The Securities and Exchange Commission has not necessarily reviewed the information in this filing and has not determined if it is accurate and complete.
The reader should not assume that the information is accurate and complete.

UNITED STATES SECURITIES AND EXCHANGE COMMISSION
Washington, D.C. 20549
FORM 13F

FORM 13F INFORMATION TABLE
OMB APPROVAL
OMB Number: 3235-0006
Estimated average burden
hours per response: 23.8
COLUMN 1 COLUMN 2 COLUMN 3 COLUMN 4 COLUMN 5 COLUMN 6 COLUMN 7 COLUMN 8
VALUE SHRS OR SH/ PUT/ INVESTMENT OTHER VOTING AUTHORITY
NAME OF ISSUER TITLE OF CLASS CUSIP (x$1000) PRN AMT PRN CALL DISCRETION MANAGER SOLE SHARED NONE
AAON INC COM 000360206 17,124 372,753 SH   DFND 2 0 372,753 0
AAON INC COM 000360206 13 286 SH   DFND 02,03 0 286 0
AAON INC COM 000360206 20 427 SH   DFND 16 0 0 427
AAR CORP COM 000361105 8,489 206,007 SH   DFND 2 0 206,007 0
AAR CORP COM 000361105 4 102 SH   DFND 16 0 0 102
AAR CORP COM 000361105 2 49 SH   DFND 18 0 49 0
ABB LTD-SPON ADR ADR 000375204 417 21,193 SH   DFND 2 0 0 21,193
ABB LTD-SPON ADR ADR 000375204 66,009 3,359,011 SH   DFND 16 0 0 3,359,011
ABB LTD-SPON ADR ADR 000375204 32 1,605 SH   DFND 16,18 0 0 1,605
ABB LTD-SPON ADR ADR 000375204 22 1,118 SH   DFND 18 0 0 1,118
ACCO BRANDS CORP COM 00081T108 2,864 290,117 SH   DFND 2 0 290,117 0
ACCO BRANDS CORP COM 00081T108 159 15,948 SH   DFND 16 0 0 15,948
ADMA BIOLOGICS INC COM 000899104 0 5 SH   DFND 2 0 5 0
ADMA BIOLOGICS INC COM 000899104 205 46,000 SH   DFND 16 0 0 46,000
ABM INDUSTRIES INC COM 000957100 12,742 350,827 SH   DFND 2 0 350,827 0
ABM INDUSTRIES INC COM 000957100 38 1,041 SH   DFND 16 0 0 1,041
ABM INDUSTRIES INC COM 000957100 1 40 SH   DFND 18 0 40 0
AFLAC INC COM 001055102 24,859 475,113 SH   DFND 2 0 475,113 0
AFLAC INC COM 001055102 140 2,679 SH   DFND 02,03 0 2,679 0
AFLAC INC COM 001055102 62,034 1,185,886 SH   DFND 10 0 1,185,886 0
AFLAC INC COM 001055102 8,145 155,655 SH   DFND 16 0 1,961 153,694
AFLAC INC COM 001055102 9 173 SH   DFND 16,18 0 173 0
AFLAC INC COM 001055102 219 4,184 SH   DFND 18 0 4,184 0
AGCO CORP COM 001084102 30,723 405,852 SH   DFND 2 0 398,652 7,200
AGCO CORP COM 001084102 863 11,403 SH   DFND 16 0 0 11,403
ACM RESEARCH INC-CLASS A COM 00108J109 1,385 99,948 SH   DFND 2 0 99,948 0
AVX CORP COM 002444107 16,019 1,053,884 SH   DFND 2 0 1,053,884 0
AVX CORP COM 002444107 378 24,862 SH   DFND 16 0 0 24,862
AZZ INC COM 002474104 9,995 229,461 SH   DFND 2 0 229,461 0
AZZ INC COM 002474104 9 200 SH   DFND 16 0 0 200
AARON'S INC COM 002535300 50,787 790,318 SH   DFND 2 0 750,018 40,300
AARON'S INC COM 002535300 273 4,250 SH   DFND 02,03 0 4,250 0
AARON'S INC COM 002535300 91 1,413 SH   DFND 16 0 0 1,413
ABBOTT LABORATORIES COM 002824100 631,310 7,545,199 SH   DFND 2 0 6,742,038 803,161
ABBOTT LABORATORIES COM 002824100 6,318 75,510 SH   DFND 02,03 0 75,510 0
ABBOTT LABORATORIES COM 002824100 10,025 119,820 SH   DFND 02,10 0 119,820 0
ABBOTT LABORATORIES COM 002824100 18,727 223,906 SH   DFND 10 0 223,906 0
ABBOTT LABORATORIES COM 002824100 69,283 829,346 SH   DFND 16 0 115,386 713,960
ABBOTT LABORATORIES COM 002824100 370 4,419 SH   DFND 16,18 0 4,419 0
ABBOTT LABORATORIES COM 002824100 845 10,094 SH   DFND 18 0 10,094 0
ABBVIE INC COM 00287Y109 309,420 4,086,364 SH   DFND 2 0 3,465,057 621,307
ABBVIE INC COM 00287Y109 405 5,351 SH   DFND 02,03 0 5,351 0
ABBVIE INC COM 00287Y109 179,449 2,298,528 SH   DFND 16 0 394,169 1,904,359
ABBVIE INC COM 00287Y109 458 6,043 SH   DFND 16,18 0 6,043 0
ABBVIE INC COM 00287Y109 1,140 15,045 SH   DFND 18 0 14,733 312
ABBVIE INC OPT 00287Y109 1,310 17,300 SH Call DFND 2 0 17,300 0
ABERCROMBIE & FITCH CO-CL A COM 002896207 5,159 330,710 SH   DFND 2 0 330,710 0
ABERCROMBIE & FITCH CO-CL A COM 002896207 1 60 SH   DFND 16 0 0 60
ABERDEEN ASIA-PAC INCOME FD COM 003009107 326 77,881 SH   DFND 16 0 0 77,881
ABERDEEN GLBL PREM PROP COM 00302L108 863 138,724 SH   DFND 16 0 0 138,724
ABERDEEN GLBL PREM PROP COM 00302L108 0 14 SH   DFND 18 0 14 0
ABERDEEN TTL DYN DIVID COM 00326L100 1,068 128,087 SH   DFND 16 0 0 128,087
ABERDEEN TTL DYN DIVID COM 00326L100 20 2,364 SH   DFND 18 0 2,364 0
ABIOMED INC COM 003654100 3,011 16,928 SH   DFND 2 0 16,928 0
ABIOMED INC COM 003654100 29 164 SH   DFND 02,03 0 164 0
ABIOMED INC COM 003654100 1,006 5,657 SH   DFND 16 0 2,007 3,650
ABRAXAS PETROLEUM CORP COM 003830106 13 25,025 SH   DFND 16 0 0 25,025
ACACIA RESEARCH CORP COM 003881307 3 1,240 SH   DFND 2 0 1,240 0
ACACIA RESEARCH CORP COM 003881307 46 17,001 SH   DFND 16 0 0 17,001
ACACIA COMMUNICATIONS INC COM 00401C108 7,399 113,105 SH   DFND 2 0 101,240 11,865
ACACIA COMMUNICATIONS INC COM 00401C108 76 1,169 SH   DFND 16 0 0 1,169
ACADIA HEALTHCARE CO INC COM 00404A109 8,821 283,816 SH   DFND 2 0 283,816 0
ACADIA HEALTHCARE CO INC COM 00404A109 571 18,345 SH   DFND 16 0 16,913 1,432
ACADIA PHARMACEUTICALS INC COM 004225108 9,370 260,385 SH   DFND 2 0 260,385 0
ACADIA PHARMACEUTICALS INC COM 004225108 31 865 SH   DFND 02,03 0 865 0
ACADIA PHARMACEUTICALS INC COM 004225108 2,291 63,653 SH   DFND 02,10 0 63,653 0
ACADIA PHARMACEUTICALS INC COM 004225108 561 15,591 SH   DFND 16 0 15,061 530
ACADIA PHARMACEUTICALS INC COM 004225108 23,753 660,000 SH   DFND 17 0 660,000 0
ACADIA REALTY TRUST COM 004239109 12,714 444,796 SH   DFND 2 0 444,796 0
ACADIA REALTY TRUST COM 004239109 3 120 SH   DFND 16 0 0 120
ACI WORLDWIDE INC COM 004498101 16,817 536,848 SH   DFND 2 0 536,848 0
ACI WORLDWIDE INC COM 004498101 413 13,161 SH   DFND 16 0 13,041 120
ACI WORLDWIDE INC COM 004498101 2 50 SH   DFND 18 0 0 50
ACORDA THERAPEUTICS INC COM 00484M106 1,186 412,795 SH   DFND 2 0 412,795 0
ACORDA THERAPEUTICS INC COM 00484M106 36 12,526 SH   DFND 16 0 12,376 150
ACORDA THERAPEUTICS INC CONV BND 00484MAA4 2,739 3,500,000 PRN   DFND 2 0 0 3,500,000
ACTIVISION BLIZZARD INC COM 00507V109 692,993 13,095,012 SH   DFND 2 0 11,997,672 1,097,340
ACTIVISION BLIZZARD INC COM 00507V109 28,568 539,835 SH   DFND 02,03 0 539,835 0
ACTIVISION BLIZZARD INC COM 00507V109 4,776 90,252 SH   DFND 02,10 0 90,252 0
ACTIVISION BLIZZARD INC COM 00507V109 92,806 1,754,027 SH   DFND 10 0 1,692,064 61,963
ACTIVISION BLIZZARD INC COM 00507V109 5,326 100,673 SH   DFND 16 0 69,484 31,189
ACTUANT CORP-A COM 00508X203 6,292 286,799 SH   DFND 2 0 286,799 0
ACUITY BRANDS INC COM 00508Y102 13,604 100,919 SH   DFND 2 0 96,744 4,175
ACUITY BRANDS INC COM 00508Y102 31 232 SH   DFND 02,03 0 232 0
ACUITY BRANDS INC COM 00508Y102 219 1,623 SH   DFND 16 0 1,419 204
ACUSHNET HOLDINGS CORP COM 005098108 2,860 108,305 SH   DFND 2 0 108,305 0
ACUSHNET HOLDINGS CORP COM 005098108 7 256 SH   DFND 16 0 0 256
ADAMAS PHARMACEUTICALS INC COM 00548A106 5,211 1,018,845 SH   DFND 2 0 1,018,845 0
ADAMAS PHARMACEUTICALS INC COM 00548A106 1 150 SH   DFND 16 0 0 150
ADAMS DIVERSIFIED EQUITY COM 006212104 193 12,207 SH   DFND 16 0 0 12,207
ADAPTIVE BIOTECHNOLOGIES COM 00650F109 7,722 249,886 SH   DFND 2 0 249,886 0
ADAPTIVE BIOTECHNOLOGIES COM 00650F109 84 2,716 SH   DFND 16 0 2,713 3
ADDUS HOMECARE CORP COM 006739106 11,513 145,213 SH   DFND 2 0 145,213 0
ADDUS HOMECARE CORP COM 006739106 352 4,451 SH   DFND 16 0 4,243 208
ADOBE INC COM 00724F101 1,123,504 4,066,990 SH   DFND 2 0 3,613,424 453,566
ADOBE INC COM 00724F101 26,306 95,223 SH   DFND 02,03 0 95,223 0
ADOBE INC COM 00724F101 23,081 83,550 SH   DFND 02,10 0 83,550 0
ADOBE INC COM 00724F101 295,153 1,068,423 SH   DFND 10 0 1,058,765 9,658
ADOBE INC COM 00724F101 42,042 152,441 SH   DFND 16 0 47,429 105,012
ADOBE INC COM 00724F101 59 214 SH   DFND 18 0 173 41
ADTALEM GLOBAL EDUCATION INC COM 00737L103 22,358 586,948 SH   DFND 2 0 547,448 39,500
ADTALEM GLOBAL EDUCATION INC COM 00737L103 196 5,150 SH   DFND 02,03 0 5,150 0
ADTALEM GLOBAL EDUCATION INC COM 00737L103 4 114 SH   DFND 16 0 0 114
ADTALEM GLOBAL EDUCATION INC COM 00737L103 24,655 647,274 SH   DFND 17 0 647,274 0
ADTRAN INC COM 00738A106 4,636 408,776 SH   DFND 2 0 372,876 35,900
ADTRAN INC COM 00738A106 61 5,400 SH   DFND 16 0 0 5,400
ADVANCE AUTO PARTS INC COM 00751Y106 8,747 52,866 SH   DFND 2 0 52,406 460
ADVANCE AUTO PARTS INC COM 00751Y106 43 262 SH   DFND 02,03 0 262 0
ADVANCE AUTO PARTS INC COM 00751Y106 272 1,644 SH   DFND 16 0 718 926
ADVANCE AUTO PARTS INC COM 00751Y106 4 24 SH   DFND 18 0 0 24
AECOM COM 00766T100 16,262 432,975 SH   DFND 2 0 432,975 0
AECOM COM 00766T100 64 1,697 SH   DFND 16 0 0 1,697
AECOM COM 00766T100 2 48 SH   DFND 18 0 48 0
AEGION CORP COM 00770F104 4,556 213,109 SH   DFND 2 0 213,109 0
AEGION CORP COM 00770F104 11 500 SH   DFND 16 0 0 500
AERIE PHARMACEUTICALS INC COM 00771V108 10,545 548,712 SH   DFND 2 0 548,712 0
AERIE PHARMACEUTICALS INC COM 00771V108 9 466 SH   DFND 02,03 0 466 0
AERIE PHARMACEUTICALS INC COM 00771V108 1,359 70,694 SH   DFND 02,10 0 70,694 0
AERIE PHARMACEUTICALS INC COM 00771V108 6,585 342,980 SH   DFND 10 0 342,980 0
AERIE PHARMACEUTICALS INC COM 00771V108 85 4,386 SH   DFND 16 0 3,886 500
ADVANSIX INC COM 00773T101 4,898 190,393 SH   DFND 2 0 182,293 8,100
ADVANSIX INC COM 00773T101 199 7,721 SH   DFND 16 0 0 7,721
ADVANSIX INC COM 00773T101 0 18 SH   DFND 16,18 0 18 0
ADVANSIX INC COM 00773T101 1 26 SH   DFND 18 0 26 0
ADVERUM BIOTECHNOLOGIES INC COM 00773U108 383 70,304 SH   DFND 2 0 70,304 0
ADVERUM BIOTECHNOLOGIES INC COM 00773U108 1 166 SH   DFND 16 0 0 166
AEROJET ROCKETDYNE HOLDINGS COM 007800105 32,431 642,067 SH   DFND 2 0 626,067 16,000
AEROJET ROCKETDYNE HOLDINGS COM 007800105 3,101 61,378 SH   DFND 10 0 61,378 0
AEROJET ROCKETDYNE HOLDINGS COM 007800105 582 11,463 SH   DFND 16 0 9,431 2,032
AEROJET ROCKETDYNE HOLDINGS COM 007800105 3 66 SH   DFND 18 0 66 0
AEROJET ROCKETDYNE HOLDINGS INC CONV BND 007800AB1 9,371 4,700,000 PRN   DFND 2 0 0 4,700,000
AEROJET ROCKETDYNE HOLDINGS INC CONV BND 007800AB1 2 1,000 PRN   DFND 16 0 0 1,000
ADVANCED MICRO DEVICES COM 007903107 55,745 1,922,922 SH   DFND 2 0 1,922,922 0
ADVANCED MICRO DEVICES COM 007903107 251 8,644 SH   DFND 02,03 0 8,644 0
ADVANCED MICRO DEVICES COM 007903107 7,607 263,117 SH   DFND 16 0 81,579 181,538
ADVANCED MICRO DEVICES COM 007903107 170 5,850 SH   DFND 17 0 5,850 0
ADVANCED MICRO DEVICES INC CONV BND 007903BD8 5,589 1,500,000 PRN   DFND 2 0 0 1,500,000
ADVANCED DRAINAGE SYSTEMS IN COM 00790R104 6,085 188,572 SH   DFND 2 0 188,572 0
ADVANCED DRAINAGE SYSTEMS IN COM 00790R104 435 13,479 SH   DFND 16 0 0 13,479
ADVANCED DISPOSAL SERVICES I COM 00790X101 10 307 SH   DFND 2 0 307 0
ADVANCED DISPOSAL SERVICES I COM 00790X101 392 12,029 SH   DFND 16 0 12,029 0
AEGON N.V.-NY REG SHR COM 007924103 61 14,895 SH   DFND 2 0 0 14,895
AEGON N.V.-NY REG SHR COM 007924103 333 80,558 SH   DFND 16 0 0 80,558
ADVANCED ENERGY INDUSTRIES COM 007973100 112,804 1,964,864 SH   DFND 2 0 1,923,324 41,540
ADVANCED ENERGY INDUSTRIES COM 007973100 54 946 SH   DFND 16 0 946 0
ADVANCED ENERGY INDUSTRIES COM 007973100 11,491 200,153 SH   DFND 17 0 200,153 0
ADVANCED ENERGY INDUSTRIES COM 007973100 1 24 SH   DFND 18 0 0 24
AEROVIRONMENT INC COM 008073108 6,100 113,892 SH   DFND 2 0 113,892 0
AEROVIRONMENT INC COM 008073108 26 486 SH   DFND 16 0 0 486
AFFILIATED MANAGERS GROUP COM 008252108 1,812 21,749 SH   DFND 2 0 21,749 0
AFFILIATED MANAGERS GROUP COM 008252108 15 180 SH   DFND 02,03 0 180 0
AFFILIATED MANAGERS GROUP COM 008252108 942 11,312 SH   DFND 16 0 5,873 5,439
AFFILIATED MANAGERS GROUP COM 008252108 15 180 SH   DFND 16,18 0 180 0
AGILENT TECHNOLOGIES INC COM 00846U101 38,215 498,575 SH   DFND 2 0 495,670 2,905
AGILENT TECHNOLOGIES INC COM 00846U101 85 1,115 SH   DFND 02,03 0 1,115 0
AGILENT TECHNOLOGIES INC COM 00846U101 3,642 47,470 SH   DFND 16 0 22,171 25,299
AGNICO EAGLE MINES LTD COM 008474108 274 5,112 SH   DFND 16 0 0 5,112
AGENUS INC COM 00847G705 27 10,333 SH   DFND 16 0 0 10,333
AGILYSYS INC COM 00847J105 2,756 107,605 SH   DFND 2 0 107,605 0
AGIOS PHARMACEUTICALS INC COM 00847X104 193 5,988 SH   DFND 2 0 5,988 0
AGIOS PHARMACEUTICALS INC COM 00847X104 0 4 SH   DFND 16 0 0 4
AGIOS PHARMACEUTICALS INC COM 00847X104 41,818 1,290,632 SH   DFND 17 0 1,290,632 0
AGREE REALTY CORP COM 008492100 16,547 226,149 SH   DFND 2 0 225,349 800
AGREE REALTY CORP COM 008492100 1,243 17,000 SH   DFND 16 0 15,681 1,319
AIR LEASE CORP COM 00912X302 681 16,250 SH   DFND 2 0 16,250 0
AIR LEASE CORP COM 00912X302 1,775 42,438 SH   DFND 16 0 39,965 2,473
AIR LEASE CORP COM 00912X302 44,919 1,074,094 SH   DFND 17 0 1,074,094 0
AIR PRODUCTS & CHEMICALS INC COM 009158106 136,487 614,958 SH   DFND 2 0 595,110 19,848
AIR PRODUCTS & CHEMICALS INC COM 009158106 3,730 16,815 SH   DFND 02,03 0 16,815 0
AIR PRODUCTS & CHEMICALS INC COM 009158106 12,333 55,605 SH   DFND 16 0 10,965 44,641
AIR PRODUCTS & CHEMICALS INC COM 009158106 49 219 SH   DFND 16,18 0 219 0
AIR PRODUCTS & CHEMICALS INC COM 009158106 133 600 SH   DFND 18 0 600 0
AIR TRANSPORT SERVICES GROUP COM 00922R105 1,871 88,994 SH   DFND 2 0 88,994 0
AIR TRANSPORT SERVICES GROUP COM 00922R105 152 7,233 SH   DFND 16 0 0 7,233
AKAMAI TECHNOLOGIES INC COM 00971T101 11,695 127,978 SH   DFND 2 0 127,978 0
AKAMAI TECHNOLOGIES INC COM 00971T101 54 595 SH   DFND 02,03 0 595 0
AKAMAI TECHNOLOGIES INC COM 00971T101 1,496 16,365 SH   DFND 16 0 6,562 9,803
AKAMAI TECHNOLOGIES INC COM 00971T101 9 97 SH   DFND 18 0 50 47
AKORN INC COM 009728106 2,181 573,735 SH   DFND 2 0 573,735 0
AKORN INC COM 009728106 3 700 SH   DFND 16 0 0 700
AKEBIA THERAPEUTICS INC COM 00972D105 1 245 SH   DFND 2 0 245 0
AKEBIA THERAPEUTICS INC COM 00972D105 132 34,556 SH   DFND 16 0 0 34,556
ALAMO GROUP INC COM 011311107 6,002 50,985 SH   DFND 2 0 50,985 0
ALAMO GROUP INC COM 011311107 1 8 SH   DFND 16 0 0 8
ALARM.COM HOLDINGS INC COM 011642105 10,087 216,257 SH   DFND 2 0 216,257 0
ALARM.COM HOLDINGS INC COM 011642105 10 225 SH   DFND 16 0 0 225
ALASKA AIR GROUP INC COM 011659109 41,698 642,409 SH   DFND 2 0 642,409 0
ALASKA AIR GROUP INC COM 011659109 113 1,748 SH   DFND 02,03 0 1,748 0
ALASKA AIR GROUP INC COM 011659109 4,793 73,833 SH   DFND 16 0 5,411 68,422
ALASKA AIR GROUP INC COM 011659109 2 24 SH   DFND 18 0 24 0
ALBANY INTL CORP-CL A COM 012348108 25,114 278,543 SH   DFND 2 0 278,543 0
ALBANY INTL CORP-CL A COM 012348108 242 2,684 SH   DFND 16 0 1,590 1,094
ALBANY INTL CORP-CL A COM 012348108 3 35 SH   DFND 18 0 0 35
ALBEMARLE CORP COM 012653101 76,289 1,097,331 SH   DFND 2 0 1,097,331 0
ALBEMARLE CORP COM 012653101 27 385 SH   DFND 02,03 0 385 0
ALBEMARLE CORP COM 012653101 65,032 935,295 SH   DFND 10 0 935,295 0
ALBEMARLE CORP COM 012653101 3,167 45,567 SH   DFND 16 0 8,152 37,415
ALBEMARLE CORP COM 012653101 22 323 SH   DFND 18 0 323 0
ALCOA CORP COM 013872106 1,153 57,464 SH   DFND 2 0 57,464 0
ALCOA CORP COM 013872106 3,187 158,765 SH   DFND 16 0 4,191 154,574
ALCOA CORP COM 013872106 0 2 SH   DFND 18 0 2 0
ALDER BIOPHARMACEUTICALS INC COM 014339105 18,646 988,625 SH   DFND 2 0 888,705 99,920
ALDER BIOPHARMACEUTICALS INC COM 014339105 75 3,969 SH   DFND 02,03 0 3,969 0
ALDER BIOPHARMACEUTICALS INC CONV BND 014339AA3 7,195 6,500,000 PRN   DFND 2 0 0 6,500,000
ALEXANDER & BALDWIN INC COM 014491104 7,799 318,213 SH   DFND 2 0 303,813 14,400
ALEXANDER & BALDWIN INC COM 014491104 2 98 SH   DFND 16 0 0 98
ALEXANDER'S INC COM 014752109 775 2,223 SH   DFND 2 0 2,223 0
ALEXANDRIA REAL ESTATE EQUIT COM 015271109 148,994 967,158 SH   DFND 2 0 966,941 217
ALEXANDRIA REAL ESTATE EQUIT COM 015271109 63 408 SH   DFND 02,03 0 408 0
ALEXANDRIA REAL ESTATE EQUIT COM 015271109 1,205 7,828 SH   DFND 10 0 7,828 0
ALEXANDRIA REAL ESTATE EQUIT COM 015271109 3,861 25,056 SH   DFND 16 0 23,854 1,202
ALEXANDRIA REAL ESTATE EQUIT COM 015271109 2 12 SH   DFND 16,18 0 12 0
ALEXION PHARMACEUTICALS INC COM 015351109 445,641 4,550,188 SH   DFND 2 0 3,879,781 670,407
ALEXION PHARMACEUTICALS INC COM 015351109 15,667 159,968 SH   DFND 02,03 0 159,968 0
ALEXION PHARMACEUTICALS INC COM 015351109 3,751 38,302 SH   DFND 02,10 0 38,302 0
ALEXION PHARMACEUTICALS INC COM 015351109 89,625 915,290 SH   DFND 10 0 905,422 9,868
ALEXION PHARMACEUTICALS INC COM 015351109 16,630 169,925 SH   DFND 16 0 16,395 153,530
ALEXION PHARMACEUTICALS INC COM 015351109 6 61 SH   DFND 18 0 34 27
ALGONQUIN POWER & UTILITIES COM 015857105 2,825 206,266 SH   DFND 16 0 204,941 1,325
ALIBABA GROUP HOLDING-SP ADR ADR 01609W102 847,147 5,065,751 SH   DFND 2 0 4,723,412 342,339
ALIBABA GROUP HOLDING-SP ADR ADR 01609W102 22,329 133,521 SH   DFND 02,03 0 133,521 0
ALIBABA GROUP HOLDING-SP ADR ADR 01609W102 29,609 177,051 SH   DFND 02,10 0 177,051 0
ALIBABA GROUP HOLDING-SP ADR ADR 01609W102 235,317 1,407,150 SH   DFND 10 0 1,384,511 22,639
ALIBABA GROUP HOLDING-SP ADR ADR 01609W102 74,307 444,505 SH   DFND 16 0 82,289 362,216
ALIBABA GROUP HOLDING-SP ADR ADR 01609W102 40 242 SH   DFND 16,18 0 242 0
ALIBABA GROUP HOLDING-SP ADR ADR 01609W102 127 761 SH   DFND 18 0 744 17
ALIGN TECHNOLOGY INC COM 016255101 154,204 852,334 SH   DFND 2 0 745,166 107,168
ALIGN TECHNOLOGY INC COM 016255101 10,113 55,903 SH   DFND 02,03 0 55,903 0
ALIGN TECHNOLOGY INC COM 016255101 4,075 22,527 SH   DFND 16 0 5,284 17,243
ALIGN TECHNOLOGY INC COM 016255101 11 64 SH   DFND 18 0 51 13
ALLEGHANY CORP COM 017175100 46,156 57,857 SH   DFND 2 0 57,682 175
ALLEGHANY CORP COM 017175100 1,775 2,224 SH   DFND 16 0 1,966 258
ALLEGHENY TECHNOLOGIES INC COM 01741R102 29,174 1,440,716 SH   DFND 2 0 1,396,201 44,515
ALLEGHENY TECHNOLOGIES INC COM 01741R102 117 5,800 SH   DFND 02,03 0 5,800 0
ALLEGHENY TECHNOLOGIES INC COM 01741R102 51 2,521 SH   DFND 16 0 0 2,521
ALLEGIANT TRAVEL CO COM 01748X102 12,728 85,043 SH   DFND 2 0 85,043 0
ALLEGIANT TRAVEL CO COM 01748X102 159 1,062 SH   DFND 16 0 0 1,062
ALLETE INC COM 018522300 17,476 199,934 SH   DFND 2 0 199,934 0
ALLETE INC COM 018522300 307 3,517 SH   DFND 16 0 0 3,517
ALLETE INC COM 018522300 3 31 SH   DFND 18 0 31 0
ALLIANCE DATA SYSTEMS CORP COM 018581108 2,358 18,397 SH   DFND 2 0 18,397 0
ALLIANCE DATA SYSTEMS CORP COM 018581108 19 152 SH   DFND 02,03 0 152 0
ALLIANCE DATA SYSTEMS CORP COM 018581108 277 2,166 SH   DFND 16 0 274 1,892
ALLIANCE DATA SYSTEMS CORP COM 018581108 19 148 SH   DFND 16,18 0 148 0
ALLIANCE NATIONAL MUNI INC COM 01864U106 1,287 93,404 SH   DFND 16 0 0 93,404
ALLIANCE RESOURCE PARTNERS LP 01877R108 218 13,615 SH   DFND 16 0 0 13,615
ALLIANCEBERNSTEIN GL HI INC COM 01879R106 599 49,951 SH   DFND 16 0 0 49,951
ALLIANCEBERNSTEIN GL HI INC COM 01879R106 44 3,696 SH   DFND 16,18 0 3,696 0
ALLIANCEBERNSTEIN GL HI INC COM 01879R106 17 1,456 SH   DFND 18 0 1,456 0
ALLIANT ENERGY CORP COM 018802108 20,224 374,999 SH   DFND 2 0 374,999 0
ALLIANT ENERGY CORP COM 018802108 46 859 SH   DFND 02,03 0 859 0
ALLIANT ENERGY CORP COM 018802108 1,286 23,850 SH   DFND 16 0 616 23,234
ALLIANT ENERGY CORP COM 018802108 101 1,878 SH   DFND 18 0 1,878 0
ALLIANCEBERNSTEIN HOLDING LP LP 01881G106 1,482 50,484 SH   DFND 16 0 0 50,484
ALLIANZGI CONV & INCOME II COM 018825109 11,727 2,335,155 SH   DFND 16 0 0 2,335,155
ALLIANZGI CONVERTIBLE & INCO COM 018828103 1,193 211,351 SH   DFND 16 0 0 211,351
ALLIANZGI CONVERTIBLE & INCO COM 018828103 3 600 SH   DFND 18 0 600 0
ALLIANZGI EQUITY & CONVERTIB COM 018829101 567 26,253 SH   DFND 16 0 0 26,253
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ALLY FINANCIAL INC COM 02005N100 3,511 105,726 SH   DFND 16 0 32,597 73,129
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ALTICE USA INC- A COM 02156K103 1,407 51,226 SH   DFND 16 0 46,373 4,853
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AMEDISYS INC COM 023436108 1,184 9,040 SH   DFND 16 0 2,305 6,735
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AMERICAN EXPRESS CO COM 025816109 83,842 708,846 SH   DFND 2 0 708,846 0
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AMERICAN EXPRESS CO COM 025816109 64 545 SH   DFND 16,18 0 545 0
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AMERICAN WATER WORKS CO INC COM 030420103 22,002 177,107 SH   DFND 2 0 177,107 0
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ARCHER-DANIELS-MIDLAND CO COM 039483102 8 202 SH   DFND 18 0 202 0
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ARLO TECHNOLOGIES INC COM 04206A101 1,719 504,178 SH   DFND 2 0 504,178 0
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ARTISAN PARTNERS ASSET MA -A COM 04316A108 2,186 77,409 SH   DFND 2 0 77,409 0
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ASPEN TECHNOLOGY INC COM 045327103 51,661 419,734 SH   DFND 2 0 419,734 0
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ASPEN TECHNOLOGY INC COM 045327103 34,413 279,600 SH   DFND 17 0 279,600 0
ASSERTIO THERAPEUTICS INC COM 04545L107 696 543,382 SH   DFND 2 0 461,682 81,700
ASSERTIO THERAPEUTICS INC COM 04545L107 1 500 SH   DFND 16 0 0 500
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AIZ 6 1/2 03/15/21 CONV PRF 04621X207 19,405 155,000 SH   DFND 2 0 0 155,000
ASTEC INDUSTRIES INC COM 046224101 6,454 207,504 SH   DFND 2 0 207,504 0
ASTEC INDUSTRIES INC COM 046224101 16 515 SH   DFND 16 0 0 515
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ASTRAZENECA PLC-SPONS ADR ADR 046353108 8,530 191,165 SH   DFND 16 0 0 191,165
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ASTRONICS CORP COM 046433108 10 338 SH   DFND 16 0 0 338
ASURE SOFTWARE INC COM 04649U102 3,805 567,040 SH   DFND 2 0 567,040 0
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AXCELIS TECHNOLOGIES INC COM 054540208 3,050 178,424 SH   DFND 2 0 170,124 8,300
AVROBIO INC COM 05455M100 1,391 98,496 SH   DFND 2 0 98,496 0
AXA EQUITABLE HOLDINGS INC COM 054561105 9,884 446,053 SH   DFND 2 0 445,011 1,042
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B&G FOODS INC COM 05508R106 9 500 SH   DFND 18 0 500 0
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BGC PARTNERS INC-CL A COM 05541T101 16 2,870 SH   DFND 2 0 2,870 0
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CSG SYSTEMS INTERNATIONAL INC CONV BND 126349AF6 9,609 8,500,000 PRN   DFND 2 0 0 8,500,000
CSW INDUSTRIALS INC COM 126402106 2,774 40,165 SH   DFND 2 0 40,165 0
CSX CORP COM 126408103 212,325 3,065,110 SH   DFND 2 0 1,942,177 1,122,933
CSX CORP COM 126408103 200 2,888 SH   DFND 02,03 0 2,888 0
CSX CORP COM 126408103 12,051 173,753 SH   DFND 16 0 1,686 172,067
CSX CORP COM 126408103 185 2,676 SH   DFND 18 0 2,614 62
CTS CORP COM 126501105 6,511 201,210 SH   DFND 2 0 201,210 0
CTS CORP COM 126501105 194 5,990 SH   DFND 16 0 5,990 0
CNX RESOURCES CORP COM 12653C108 3,332 458,978 SH   DFND 2 0 458,978 0
CNX RESOURCES CORP COM 12653C108 220 30,360 SH   DFND 16 0 0 30,360
CNX MIDSTREAM PARTNERS LP LP 12654A101 646 45,800 SH   DFND 2 0 45,800 0
CNX MIDSTREAM PARTNERS LP LP 12654A101 42 3,000 SH   DFND 16 0 0 3,000
CVB FINANCIAL CORP COM 126600105 14,759 707,177 SH   DFND 2 0 707,177 0
CVB FINANCIAL CORP COM 126600105 80 3,852 SH   DFND 16 0 0 3,852
CVR ENERGY INC COM 12662P108 1,430 32,485 SH   DFND 2 0 32,485 0
CVR ENERGY INC COM 12662P108 374 8,498 SH   DFND 16 0 0 8,498
CVS HEALTH CORP COM 126650100 69,221 1,097,540 SH   DFND 2 0 1,097,540 0
CVS HEALTH CORP COM 126650100 296 4,701 SH   DFND 02,03 0 4,701 0
CVS HEALTH CORP COM 126650100 70,057 1,109,572 SH   DFND 16 0 300,918 808,653
CVS HEALTH CORP COM 126650100 82 1,306 SH   DFND 16,18 0 1,306 0
CVS HEALTH CORP COM 126650100 242 3,838 SH   DFND 18 0 3,740 98
C&J ENERGY SERVICES INC COM 12674R100 7,430 692,412 SH   DFND 2 0 692,412 0
CABLE ONE INC COM 12685J105 28,595 22,791 SH   DFND 2 0 22,421 370
CABLE ONE INC COM 12685J105 2,826 2,252 SH   DFND 16 0 2,223 29
CABLE ONE INC COM 12685J105 28,669 22,849 SH   DFND 17 0 22,849 0
CABLE ONE INC COM 12685J105 1 1 SH   DFND 18 0 0 1
CABOT CORP COM 127055101 7,115 157,020 SH   DFND 2 0 157,020 0
CABOT CORP COM 127055101 461 10,170 SH   DFND 16 0 9,625 545
CABOT OIL & GAS CORP COM 127097103 8,927 508,064 SH   DFND 2 0 508,064 0
CABOT OIL & GAS CORP COM 127097103 27 1,511 SH   DFND 02,03 0 1,511 0
CABOT OIL & GAS CORP COM 127097103 637 36,286 SH   DFND 16 0 27,935 8,351
CABOT MICROELECTRONICS CORP COM 12709P103 21,826 154,563 SH   DFND 2 0 153,563 1,000
CABOT MICROELECTRONICS CORP COM 12709P103 2,506 17,751 SH   DFND 16 0 17,562 189
CABOT MICROELECTRONICS CORP COM 12709P103 4 25 SH   DFND 18 0 0 25
CACI INTERNATIONAL INC -CL A COM 127190304 39,525 170,909 SH   DFND 2 0 165,622 5,287
CACI INTERNATIONAL INC -CL A COM 127190304 1,344 5,806 SH   DFND 16 0 5,391 415
CACI INTERNATIONAL INC -CL A COM 127190304 0 2 SH   DFND 18 0 2 0
CACTUS INC - A COM 127203107 1,543 53,339 SH   DFND 2 0 53,339 0
CADENCE DESIGN SYS INC COM 127387108 62,424 944,634 SH   DFND 2 0 944,634 0
CADENCE DESIGN SYS INC COM 127387108 67 1,015 SH   DFND 02,03 0 1,015 0
CADENCE DESIGN SYS INC COM 127387108 1,499 22,685 SH   DFND 16 0 0 22,685
CADENCE DESIGN SYS INC COM 127387108 28,202 426,787 SH   DFND 17 0 426,787 0
CADENCE BANCORP COM 12739A100 12,337 703,461 SH   DFND 2 0 703,461 0
CADENCE BANCORP COM 12739A100 78 4,419 SH   DFND 16 0 3,866 553
CADENCE BANCORP COM 12739A100 3 188 SH   DFND 18 0 188 0
CAESARS ENTERTAINMENT CORP COM 127686103 16,508 1,415,765 SH   DFND 2 0 1,415,765 0
CAESARS ENTERTAINMENT CORP COM 127686103 49 4,181 SH   DFND 16 0 0 4,181
CAESARS ENTERTAINMENT CORP CONV BND 127686AA1 21,908 12,900,000 PRN   DFND 2 0 0 12,900,000
CAESARS ENTERTAINMENT CORP CONV BND 127686AA1 17 10,150 PRN   DFND 16 0 0 10,150
CAL-MAINE FOODS INC COM 128030202 6,328 158,397 SH   DFND 2 0 158,397 0
CAL-MAINE FOODS INC COM 128030202 858 21,462 SH   DFND 16 0 21,281 181
CALAMOS CONVERTIBLE OPP&INC COM 128117108 2,933 281,789 SH   DFND 16 0 0 281,789
CALAMOS CONVERTIBLE OPP&INC COM 128117108 40 3,862 SH   DFND 18 0 3,862 0
CALAMOS GLOBAL TOTAL RETURN COM 128118106 130 10,738 SH   DFND 16 0 0 10,738
CALAMOS GLOBAL DYNAMIC INCOM COM 12811L107 1,981 251,198 SH   DFND 16 0 0 251,198
CALAMOS GLOBAL DYNAMIC INCOM COM 12811L107 8 917 SH   DFND 16,18 0 917 0
CALAMOS CVT AND HIGH INCOME COM 12811P108 3,370 306,327 SH   DFND 16 0 0 306,327
CALAMOS CVT AND HIGH INCOME COM 12811P108 42 3,860 SH   DFND 18 0 3,860 0
CALAMOS DYNAMIC CONVERTIBLE COM 12811V105 1,964 96,348 SH   DFND 16 0 0 96,348
CALAMOS DYNAMIC CONVERTIBLE COM 12811V105 81 4,000 SH   DFND 18 0 4,000 0
CALAMOS STRAT TOT RETURN FD COM 128125101 2,527 198,653 SH   DFND 16 0 0 198,653
CALAMOS STRAT TOT RETURN FD COM 128125101 25 1,992 SH   DFND 18 0 1,992 0
CALAMP CORP COM 128126109 2,080 180,600 SH   DFND 2 0 180,600 0
CALAMP CORP COM 128126109 133 11,577 SH   DFND 16 0 11,023 554
CALAVO GROWERS INC COM 128246105 8,301 87,208 SH   DFND 2 0 87,208 0
CALAVO GROWERS INC COM 128246105 2,400 25,224 SH   DFND 16 0 24,842 382
CALAVO GROWERS INC COM 128246105 4,725 49,635 SH   DFND 17 0 49,635 0
CALERES INC COM 129500104 5,275 225,314 SH   DFND 2 0 225,314 0
CALERES INC COM 129500104 149 6,356 SH   DFND 16 0 5,751 605
CALIFORNIA RESOURCES CORP COM 13057Q206 566 55,526 SH   DFND 2 0 55,526 0
CALIFORNIA RESOURCES CORP COM 13057Q206 30 2,926 SH   DFND 16 0 0 2,926
CALIFORNIA WATER SERVICE GRP COM 130788102 18,284 345,449 SH   DFND 2 0 345,449 0
CALIFORNIA WATER SERVICE GRP COM 130788102 371 7,005 SH   DFND 16 0 0 7,005
CALIX INC COM 13100M509 273 42,708 SH   DFND 2 0 42,708 0
CALLAWAY GOLF COMPANY COM 131193104 11,390 586,823 SH   DFND 2 0 586,823 0
CALLAWAY GOLF COMPANY COM 131193104 5 257 SH   DFND 16 0 0 257
CALLON PETROLEUM CO COM 13123X102 29,088 6,702,097 SH   DFND 2 0 6,452,820 249,277
CALLON PETROLEUM CO COM 13123X102 124 28,550 SH   DFND 02,03 0 28,550 0
CALLON PETROLEUM CO COM 13123X102 140 32,246 SH   DFND 16 0 26,924 5,322
CALLON PETROLEUM CO COM 13123X102 4,219 972,116 SH   DFND 17 0 972,116 0
CAMBREX CORP COM 132011107 10,654 179,047 SH   DFND 2 0 179,047 0
CAMBREX CORP COM 132011107 4 62 SH   DFND 16 0 0 62
CAMBREX CORP COM 132011107 2 30 SH   DFND 18 0 30 0
CAMDEN NATIONAL CORP COM 133034108 694 16,012 SH   DFND 2 0 16,012 0
CAMDEN NATIONAL CORP COM 133034108 28 651 SH   DFND 16 0 0 651
CAMDEN PROPERTY TRUST COM 133131102 40,274 362,738 SH   DFND 2 0 361,313 1,425
CAMDEN PROPERTY TRUST COM 133131102 93 835 SH   DFND 16 0 0 835
CAMDEN PROPERTY TRUST COM 133131102 0 4 SH   DFND 18 0 4 0
CAMECO CORP COM 13321L108 20,521 2,160,025 SH   DFND 2 0 2,160,025 0
CAMECO CORP COM 13321L108 9,814 1,033,059 SH   DFND 02,10 0 1,033,059 0
CAMECO CORP COM 13321L108 362 38,243 SH   DFND 16 0 0 38,243
CAMPBELL SOUP CO COM 134429109 2,329 49,642 SH   DFND 2 0 49,642 0
CAMPBELL SOUP CO COM 134429109 28 607 SH   DFND 02,03 0 607 0
CAMPBELL SOUP CO COM 134429109 878 18,712 SH   DFND 16 0 0 18,712
CAMPBELL SOUP CO COM 134429109 5 110 SH   DFND 18 0 110 0
CAMPING WORLD HOLDINGS INC-A COM 13462K109 688 77,310 SH   DFND 2 0 77,310 0
CAMPING WORLD HOLDINGS INC-A COM 13462K109 78 8,799 SH   DFND 16 0 0 8,799
CANADA GOOSE HOLDINGS INC COM 135086106 210,358 4,784,194 SH   DFND 2 0 4,147,833 636,361
CANADA GOOSE HOLDINGS INC COM 135086106 14,328 325,860 SH   DFND 02,03 0 325,860 0
CANADA GOOSE HOLDINGS INC COM 135086106 7,223 164,194 SH   DFND 10 0 164,194 0
CANADA GOOSE HOLDINGS INC COM 135086106 3,064 69,686 SH   DFND 16 0 46,368 23,318
CANADA GOOSE HOLDINGS INC COM 135086106 18 400 SH   DFND 16,18 0 400 0
CAN IMPERIAL BK OF COMMERCE COM 136069101 35 414 SH   DFND 2 0 414 0
CAN IMPERIAL BK OF COMMERCE COM 136069101 20,061 242,875 SH   DFND 16 0 232,746 10,128
CAN IMPERIAL BK OF COMMERCE COM 136069101 16 193 SH   DFND 18 0 0 193
CANADIAN NATL RAILWAY CO COM 136375102 10,741 119,542 SH   DFND 2 0 119,542 0
CANADIAN NATL RAILWAY CO COM 136375102 18,060 201,562 SH   DFND 16 0 71,997 129,565
CANADIAN NATL RAILWAY CO COM 136375102 117 1,301 SH   DFND 18 0 1,112 189
CANADIAN NATURAL RESOURCES COM 136385101 147,735 5,547,664 SH   DFND 2 0 5,373,609 174,055
CANADIAN NATURAL RESOURCES COM 136385101 5,395 202,577 SH   DFND 02,03 0 202,577 0
CANADIAN NATURAL RESOURCES COM 136385101 684 25,681 SH   DFND 16 0 11,550 14,131
CANADIAN NATURAL RESOURCES COM 136385101 5 181 SH   DFND 18 0 181 0
CANADIAN PACIFIC RAILWAY LTD COM 13645T100 4,439 19,955 SH   DFND 2 0 19,955 0
CANADIAN PACIFIC RAILWAY LTD COM 13645T100 1,073 4,802 SH   DFND 16 0 1,092 3,710
CANADIAN PACIFIC RAILWAY LTD COM 13645T100 19 86 SH   DFND 18 0 0 86
CANNAE HOLDINGS INC COM 13765N107 213 7,766 SH   DFND 2 0 7,766 0
CANNAE HOLDINGS INC COM 13765N107 0 2 SH   DFND 16 0 0 2
CANON INC-SPONS ADR ADR 138006309 397 14,879 SH   DFND 2 0 0 14,879
CANON INC-SPONS ADR ADR 138006309 703 26,352 SH   DFND 16 0 0 26,352
CANOPY GROWTH CORP COM 138035100 5,082 221,608 SH   DFND 16 0 0 221,608
CANTEL MEDICAL CORP COM 138098108 38,533 515,144 SH   DFND 2 0 515,144 0
CANTEL MEDICAL CORP COM 138098108 2,186 29,234 SH   DFND 16 0 27,837 1,397
CAPITAL CITY BANK GROUP INC COM 139674105 10,746 391,455 SH   DFND 2 0 391,455 0
CAPITAL ONE FINANCIAL CORP COM 14040H105 339,371 3,730,181 SH   DFND 2 0 2,467,546 1,262,635
CAPITAL ONE FINANCIAL CORP COM 14040H105 155 1,700 SH   DFND 02,03 0 1,700 0
CAPITAL ONE FINANCIAL CORP COM 14040H105 4,021 44,288 SH   DFND 16 0 786 43,502
CAPITAL ONE FINANCIAL CORP COM 14040H105 1 13 SH   DFND 16,18 0 13 0
CAPITAL ONE FINANCIAL CORP COM 14040H105 39 422 SH   DFND 18 0 371 51
CAPITOL FEDERAL FINANCIAL IN COM 14057J101 763 55,323 SH   DFND 2 0 55,323 0
CAPITOL FEDERAL FINANCIAL IN COM 14057J101 29 2,070 SH   DFND 16 0 0 2,070
CAPSTEAD MORTGAGE CORP COM 14067E506 3,839 522,020 SH   DFND 2 0 512,420 9,600
CAPSTEAD MORTGAGE CORP COM 14067E506 1 88 SH   DFND 16 0 0 88
CARBON BLACK INC COM 14081R103 7,252 279,037 SH   DFND 2 0 279,037 0
CARBON BLACK INC COM 14081R103 6 234 SH   DFND 16 0 0 234
CARBONITE INC COM 141337105 9,938 641,425 SH   DFND 2 0 635,625 5,800
CARBONITE INC COM 141337105 11 683 SH   DFND 16 0 0 683
CARDINAL HEALTH INC COM 14149Y108 242,870 5,146,063 SH   DFND 2 0 3,309,613 1,836,450
CARDINAL HEALTH INC COM 14149Y108 51 1,082 SH   DFND 02,03 0 1,082 0
CARDINAL HEALTH INC COM 14149Y108 6,878 145,780 SH   DFND 16 0 33,903 111,877
CARDINAL HEALTH INC COM 14149Y108 89 1,880 SH   DFND 18 0 1,880 0
CARDIOVASCULAR SYSTEMS INC COM 141619106 10,706 225,283 SH   DFND 2 0 225,283 0
CARDIOVASCULAR SYSTEMS INC COM 141619106 1,094 23,021 SH   DFND 16 0 0 23,021
CARE.COM INC COM 141633107 1,623 155,335 SH   DFND 2 0 155,335 0
CARE.COM INC COM 141633107 4 428 SH   DFND 16 0 0 428
CAREER EDUCATION CORP COM 141665109 6,386 401,888 SH   DFND 2 0 389,088 12,800
CAREER EDUCATION CORP COM 141665109 1 75 SH   DFND 16 0 0 75
CAREDX INC COM 14167L103 3,252 143,840 SH   DFND 2 0 143,840 0
CAREDX INC COM 14167L103 21 929 SH   DFND 16 0 0 929
CARETRUST REIT INC COM 14174T107 23,445 997,384 SH   DFND 2 0 970,684 26,700
CARETRUST REIT INC COM 14174T107 1,072 45,586 SH   DFND 16 0 2,418 43,168
CARGURUS INC COM 141788109 3 90 SH   DFND 2 0 90 0
CARGURUS INC COM 141788109 2,538 82,025 SH   DFND 16 0 78,505 3,520
CARLISLE COS INC COM 142339100 24,006 164,942 SH   DFND 2 0 160,292 4,650
CARLISLE COS INC COM 142339100 3,885 29,528 SH   DFND 16 0 823 28,705
CARLISLE COS INC COM 142339100 3 20 SH   DFND 18 0 0 20
CARLYLE GROUP/THE LP 14309L102 1,759 68,827 SH   DFND 16 0 0 68,827
CARMAX INC COM 143130102 5,714 64,934 SH   DFND 2 0 64,934 0
CARMAX INC COM 143130102 53 597 SH   DFND 02,03 0 597 0
CARMAX INC COM 143130102 839 9,541 SH   DFND 16 0 0 9,541
CARMAX INC COM 143130102 22 250 SH   DFND 18 0 250 0
CARNIVAL CORP COM 143658300 49,615 1,135,053 SH   DFND 2 0 1,134,709 344
CARNIVAL CORP COM 143658300 63 1,444 SH   DFND 02,03 0 1,444 0
CARNIVAL CORP COM 143658300 25,578 582,441 SH   DFND 16 0 213,697 368,744
CARNIVAL CORP COM 143658300 81 1,858 SH   DFND 16,18 0 1,858 0
CARNIVAL CORP COM 143658300 161 3,680 SH   DFND 18 0 3,680 0
CARNIVAL PLC-ADR ADR 14365C103 169 4,024 SH   DFND 2 0 0 4,024
CARNIVAL PLC-ADR ADR 14365C103 281 6,681 SH   DFND 16 0 0 6,681
CAROLINA FINANCIAL CORP COM 143873107 981 27,600 SH   DFND 2 0 27,600 0
CAROLINA FINANCIAL CORP COM 143873107 223 6,275 SH   DFND 16 0 6,075 200
CARRIAGE SERVICES INC COM 143905107 5,701 278,930 SH   DFND 2 0 278,930 0
CARRIAGE SERVICES INC COM 143905107 25 1,221 SH   DFND 16 0 0 1,221
CARPENTER TECHNOLOGY COM 144285103 27,666 535,530 SH   DFND 2 0 502,030 33,500
CARPENTER TECHNOLOGY COM 144285103 225 4,350 SH   DFND 02,03 0 4,350 0
CARPENTER TECHNOLOGY COM 144285103 1,133 21,779 SH   DFND 16 0 9,707 12,072
CARRIZO OIL & GAS INC COM 144577103 9,170 1,068,172 SH   DFND 2 0 1,068,172 0
CARRIZO OIL & GAS INC COM 144577103 6 649 SH   DFND 02,03 0 649 0
CARRIZO OIL & GAS INC COM 144577103 345 40,128 SH   DFND 16 0 39,663 465
CARS.COM INC COM 14575E105 844 94,012 SH   DFND 2 0 94,012 0
CARS.COM INC COM 14575E105 1 112 SH   DFND 16 0 0 112
CARS.COM INC COM 14575E105 7 775 SH   DFND 16,18 0 775 0
CARTER'S INC COM 146229109 22,085 242,130 SH   DFND 2 0 242,130 0
CARTER'S INC COM 146229109 1,019 11,168 SH   DFND 16 0 9,301 1,867
CARTER'S INC COM 146229109 30,118 330,200 SH   DFND 17 0 330,200 0
CARVANA CO COM 146869102 5 80 SH   DFND 2 0 80 0
CARVANA CO COM 146869102 2,679 40,611 SH   DFND 16 0 11,075 29,536
CASA SYSTEMS INC COM 14713L102 4,546 578,768 SH   DFND 2 0 578,768 0
CASA SYSTEMS INC COM 14713L102 61 7,759 SH   DFND 16 0 0 7,759
CASELLA WASTE SYSTEMS INC-A COM 147448104 12,826 298,728 SH   DFND 2 0 298,728 0
CASELLA WASTE SYSTEMS INC-A COM 147448104 8,782 204,601 SH   DFND 10 0 204,601 0
CASELLA WASTE SYSTEMS INC-A COM 147448104 1,514 35,272 SH   DFND 16 0 31,287 3,985
CASEY'S GENERAL STORES INC COM 147528103 16,698 103,610 SH   DFND 2 0 101,285 2,325
CASEY'S GENERAL STORES INC COM 147528103 5,812 36,101 SH   DFND 16 0 24,874 11,227
CASI PHARMACEUTICALS INC COM 14757U109 104 31,054 SH   DFND 16 0 0 31,054
CASS INFORMATION SYSTEMS INC COM 14808P109 1,081 19,978 SH   DFND 2 0 19,978 0
CASS INFORMATION SYSTEMS INC COM 14808P109 11 199 SH   DFND 16 0 0 199
CATALENT INC COM 148806102 16,044 336,625 SH   DFND 2 0 336,625 0
CATALENT INC COM 148806102 6,141 128,956 SH   DFND 10 0 128,956 0
CATALENT INC COM 148806102 986 20,680 SH   DFND 16 0 20,620 60
CATERPILLAR INC COM 149123101 96,006 760,081 SH   DFND 2 0 747,972 12,109
CATERPILLAR INC COM 149123101 1,373 10,872 SH   DFND 02,03 0 10,872 0
CATERPILLAR INC COM 149123101 105,723 836,816 SH   DFND 10 0 836,816 0
CATERPILLAR INC COM 149123101 23,631 186,925 SH   DFND 16 0 10,536 176,389
CATERPILLAR INC COM 149123101 31 247 SH   DFND 16,18 0 247 0
CATERPILLAR INC COM 149123101 551 4,365 SH   DFND 18 0 4,365 0
CATCHMARK TIMBER TRUST INC-A COM 14912Y202 57 5,350 SH   DFND 2 0 5,350 0
CATCHMARK TIMBER TRUST INC-A COM 14912Y202 488 45,729 SH   DFND 16 0 41,773 3,956
CATHAY GENERAL BANCORP COM 149150104 37,948 1,092,459 SH   DFND 2 0 1,027,159 65,300
CATHAY GENERAL BANCORP COM 149150104 236 6,800 SH   DFND 02,03 0 6,800 0
CATHAY GENERAL BANCORP COM 149150104 811 23,314 SH   DFND 16 0 23,242 72
CATO CORP-CLASS A COM 149205106 2,126 120,748 SH   DFND 2 0 116,548 4,200
CAVCO INDUSTRIES INC COM 149568107 15,712 81,795 SH   DFND 2 0 81,795 0
CAVCO INDUSTRIES INC COM 149568107 1 7 SH   DFND 16 0 0 7
CAVCO INDUSTRIES INC COM 149568107 6,444 33,549 SH   DFND 17 0 33,549 0
CEDAR FAIR LP LP 150185106 477 8,170 SH   DFND 2 0 8,170 0
CEDAR FAIR LP LP 150185106 1,386 23,747 SH   DFND 16 0 6,013 17,734
CEDAR FAIR LP LP 150185106 19,516 334,396 SH   DFND 17 0 334,396 0
CEDAR REALTY TRUST INC COM 150602209 1,672 557,406 SH   DFND 2 0 456,406 101,000
CEL-SCI CORP COM 150837607 102 11,420 SH   DFND 16 0 0 11,420
CELANESE CORP COM 150870103 59,460 486,235 SH   DFND 2 0 460,335 25,900
CELANESE CORP COM 150870103 54 444 SH   DFND 02,03 0 444 0
CELANESE CORP COM 150870103 3,101 25,367 SH   DFND 16 0 59 25,307
CELANESE CORP COM 150870103 7 61 SH   DFND 16,18 0 61 0
CELANESE CORP COM 150870103 3 24 SH   DFND 18 0 24 0
CELGENE CORP COM 151020104 33,169 334,029 SH   DFND 2 0 334,029 0
CELGENE CORP COM 151020104 255 2,566 SH   DFND 02,03 0 2,566 0
CELGENE CORP COM 151020104 19,495 196,316 SH   DFND 16 0 200 196,116
CELGENE CORP COM 151020104 21 212 SH   DFND 16,18 0 212 0
CELGENE CORP COM 151020104 256 2,581 SH   DFND 18 0 2,527 54
CEMEX SAB-SPONS ADR PART CER ADR 151290889 33 8,457 SH   DFND 2 0 0 8,457
CEMEX SAB-SPONS ADR PART CER ADR 151290889 315 80,487 SH   DFND 16 0 0 80,487
CENTENE CORP COM 15135B101 8,497 196,407 SH   DFND 2 0 196,407 0
CENTENE CORP COM 15135B101 65 1,498 SH   DFND 02,03 0 1,498 0
CENTENE CORP COM 15135B101 149,259 3,451,140 SH   DFND 10 0 3,382,092 69,048
CENTENE CORP COM 15135B101 59,033 1,365,169 SH   DFND 16 0 6,079 1,359,090
CENTENE CORP COM 15135B101 87 2,015 SH   DFND 16,18 0 2,015 0
CENTENE CORP COM 15135B101 264 6,092 SH   DFND 18 0 6,092 0
CENTENNIAL RESOURCE DEVELO-A COM 15136A102 637 141,076 SH   DFND 2 0 141,076 0
CENTENNIAL RESOURCE DEVELO-A COM 15136A102 12,740 2,824,770 SH   DFND 10 0 2,824,770 0
CENTERPOINT ENERGY INC COM 15189T107 12,694 420,633 SH   DFND 2 0 420,145 488
CENTERPOINT ENERGY INC COM 15189T107 55 1,815 SH   DFND 02,03 0 1,815 0
CENTERPOINT ENERGY INC COM 15189T107 5,579 184,931 SH   DFND 16 0 148,834 36,097
CENTERPOINT ENERGY INC COM 15189T107 11 350 SH   DFND 18 0 350 0
CENTERPOINT ENERGY INC CONV PRF 15189T503 53,958 1,031,100 SH   DFND 2 0 0 1,031,100
CENTERSTATE BANK CORP COM 15201P109 9,890 412,356 SH   DFND 2 0 412,356 0
CENTERSTATE BANK CORP COM 15201P109 33 1,388 SH   DFND 16 0 0 1,388
CENTERSTATE BANK CORP COM 15201P109 4 149 SH   DFND 18 0 0 149
CENTRAL GARDEN & PET CO COM 153527106 1,573 53,845 SH   DFND 2 0 53,845 0
CENTRAL GARDEN & PET CO COM 153527106 9 300 SH   DFND 16 0 0 300
CENTRAL GARDEN AND PET CO-A COM 153527205 11,470 413,670 SH   DFND 2 0 0 413,670
CENTRAL GARDEN AND PET CO-A COM 153527205 408 14,729 SH   DFND 16 0 0 14,729
CENTRAL PACIFIC FINANCIAL CO COM 154760409 8,480 298,625 SH   DFND 2 0 282,825 15,800
CENTRAL PACIFIC FINANCIAL CO COM 154760409 12 431 SH   DFND 16 0 0 431
CENTURY ALUMINUM COMPANY COM 156431108 8,812 1,328,182 SH   DFND 2 0 1,328,182 0
CENTURY COMMUNITIES INC COM 156504300 5,325 173,850 SH   DFND 2 0 173,850 0
CENTURY COMMUNITIES INC COM 156504300 0 14 SH   DFND 16 0 0 14
CENTURYLINK INC COM 156700106 7,287 583,972 SH   DFND 2 0 583,972 0
CENTURYLINK INC COM 156700106 44 3,547 SH   DFND 02,03 0 3,547 0
CENTURYLINK INC COM 156700106 6,947 555,461 SH   DFND 16 0 11,421 544,040
CENTURYLINK INC COM 156700106 6 457 SH   DFND 16,18 0 457 0
CENTURYLINK INC COM 156700106 39 3,153 SH   DFND 18 0 3,153 0
CERNER CORP COM 156782104 78,406 1,150,037 SH   DFND 2 0 1,150,037 0
CERNER CORP COM 156782104 79 1,152 SH   DFND 02,03 0 1,152 0
CERNER CORP COM 156782104 1,032 15,143 SH   DFND 16 0 2,315 12,828
CERNER CORP COM 156782104 22 332 SH   DFND 18 0 297 35
CEVA INC COM 157210105 3,432 114,940 SH   DFND 2 0 114,940 0
CHANGE HEALTHCARE INC COM 15912K100 384 31,781 SH   DFND 2 0 31,781 0
CHANGE HEALTHCARE INC COM 15912K100 31 2,587 SH   DFND 16 0 2,387 200
CHNG 6 06/30/22 CONV PRF 15912K209 16,146 335,000 SH   DFND 2 0 0 335,000
CHARLES RIVER LABORATORIES COM 159864107 35,917 271,337 SH   DFND 2 0 271,337 0
CHARLES RIVER LABORATORIES COM 159864107 53,828 406,521 SH   DFND 10 0 406,521 0
CHARLES RIVER LABORATORIES COM 159864107 167 1,261 SH   DFND 16 0 592 669
CHART INDUSTRIES INC COM 16115Q308 19,334 310,036 SH   DFND 2 0 310,036 0
CHART INDUSTRIES INC COM 16115Q308 371 5,945 SH   DFND 16 0 5,836 109
CHARTER COMMUNICATIONS INC-A COM 16119P108 65,750 159,540 SH   DFND 2 0 159,540 0
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EXLSERVICE HOLDINGS INC COM 302081104 12,231 182,649 SH   DFND 2 0 182,649 0
EXLSERVICE HOLDINGS INC COM 302081104 315 4,704 SH   DFND 16 0 4,244 460
EXPEDIA GROUP INC COM 30212P303 289,387 2,153,023 SH   DFND 2 0 1,485,944 667,079
EXPEDIA GROUP INC COM 30212P303 137 1,026 SH   DFND 02,03 0 1,026 0
EXPEDIA GROUP INC COM 30212P303 8,192 60,949 SH   DFND 16 0 54,955 5,994
EXPEDIA GROUP INC COM 30212P303 5 41 SH   DFND 18 0 15 26
EXPEDITORS INTL WASH INC COM 302130109 96,815 1,303,208 SH   DFND 2 0 1,303,208 0
EXPEDITORS INTL WASH INC COM 302130109 46 622 SH   DFND 02,03 0 622 0
EXPEDITORS INTL WASH INC COM 302130109 1,290 17,373 SH   DFND 16 0 12,534 4,839
EXPEDITORS INTL WASH INC COM 302130109 46,375 624,237 SH   DFND 17 0 624,237 0
EXPEDITORS INTL WASH INC COM 302130109 49 656 SH   DFND 18 0 656 0
EXPONENT INC COM 30214U102 35,321 505,320 SH   DFND 2 0 503,620 1,700
EXPONENT INC COM 30214U102 2,272 32,501 SH   DFND 16 0 32,235 266
EXPONENT INC COM 30214U102 43,634 624,240 SH   DFND 17 0 624,240 0
EXPRESS INC COM 30219E103 1,213 352,575 SH   DFND 2 0 352,575 0
EXTENDED STAY AMERICA INC COM 30224P200 34,463 2,353,966 SH   DFND 2 0 2,352,887 1,079
EXTENDED STAY AMERICA INC COM 30224P200 37,482 2,561,932 SH   DFND 10 0 2,561,932 0
EXTENDED STAY AMERICA INC COM 30224P200 1,649 112,649 SH   DFND 16 0 112,097 553
EXTENDED STAY AMERICA INC COM 30224P200 34,363 2,347,180 SH   DFND 17 0 2,347,180 0
EXTENDED STAY AMERICA INC COM 30224P200 0 18 SH   DFND 18 0 18 0
EXTRA SPACE STORAGE INC COM 30225T102 27,779 237,796 SH   DFND 2 0 237,796 0
EXTRA SPACE STORAGE INC COM 30225T102 54 462 SH   DFND 02,03 0 462 0
EXTRA SPACE STORAGE INC COM 30225T102 126 1,083 SH   DFND 10 0 1,083 0
EXTRA SPACE STORAGE INC COM 30225T102 3,082 32,524 SH   DFND 16 0 7,414 25,110
EXTRA SPACE STORAGE INC COM 30225T102 29 252 SH   DFND 16,18 0 252 0
EXTREME NETWORKS INC COM 30226D106 20,216 2,779,017 SH   DFND 2 0 2,673,902 105,115
EXTREME NETWORKS INC COM 30226D106 68 9,340 SH   DFND 16 0 0 9,340
EXTERRAN CORP COM 30227H106 9,544 730,792 SH   DFND 2 0 719,614 11,178
EXTERRAN CORP COM 30227H106 8 629 SH   DFND 16 0 0 629
EXTRACTION OIL & GAS INC COM 30227M105 0 110 SH   DFND 2 0 110 0
EXTRACTION OIL & GAS INC COM 30227M105 273 92,939 SH   DFND 16 0 0 92,939
EZCORP INC-CL A COM 302301106 9,403 1,456,825 SH   DFND 2 0 0 1,456,825
EXXON MOBIL CORP COM 30231G102 457,819 6,483,784 SH   DFND 2 0 6,256,625 227,159
EXXON MOBIL CORP COM 30231G102 1,720 24,364 SH   DFND 02,03 0 24,364 0
EXXON MOBIL CORP COM 30231G102 3,072 43,519 SH   DFND 10 0 43,519 0
EXXON MOBIL CORP COM 30231G102 108,242 1,537,046 SH   DFND 16 0 280,628 1,256,418
EXXON MOBIL CORP COM 30231G102 530 7,505 SH   DFND 16,18 0 7,505 0
EXXON MOBIL CORP COM 30231G102 1,587 22,476 SH   DFND 18 0 22,159 317
FBL FINANCIAL GROUP INC-CL A COM 30239F106 9,361 157,303 SH   DFND 2 0 157,303 0
FLIR SYSTEMS INC COM 302445101 3,002 57,089 SH   DFND 2 0 57,089 0
FLIR SYSTEMS INC COM 302445101 78 1,474 SH   DFND 02,03 0 1,474 0
FLIR SYSTEMS INC COM 302445101 3,578 68,011 SH   DFND 16 0 1,915 66,096
FLIR SYSTEMS INC COM 302445101 96 1,824 SH   DFND 16,18 0 1,824 0
FLIR SYSTEMS INC COM 302445101 20 375 SH   DFND 18 0 375 0
FMC CORP COM 302491303 346,381 3,950,332 SH   DFND 2 0 2,556,198 1,394,134
FMC CORP COM 302491303 41 473 SH   DFND 02,03 0 473 0
FMC CORP COM 302491303 3,317 38,196 SH   DFND 16 0 25,827 12,369
FMC CORP COM 302491303 10 114 SH   DFND 18 0 3 111
FNB CORP COM 302520101 13,384 1,160,835 SH   DFND 2 0 1,119,435 41,400
FNB CORP COM 302520101 70 6,076 SH   DFND 16 0 0 6,076
FB FINANCIAL CORP COM 30257X104 363 9,657 SH   DFND 2 0 9,657 0
FB FINANCIAL CORP COM 30257X104 10 267 SH   DFND 16 0 0 267