAMPUS COMMUNITIES COM 024835100 313 8,986 SH   DFND 16 0 7,286 1,700
AMERICAN CAMPUS COMMUNITIES COM 024835100 2 60 SH   DFND 18 0 0 60
AMERICAN ELECTRIC POWER COM 025537101 247,909 3,033,272 SH   DFND 2 0 2,847,803 185,469
AMERICAN ELECTRIC POWER COM 025537101 1,564 19,139 SH   DFND 2,3 0 19,139 0
AMERICAN ELECTRIC POWER COM 025537101 89,480 1,094,823 SH   DFND 10 0 1,094,823 0
AMERICAN ELECTRIC POWER COM 025537101 19,714 241,309 SH   DFND 16 0 74,695 166,615
AMERICAN ELECTRIC POWER COM 025537101 42 513 SH   DFND 16,18 0 513 0
AMERICAN ELECTRIC POWER COM 025537101 129 1,582 SH   DFND 18 0 1,516 66
AEP 6 1/8 03/15/22 CONV PRF 025537127 376 7,750 SH   DFND 2 0 0 7,750
AMERICAN EAGLE OUTFITTERS COM 02553E106 11,793 796,276 SH   DFND 2 0 756,276 40,000
AMERICAN EAGLE OUTFITTERS COM 02553E106 241 16,300 SH   DFND 2,3 0 16,300 0
AMERICAN EAGLE OUTFITTERS COM 02553E106 356 24,062 SH   DFND 16 0 0 24,062
AMERICAN EQUITY INVT LIFE HL COM 025676206 20,354 925,622 SH   DFND 2 0 915,922 9,700
AMERICAN EQUITY INVT LIFE HL COM 025676206 68 3,080 SH   DFND 16 0 2,396 684
AMERICAN EXPRESS CO COM 025816109 217,774 2,172,310 SH   DFND 2 0 2,089,253 83,057
AMERICAN EXPRESS CO COM 025816109 6,767 67,508 SH   DFND 2,3 0 67,508 0
AMERICAN EXPRESS CO COM 025816109 37,840 377,650 SH   DFND 16 0 165,926 211,724
AMERICAN EXPRESS CO COM 025816109 52 520 SH   DFND 16,18 0 520 0
AMERICAN EXPRESS CO COM 025816109 141 1,410 SH   DFND 18 0 1,377 33
AMERICAN FINANCIAL GROUP INC COM 025932104 21,600 322,483 SH   DFND 2 0 319,068 3,415
AMERICAN FINANCIAL GROUP INC COM 025932104 93 1,382 SH   DFND 16 0 0 1,382
AMERICAN FINANCIAL GROUP INC COM 025932104 3 51 SH   DFND 18 0 0 51
AMERICAN HOMES 4 RENT- A COM 02665T306 62,839 2,206,337 SH   DFND 2 0 2,177,626 28,711
AMERICAN HOMES 4 RENT- A COM 02665T306 1,307 45,819 SH   DFND 16 0 40,170 5,649
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ARTISAN PARTNERS ASSET MA -A COM 04316A108 2,257 57,875 SH   DFND 2 0 57,875 0
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AIZ 6 1/2 03/15/21 CONV PRF 04621X207 23,026 195,000 SH   DFND 2 0 0 195,000
ASTEC INDUSTRIES INC COM 046224101 11,611 214,034 SH   DFND 2 0 210,734 3,300
ASTEC INDUSTRIES INC COM 046224101 32 587 SH   DFND 16 0 0 587
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ASURE SOFTWARE INC COM 04649U102 5,102 675,882 SH   DFND 2 0 675,882 0
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CAE INC COM 124765108 2,796 191,356 SH   DFND 16 0 190,965 391
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CRH PLC-SPONSORED ADR ADR 12626K203 2,113 58,524 SH   DFND 16 0 0 58,524
CRH PLC-SPONSORED ADR ADR 12626K203 3 76 SH   DFND 18 0 0 76
CSG SYSTEMS INTL INC COM 126349109 7,755 189,370 SH   DFND 2 0 184,820 4,550
CSG SYSTEMS INTL INC COM 126349109 361 8,787 SH   DFND 16 0 6,688 2,099
CSG SYSTEMS INTL INC COM 126349109 1 27 SH   DFND 18 0 27 0
CSW INDUSTRIALS INC COM 126402106 1,757 22,732 SH   DFND 2 0 22,732 0
CSW INDUSTRIALS INC COM 126402106 102 1,325 SH   DFND 16 0 1,325 0
CSX CORP COM 126408103 271,788 3,499,271 SH   DFND 2 0 2,447,540 1,051,731
CSX CORP COM 126408103 205 2,638 SH   DFND 2,3 0 2,638 0
CSX CORP COM 126408103 16,899 217,678 SH   DFND 16 0 12,204 205,474
CSX CORP COM 126408103 5 60 SH   DFND 16,18 0 60 0
CSX CORP COM 126408103 276 3,560 SH   DFND 18 0 3,504 56
CTS CORP COM 126501105 3,732 169,369 SH   DFND 2 0 165,669 3,700
CTS CORP COM 126501105 103 4,677 SH   DFND 16 0 4,572 105
CNX RESOURCES CORP COM 12653C108 5,041 533,867 SH   DFND 2 0 526,467 7,400
CNX RESOURCES CORP COM 12653C108 28 2,974 SH   DFND 16 0 0 2,974
CVB FINANCIAL CORP COM 126600105 9,461 568,904 SH   DFND 2 0 562,404 6,500
CVS HEALTH CORP COM 126650100 59,824 1,024,390 SH   DFND 2 0 992,670 31,720
CVS HEALTH CORP COM 126650100 263 4,511 SH   DFND 2,3 0 4,511 0
CVS HEALTH CORP COM 126650100 184,353 3,188,043 SH   DFND 16 0 29,537 3,158,505
CVS HEALTH CORP COM 126650100 353 6,047 SH   DFND 16,18 0 6,047 0
CVS HEALTH CORP COM 126650100 534 9,166 SH   DFND 18 0 9,090 76
CABLE ONE INC COM 12685J105 36,535 19,377 SH   DFND 2 0 19,156 221
CABLE ONE INC COM 12685J105 307 163 SH   DFND 16 0 61 102
CABOT CORP COM 127055101 4,657 129,256 SH   DFND 2 0 126,031 3,225
CABOT CORP COM 127055101 318 8,828 SH   DFND 16 0 8,144 684
CABOT OIL & GAS CORP COM 127097103 4,752 273,748 SH   DFND 2 0 273,728 20
CABOT OIL & GAS CORP COM 127097103 24 1,371 SH   DFND 2,3 0 1,371 0
CABOT OIL & GAS CORP COM 127097103 1,537 88,451 SH   DFND 16 0 0 88,451
CABOT OIL & GAS CORP COM 127097103 3 146 SH   DFND 18 0 146 0
CACI INTERNATIONAL INC -CL A COM 127190304 46,081 216,178 SH   DFND 2 0 214,345 1,833
CACI INTERNATIONAL INC -CL A COM 127190304 1,059 4,971 SH   DFND 16 0 2,768 2,203
CACI INTERNATIONAL INC -CL A COM 127190304 0 2 SH   DFND 18 0 2 0
CACTUS INC - A COM 127203107 386 20,110 SH   DFND 2 0 20,110 0
CACTUS INC - A COM 127203107 0 23 SH   DFND 16 0 0 23
CACTUS INC - A COM 127203107 2 81 SH   DFND 18 0 0 81
CADENCE DESIGN SYS INC COM 127387108 106,903 1,002,540 SH   DFND 2 0 994,222 8,318
CADENCE DESIGN SYS INC COM 127387108 102 955 SH   DFND 2,3 0 955 0
CADENCE DESIGN SYS INC COM 127387108 16,267 152,648 SH   DFND 10 0 152,648 0
CADENCE DESIGN SYS INC COM 127387108 12,530 117,506 SH   DFND 16 0 35,908 81,597
CADENCE DESIGN SYS INC COM 127387108 39,625 371,612 SH   DFND 17 0 371,612 0
CADENCE DESIGN SYS INC COM 127387108 6 55 SH   DFND 18 0 55 0
CADENCE BANCORP COM 12739A100 8,691 1,011,770 SH   DFND 2 0 1,001,870 9,900
CADENCE BANCORP COM 12739A100 1 120 SH   DFND 16 0 0 120
CAESARS ENTERTAINMENT INC COM 12769G100 103,290 1,842,488 SH   DFND 2 0 1,814,378 28,110
CAESARS ENTERTAINMENT INC COM 12769G100 693 12,377 SH   DFND 2,3 0 12,377 0
CAESARS ENTERTAINMENT INC COM 12769G100 945 16,867 SH   DFND 16 0 16,348 519
CAL-MAINE FOODS INC COM 128030202 6,267 163,297 SH   DFND 2 0 161,197 2,100
CAL-MAINE FOODS INC COM 128030202 1,989 51,834 SH   DFND 16 0 51,390 444
CALAMOS CONVERTIBLE OPP&INC COM 128117108 3,725 338,596 SH   DFND 16 0 0 338,596
CALAMOS CONVERTIBLE OPP&INC COM 128117108 54 4,887 SH   DFND 18 0 4,887 0
CALAMOS GLOBAL DYNAMIC INCOM COM 12811L107 2,079 255,733 SH   DFND 16 0 0 255,733
CALAMOS CVT AND HIGH INCOME COM 12811P108 2,562 219,308 SH   DFND 16 0 0 219,308
CALAMOS CVT AND HIGH INCOME COM 12811P108 45 3,860 SH   DFND 18 0 3,860 0
CALAMOS DYNAMIC CONVERTIBLE COM 12811V105 3,131 136,863 SH   DFND 16 0 0 136,863
CALAMOS DYNAMIC CONVERTIBLE COM 12811V105 133 5,810 SH   DFND 18 0 5,810 0
CALAMOS STRAT TOT RETURN FD COM 128125101 3,608 268,069 SH   DFND 16 0 0 268,069
CALAMOS STRAT TOT RETURN FD COM 128125101 19 1,387 SH   DFND 18 0 1,387 0
CALAMP CORP COM 128126109 1,087 151,145 SH   DFND 2 0 151,145 0
CALAMP CORP COM 128126109 6 900 SH   DFND 16 0 0 900
CALAVO GROWERS INC COM 128246105 4,876 73,574 SH   DFND 2 0 72,774 800
CALAVO GROWERS INC COM 128246105 2,750 41,462 SH   DFND 16 0 41,074 388
CALERES INC COM 129500104 1,699 177,664 SH   DFND 2 0 173,364 4,300
CALERES INC COM 129500104 229 23,981 SH   DFND 16 0 23,348 633
CALIFORNIA WATER SERVICE GRP COM 130788102 12,777 294,077 SH   DFND 2 0 289,777 4,300
CALIFORNIA WATER SERVICE GRP COM 130788102 719 16,538 SH   DFND 16 0 9,645 6,893
CALLAWAY GOLF COMPANY COM 131193104 8,227 429,855 SH   DFND 2 0 423,355 6,500
CALLAWAY GOLF COMPANY COM 131193104 103 5,406 SH   DFND 16 0 0 5,406
CALLON PETROLEUM CO COM 13123X508 836 173,553 SH   DFND 2 0 173,553 0
CALLON PETROLEUM CO COM 13123X508 1 160 SH   DFND 16 0 0 160
CALYXT INC COM 13173L107 279 50,765 SH   DFND 2 0 50,765 0
CALYXT INC COM 13173L107 1 125 SH   DFND 16 0 0 125
CAMDEN NATIONAL CORP COM 133034108 308 10,171 SH   DFND 2 0 10,171 0
CAMDEN NATIONAL CORP COM 133034108 188 6,213 SH   DFND 16 0 0 6,213
CAMDEN PROPERTY TRUST COM 133131102 25,526 286,802 SH   DFND 2 0 283,852 2,950
CAMDEN PROPERTY TRUST COM 133131102 493 5,536 SH   DFND 16 0 4,597 939
CAMDEN PROPERTY TRUST COM 133131102 3 42 SH   DFND 18 0 4 38
CAMECO CORP COM 13321L108 34,152 3,381,478 SH   DFND 2 0 3,381,478 0
CAMECO CORP COM 13321L108 26,394 2,613,283 SH   DFND 2,10 0 2,613,283 0
CAMECO CORP COM 13321L108 1,556 154,122 SH   DFND 16 0 0 154,122
CAMPBELL SOUP CO COM 134429109 3,905 80,765 SH   DFND 2 0 74,265 6,500
CAMPBELL SOUP CO COM 134429109 34 697 SH   DFND 2,3 0 697 0
CAMPBELL SOUP CO COM 134429109 2,933 60,646 SH   DFND 16 0 0 60,646
CAMPBELL SOUP CO COM 134429109 5 113 SH   DFND 18 0 113 0
CAMPING WORLD HOLDINGS INC-A COM 13462K109 215 7,240 SH   DFND 2 0 7,240 0
CAMPING WORLD HOLDINGS INC-A COM 13462K109 901 30,285 SH   DFND 16 0 0 30,285
CANADA GOOSE HOLDINGS INC COM 135086106 7,581 235,633 SH   DFND 2 0 235,633 0
CANADA GOOSE HOLDINGS INC COM 135086106 12,923 401,719 SH   DFND 10 0 401,719 0
CANADA GOOSE HOLDINGS INC COM 135086106 563 17,508 SH   DFND 16 0 14,925 2,583
CANADA GOOSE HOLDINGS INC COM 135086106 13 400 SH   DFND 16,18 0 400 0
CAN IMPERIAL BK OF COMMERCE COM 136069101 18 238 SH   DFND 2 0 238 0
CAN IMPERIAL BK OF COMMERCE COM 136069101 16,982 227,305 SH   DFND 16 0 220,902 6,403
CAN IMPERIAL BK OF COMMERCE COM 136069101 15 201 SH   DFND 18 0 0 201
CANADIAN NATL RAILWAY CO COM 136375102 11,829 111,109 SH   DFND 2 0 111,109 0
CANADIAN NATL RAILWAY CO COM 136375102 7,875 74,053 SH   DFND 16 0 63,155 10,898
CANADIAN NATL RAILWAY CO COM 136375102 28 259 SH   DFND 18 0 83 176
CANADIAN NATURAL RESOURCES COM 136385101 157,898 9,862,585 SH   DFND 2 0 9,528,811 333,774
CANADIAN NATURAL RESOURCES COM 136385101 6,007 375,162 SH   DFND 2,3 0 375,162 0
CANADIAN NATURAL RESOURCES COM 136385101 363 22,678 SH   DFND 16 0 3,442 19,236
CANADIAN NATURAL RESOURCES COM 136385101 5 287 SH   DFND 18 0 287 0
CANADIAN PACIFIC RAILWAY LTD COM 13645T100 79 259 SH   DFND 2 0 259 0
CANADIAN PACIFIC RAILWAY LTD COM 13645T100 5,238 17,217 SH   DFND 16 0 650 16,567
CANADIAN PACIFIC RAILWAY LTD COM 13645T100 19 61 SH   DFND 16,18 0 61 0
CANADIAN PACIFIC RAILWAY LTD COM 13645T100 22 73 SH   DFND 18 0 0 73
CANNAE HOLDINGS INC COM 13765N107 271 7,279 SH   DFND 2 0 7,279 0
CANNAE HOLDINGS INC COM 13765N107 4 104 SH   DFND 16 0 0 104
CANOPY GROWTH CORP COM 138035100 4,932 344,447 SH   DFND 16 0 0 344,447
CANTEL MEDICAL CORP COM 138098108 30,254 688,520 SH   DFND 2 0 687,420 1,100
CANTEL MEDICAL CORP COM 138098108 149 3,378 SH   DFND 16 0 649 2,729
CAPITAL CITY BANK GROUP INC COM 139674105 5,950 316,696 SH   DFND 2 0 316,696 0
CAPITAL BANCORP INC/MD COM 139737100 3,529 373,008 SH   DFND 2 0 373,008 0
CAPITAL ONE FINANCIAL CORP COM 14040H105 36,587 509,148 SH   DFND 2 0 498,848 10,300
CAPITAL ONE FINANCIAL CORP COM 14040H105 113 1,570 SH   DFND 2,3 0 1,570 0
CAPITAL ONE FINANCIAL CORP COM 14040H105 3,149 43,921 SH   DFND 16 0 380 43,541
CAPITAL ONE FINANCIAL CORP COM 14040H105 28 389 SH   DFND 18 0 389 0
CAPSTEAD MORTGAGE CORP COM 14067E506 2,410 428,765 SH   DFND 2 0 418,665 10,100
CAPSTAR FINANCIAL HOLDINGS I COM 14070T102 104 10,576 SH   DFND 2 0 10,576 0
CARDINAL HEALTH INC COM 14149Y108 19,210 408,565 SH   DFND 2 0 283,325 125,240
CARDINAL HEALTH INC COM 14149Y108 47 1,002 SH   DFND 2,3 0 1,002 0
CARDINAL HEALTH INC COM 14149Y108 7,395 157,506 SH   DFND 16 0 68,461 89,045
CARDINAL HEALTH INC COM 14149Y108 89 1,905 SH   DFND 18 0 1,880 25
CARDIOVASCULAR SYSTEMS INC COM 141619106 8,549 217,256 SH   DFND 2 0 214,356 2,900
CARDIOVASCULAR SYSTEMS INC COM 141619106 7 168 SH   DFND 16 0 0 168
CARDLYTICS INC COM 14161W105 2,204 31,230 SH   DFND 2 0 31,230 0
CARDLYTICS INC COM 14161W105 44,719 633,695 SH   DFND 10 0 464,704 168,991
CARDLYTICS INC COM 14161W105 1,320 18,698 SH   DFND 16 0 0 18,698
CAREDX INC COM 14167L103 1,054 27,784 SH   DFND 2 0 27,784 0
CAREDX INC COM 14167L103 26 675 SH   DFND 16 0 0 675
CARETRUST REIT INC COM 14174T107 9,251 519,785 SH   DFND 2 0 508,985 10,800
CARETRUST REIT INC COM 14174T107 421 23,707 SH   DFND 16 0 3,062 20,645
CARGURUS INC COM 141788109 6 275 SH   DFND 2 0 275 0
CARGURUS INC COM 141788109 2,426 112,172 SH   DFND 16 0 108,400 3,772
CARLISLE COS INC COM 142339100 16,193 132,337 SH   DFND 2 0 130,158 2,179
CARLISLE COS INC COM 142339100 457 3,766 SH   DFND 16 0 1,245 2,521
CARLISLE COS INC COM 142339100 3 23 SH   DFND 18 0 0 23
CARMAX INC COM 143130102 8,773 95,457 SH   DFND 2 0 91,257 4,200
CARMAX INC COM 143130102 51 557 SH   DFND 2,3 0 557 0
CARMAX INC COM 143130102 4,088 44,469 SH   DFND 16 0 36,831 7,638
CARMAX INC COM 143130102 23 250 SH   DFND 18 0 250 0
CARLYLE GROUP INC/THE LP 14316J108 6 235 SH   DFND 2 0 235 0
CARLYLE GROUP INC/THE LP 14316J108 413 16,723 SH   DFND 16 0 0 16,723
CARNIVAL CORP COM 143658300 3,567 235,014 SH   DFND 2 0 234,989 25
CARNIVAL CORP COM 143658300 27 1,784 SH   DFND 2,3 0 1,784 0
CARNIVAL CORP COM 143658300 3,904 257,600 SH   DFND 16 0 295 257,305
CARNIVAL CORP COM 143658300 18 1,156 SH   DFND 18 0 1,156 0
CARNIVAL PLC-ADR ADR 14365C103 275 21,293 SH   DFND 2 0 0 21,293
CARNIVAL PLC-ADR ADR 14365C103 16 1,223 SH   DFND 16 0 0 1,223
CARRIAGE SERVICES INC COM 143905107 8,478 380,044 SH   DFND 2 0 380,044 0
CARRIAGE SERVICES INC COM 143905107 14,733 660,666 SH   DFND 10 0 660,666 0
CARRIAGE SERVICES INC COM 143905107 16 721 SH   DFND 16 0 0 721
CARPARTS.COM INC COM 14427M107 2,882 266,599 SH   DFND 2 0 266,599 0
CARPARTS.COM INC COM 14427M107 40 4,200 SH   DFND 16 0 0 4,200
CARPENTER TECHNOLOGY COM 144285103 8,665 477,064 SH   DFND 2 0 457,164 19,900
CARPENTER TECHNOLOGY COM 144285103 134 7,376 SH   DFND 2,3 0 7,376 0
CARPENTER TECHNOLOGY COM 144285103 838 46,245 SH   DFND 16 0 17,618 28,627
CARRIER GLOBAL CORP COM 14448C104 255,609 8,369,637 SH   DFND 2 0 8,013,932 355,705
CARRIER GLOBAL CORP COM 14448C104 8,279 271,054 SH   DFND 2,3 0 271,054 0
CARRIER GLOBAL CORP COM 14448C104 5,553 181,826 SH   DFND 16 0 4,702 177,124
CARRIER GLOBAL CORP COM 14448C104 1 37 SH   DFND 16,18 0 37 0
CARRIER GLOBAL CORP COM 14448C104 141 4,626 SH   DFND 18 0 4,546 80
CARROLS RESTAURANT GROUP INC COM 14574X104 7,662 1,187,919 SH   DFND 2 0 1,084,919 103,000
CARROLS RESTAURANT GROUP INC COM 14574X104 271 42,000 SH   DFND 2,3 0 42,000 0
CARROLS RESTAURANT GROUP INC COM 14574X104 15 2,300 SH   DFND 16 0 0 2,300
CARTER BANK & TRUST COM 146102108 252 37,757 SH   DFND 2 0 37,757 0
CARTER BANK & TRUST COM 146102108 6 885 SH   DFND 16 0 0 885
CARTER'S INC COM 146229109 7,398 85,448 SH   DFND 2 0 82,948 2,500
CARTER'S INC COM 146229109 836 9,656 SH   DFND 16 0 8,421 1,235
CARVANA CO COM 146869102 296 1,327 SH   DFND 2 0 1,327 0
CARVANA CO COM 146869102 1,062 4,761 SH   DFND 16 0 3,270 1,491
CASA SYSTEMS INC COM 14713L102 4,117 1,021,540 SH   DFND 2 0 1,021,540 0
CASA SYSTEMS INC COM 14713L102 31 7,759 SH   DFND 16 0 0 7,759
CASELLA WASTE SYSTEMS INC-A COM 147448104 29,397 526,366 SH   DFND 2 0 526,366 0
CASELLA WASTE SYSTEMS INC-A COM 147448104 23,335 417,822 SH   DFND 10 0 303,510 114,312
CASELLA WASTE SYSTEMS INC-A COM 147448104 3,857 69,082 SH   DFND 16 0 65,733 3,349
CASEY'S GENERAL STORES INC COM 147528103 14,961 84,219 SH   DFND 2 0 82,664 1,555
CASEY'S GENERAL STORES INC COM 147528103 4,594 25,844 SH   DFND 16 0 23,189 2,655
CASI PHARMACEUTICALS INC COM 14757U109 26 17,000 SH   DFND 16 0 0 17,000
CASS INFORMATION SYSTEMS INC COM 14808P109 879 21,848 SH   DFND 2 0 21,848 0
CASS INFORMATION SYSTEMS INC COM 14808P109 90 2,228 SH   DFND 16 0 2,188 40
CASTLE BIOSCIENCES INC COM 14843C105 7,497 145,719 SH   DFND 2 0 145,719 0
CASTLE BIOSCIENCES INC COM 14843C105 10 197 SH   DFND 16 0 0 197
CATALENT INC COM 148806102 3,694 43,127 SH   DFND 2 0 37,627 5,500
CATALENT INC COM 148806102 49 570 SH   DFND 2,3 0 570 0
CATALENT INC COM 148806102 4,040 47,143 SH   DFND 16 0 46,388 755
CATALYST BIOSCIENCES INC COM 14888D208 54 12,532 SH   DFND 2 0 12,532 0
CATALYST PHARMACEUTICALS INC COM 14888U101 2 550 SH   DFND 2 0 550 0
CATALYST PHARMACEUTICALS INC COM 14888U101 84 28,316 SH   DFND 16 0 0 28,316
CATERPILLAR INC COM 149123101 190,720 1,278,713 SH   DFND 2 0 1,168,326 110,387
CATERPILLAR INC COM 149123101 278 1,867 SH   DFND 2,3 0 1,867 0
CATERPILLAR INC COM 149123101 156,498 1,050,290 SH   DFND 10 0 1,050,290 0
CATERPILLAR INC COM 149123101 30,163 202,194 SH   DFND 16 0 37,300 164,894
CATERPILLAR INC COM 149123101 53 356 SH   DFND 16,18 0 356 0
CATERPILLAR INC COM 149123101 552 3,698 SH   DFND 18 0 3,698 0
CATCHMARK TIMBER TRUST INC-A COM 14912Y202 11 1,279 SH   DFND 2 0 1,279 0
CATCHMARK TIMBER TRUST INC-A COM 14912Y202 382 42,801 SH   DFND 16 0 41,466 1,335
CATHAY GENERAL BANCORP COM 149150104 20,279 935,385 SH   DFND 2 0 878,685 56,700
CATHAY GENERAL BANCORP COM 149150104 418 19,300 SH   DFND 2,3 0 19,300 0
CATHAY GENERAL BANCORP COM 149150104 60 2,773 SH   DFND 16 0 0 2,773
CATO CORP-CLASS A COM 149205106 1,389 177,635 SH   DFND 2 0 177,635 0
CATO CORP-CLASS A COM 149205106 9 1,097 SH   DFND 16 0 0 1,097
CAVCO INDUSTRIES INC COM 149568107 13,832 76,714 SH   DFND 2 0 75,999 715
CAVCO INDUSTRIES INC COM 149568107 19 103 SH   DFND 16 0 0 103
CEDAR FAIR LP LP 150185106 216 7,699 SH   DFND 16 0 0 7,699
CEL-SCI CORP COM 150837607 2 185 SH   DFND 2 0 185 0
CEL-SCI CORP COM 150837607 432 33,850 SH   DFND 16 0 0 33,850
CELANESE CORP COM 150870103 51,606 480,280 SH   DFND 2 0 480,264 16
CELANESE CORP COM 150870103 43 404 SH   DFND 2,3 0 404 0
CELANESE CORP COM 150870103 979 9,119 SH   DFND 16 0 1,823 7,296
CELANESE CORP COM 150870103 7 61 SH   DFND 16,18 0 61 0
CELANESE CORP COM 150870103 3 24 SH   DFND 18 0 24 0
CELLULAR BIOMEDICINE GROUP I COM 15117P102 582 31,732 SH   DFND 2 0 31,732 0
CEMEX SAB-SPONS ADR PART CER ADR 151290889 1 328 SH   DFND 2 0 0 328
CEMEX SAB-SPONS ADR PART CER ADR 151290889 134 35,243 SH   DFND 16 0 0 35,243
CENTENE CORP COM 15135B101 187,643 3,216,960 SH   DFND 2 0 2,509,231 707,729
CENTENE CORP COM 15135B101 117 1,999 SH   DFND 2,3 0 1,999 0
CENTENE CORP COM 15135B101 204,025 3,498,515 SH   DFND 10 0 3,498,515 0
CENTENE CORP COM 15135B101 32,930 564,223 SH   DFND 16 0 4,415 559,808
CENTENE CORP COM 15135B101 95 1,627 SH   DFND 16,18 0 1,627 0
CENTENE CORP COM 15135B101 94 1,613 SH   DFND 18 0 1,555 58
CENTENNIAL RESOURCE DEVELO-A COM 15136A102 24 39,790 SH   DFND 2 0 39,790 0
CENTERPOINT ENERGY INC COM 15189T107 3,474 179,589 SH   DFND 2 0 179,589 0
CENTERPOINT ENERGY INC COM 15189T107 36 1,875 SH   DFND 2,3 0 1,875 0
CENTERPOINT ENERGY INC COM 15189T107 3,479 180,074 SH   DFND 16 0 105,308 74,766
CENTERPOINT ENERGY INC COM 15189T107 7 350 SH   DFND 18 0 350 0
CENTRAL GARDEN & PET CO COM 153527106 1,690 42,330 SH   DFND 2 0 42,330 0
CENTRAL GARDEN AND PET CO-A COM 153527205 10,462 289,506 SH   DFND 2 0 0 289,506
CENTRAL GARDEN AND PET CO-A COM 153527205 243 6,741 SH   DFND 16 0 0 6,741
CENTRAL PACIFIC FINANCIAL CO COM 154760409 7,187 529,576 SH   DFND 2 0 524,776 4,800
CENTRAL PACIFIC FINANCIAL CO COM 154760409 4 302 SH   DFND 16 0 0 302
CENTURY ALUMINUM COMPANY COM 156431108 1,584 222,381 SH   DFND 2 0 217,881 4,500
CENTURY ALUMINUM COMPANY COM 156431108 9 1,200 SH   DFND 16 0 0 1,200
CENTURY COMMUNITIES INC COM 156504300 6,250 147,647 SH   DFND 2 0 144,947 2,700
CENTURY COMMUNITIES INC COM 156504300 1 15 SH   DFND 16 0 0 15
CENTURYLINK INC COM 156700106 7,144 708,039 SH   DFND 2 0 671,689 36,350
CENTURYLINK INC COM 156700106 34 3,397 SH   DFND 2,3 0 3,397 0
CENTURYLINK INC COM 156700106 3,862 382,788 SH   DFND 16 0 84 382,704
CENTURYLINK INC COM 156700106 32 3,156 SH   DFND 18 0 3,156 0
CERENCE INC COM 156727109 165,358 3,383,636 SH   DFND 2 0 3,317,008 66,628
CERENCE INC COM 156727109 411 8,400 SH   DFND 2,3 0 8,400 0
CERENCE INC COM 156727109 1,442 29,518 SH   DFND 16 0 27,375 2,143
CERIDIAN HCM HOLDING INC COM 15677J108 19,293 233,429 SH   DFND 2 0 229,379 4,050
CERIDIAN HCM HOLDING INC COM 15677J108 0 4 SH   DFND 16 0 0 4
CERNER CORP COM 156782104 77,393 1,070,476 SH   DFND 2 0 1,070,476 0
CERNER CORP COM 156782104 76 1,052 SH   DFND 2,3 0 1,052 0
CERNER CORP COM 156782104 695 9,617 SH   DFND 16 0 0 9,617
CERNER CORP COM 156782104 24 323 SH   DFND 18 0 299 24
CERUS CORP COM 157085101 0 10 SH   DFND 2 0 10 0
CERUS CORP COM 157085101 559 89,247 SH   DFND 16 0 0 89,247
CEVA INC COM 157210105 3,839 97,529 SH   DFND 2 0 96,729 800
CHAMPIONX CORP COM 15872M104 9,840 1,231,607 SH   DFND 2 0 1,224,507 7,100
CHAMPIONX CORP COM 15872M104 61 7,568 SH   DFND 16 0 6,823 745
CHAMPIONX CORP COM 15872M104 1 97 SH   DFND 18 0 0 97
CHANGE HEALTHCARE INC COM 15912K100 976 67,202 SH   DFND 2 0 67,202 0
CHANGE HEALTHCARE INC COM 15912K100 818 56,365 SH   DFND 16 0 52,294 4,071
CHNG 6 06/30/22 CONV PRF 15912K209 26,866 505,000 SH   DFND 2 0 0 505,000
CHANNELADVISOR CORP COM 159179100 1,749 120,909 SH   DFND 2 0 120,909 0
CHARLES RIVER LABORATORIES COM 159864107 94,138 415,710 SH   DFND 2 0 413,680 2,030
CHARLES RIVER LABORATORIES COM 159864107 48 212 SH   DFND 2,3 0 212 0
CHARLES RIVER LABORATORIES COM 159864107 77,556 342,484 SH   DFND 10 0 286,511 55,973
CHARLES RIVER LABORATORIES COM 159864107 715 3,156 SH   DFND 16 0 712 2,444
CHART INDUSTRIES INC COM 16115Q308 17,480 248,756 SH   DFND 2 0 245,706 3,050
CHART INDUSTRIES INC COM 16115Q308 403 5,735 SH   DFND 16 0 5,607 128
CHARTER COMMUNICATIONS INC-A COM 16119P108 97,611 156,345 SH   DFND 2 0 153,970 2,375
CHARTER COMMUNICATIONS INC-A COM 16119P108 320 513 SH   DFND 2,3 0 513 0
CHARTER COMMUNICATIONS INC-A COM 16119P108 33,641 53,913 SH   DFND 10 0 53,913 0
CHARTER COMMUNICATIONS INC-A COM 16119P108 3,016 4,829 SH   DFND 16 0 3 4,826
CHARTER COMMUNICATIONS INC-A COM 16119P108 105 168 SH   DFND 18 0 168 0
CHATHAM LODGING TRUST COM 16208T102 2,329 305,603 SH   DFND 2 0 305,393 210
CHATHAM LODGING TRUST COM 16208T102 41 5,389 SH   DFND 16 0 0 5,389
CHEESECAKE FACTORY INC/THE COM 163072101 6,679 240,768 SH   DFND 2 0 237,468 3,300
CHEESECAKE FACTORY INC/THE COM 163072101 75 2,693 SH   DFND 16 0 0 2,693
CHEESECAKE FACTORY INC/THE COM 163072101 3 118 SH   DFND 18 0 118 0
CHEFS' WAREHOUSE INC/THE COM 163086101 3,659 251,676 SH   DFND 2 0 251,676 0
CHEFS' WAREHOUSE INC/THE COM 163086101 0 30 SH   DFND 16 0 0 30
CHEGG INC COM 163092109 52 727 SH   DFND 2 0 727 0
CHEGG INC COM 163092109 18,020 252,660 SH   DFND 16 0 67,967 184,693
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EXPEDIA GROUP INC COM 30212P303 9,692 105,699 SH   DFND 2 0 105,693 6
EXPEDIA GROUP INC COM 30212P303 43 472 SH   DFND 2,3 0 472 0
EXPEDIA GROUP INC COM 30212P303 1,998 21,800 SH   DFND 16 0 3,849 17,951
EXPEDIA GROUP INC COM 30212P303 1 15 SH   DFND 18 0 15 0
EXPEDITORS INTL WASH INC COM 302130109 109,107 1,205,319 SH   DFND 2 0 1,197,519 7,800
EXPEDITORS INTL WASH INC COM 302130109 53 582 SH   DFND 2,3 0 582 0
EXPEDITORS INTL WASH INC COM 302130109 2,128 23,498 SH   DFND 16 0 17,709 5,789
EXPEDITORS INTL WASH INC COM 302130109 59 656 SH   DFND 18 0 656 0
EXPONENT INC COM 30214U102 24,119 334,853 SH   DFND 2 0 330,253 4,600
EXPONENT INC COM 30214U102 25,885 359,405 SH   DFND 10 0 263,560 95,845
EXPONENT INC COM 30214U102 2,713 37,630 SH   DFND 16 0 37,280 350
EXPONENT INC COM 30214U102 63,836 886,249 SH   DFND 17 0 886,249 0
EXTENDED STAY AMERICA INC COM 30224P200 29,102 2,435,330 SH   DFND 2 0 2,430,097 5,233
EXTENDED STAY AMERICA INC COM 30224P200 839 70,221 SH   DFND 16 0 50,210 20,011
EXTENDED STAY AMERICA INC COM 30224P200 0 18 SH   DFND 18 0 18 0
EXTRA SPACE STORAGE INC COM 30225T102 173,783 1,624,307 SH   DFND 2 0 1,533,159 91,148
EXTRA SPACE STORAGE INC COM 30225T102 1,063 9,942 SH   DFND 2,3 0 9,942 0
EXTRA SPACE STORAGE INC COM 30225T102 80 745 SH   DFND 10 0 745 0
EXTRA SPACE STORAGE INC COM 30225T102 1,556 14,554 SH   DFND 16 0 12,424 2,130
EXTREME NETWORKS INC COM 30226D106 9,900 2,462,904 SH   DFND 2 0 2,462,904 0
EXTREME NETWORKS INC COM 30226D106 2 610 SH   DFND 16 0 0 610
EXTERRAN CORP COM 30227H106 2,864 688,273 SH   DFND 2 0 688,273 0
EXTERRAN CORP COM 30227H106 3 629 SH   DFND 16 0 0 629
EZCORP INC-CL A COM 302301106 10,086 2,004,898 SH   DFND 2 0 0 2,004,898
EZCORP INC-CL A COM 302301106 1 106 SH   DFND 16 0 0 106
EXXON MOBIL CORP COM 30231G102 112,545 3,278,356 SH   DFND 2 0 3,212,078 66,278
EXXON MOBIL CORP COM 30231G102 500 14,560 SH   DFND 2,3 0 14,560 0
EXXON MOBIL CORP COM 30231G102 43,787 1,276,257 SH   DFND 16 0 364,056 912,201
EXXON MOBIL CORP COM 30231G102 207 6,021 SH   DFND 16,18 0 6,021 0
EXXON MOBIL CORP COM 30231G102 651 18,978 SH   DFND 18 0 18,507 471
FBL FINANCIAL GROUP INC-CL A COM 30239F106 10,696 221,890 SH   DFND 2 0 221,890 0
FLIR SYSTEMS INC COM 302445101 2,594 72,350 SH   DFND 2 0 72,350 0
FLIR SYSTEMS INC COM 302445101 16 450 SH   DFND 2,3 0 450 0
FLIR SYSTEMS INC COM 302445101 56,960 1,598,557 SH   DFND 16 0 17,323 1,581,234
FLIR SYSTEMS INC COM 302445101 72 2,000 SH   DFND 16,18 0 2,000 0
FLIR SYSTEMS INC COM 302445101 54 1,503 SH   DFND 18 0 1,503 0
FMC CORP COM 302491303 304,521 2,875,120 SH   DFND 2 0 1,949,923 925,197
FMC CORP COM 302491303 47 443 SH   DFND 2,3 0 443 0
FMC CORP COM 302491303 5,952 56,424 SH   DFND 16 0 25,300 31,124
FMC CORP COM 302491303 19 184 SH   DFND 18 0 108 76
FNB CORP COM 302520101 4,969 732,819 SH   DFND 2 0 712,219 20,600
FNB CORP COM 302520101 430 63,451 SH   DFND 16 0 0 63,451
FNB CORP COM 302520101 11 1,576 SH   DFND 16,18 0 1,576 0
FB FINANCIAL CORP COM 30257X104 4,763 189,646 SH   DFND 2 0 188,046 1,600
FB FINANCIAL CORP COM 30257X104 15 595 SH   DFND 16 0 0 595
FS KKR CAPITAL CORP COM 302635206 7,549 459,326 SH   DFND 16 0 0 459,326
FS KKR CAPITAL CORP COM 302635206 47 2,993 SH   DFND 18 0 2,993 0
FTI CONSULTING INC COM 302941109 9,469 89,348 SH   DFND 2 0 87,848 1,500
FTI CONSULTING INC COM 302941109 3,305 31,187 SH   DFND 16 0 30,909 278
FACEBOOK INC-CLASS A COM 30303M102 2,316,778 8,846,036 SH   DFND 2 0 8,146,213 699,823
FACEBOOK INC-CLASS A COM 30303M102 43,446 165,891 SH   DFND 2,3 0 165,891 0
FACEBOOK INC-CLASS A COM 30303M102 14,502 55,373 SH   DFND 2,10 0 55,373 0
FACEBOOK INC-CLASS A COM 30303M102 568,837 2,173,672 SH   DFND 10 0 2,173,672 0
FACEBOOK INC-CLASS A COM 30303M102 195,509 745,686 SH   DFND 16 0 193,661 552,025
FACEBOOK INC-CLASS A COM 30303M102 421 1,608 SH   DFND 16,18 0 1,608 0
FACEBOOK INC-CLASS A COM 30303M102 789 3,013