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Stock-based Compensation (Tables)
 9 Months Ended
Sep. 30, 2013
Stock-based Compensation [Abstract]  
Weighted average assumptions
The fair value of the stock options granted was estimated based on the following weighted average assumptions for three and nine months ended September 30, 2013 and 2012:

 
 
Three Months Ended September 30,
  
Nine Months Ended September 30,
 
 
 
2013
  
2012
  
2013
  
2012
 
Dividend yield
  
0
%
  
0
%
  
0
%
  
0
%
Expected volatility
  
92
%
  
92
%
  
92
%
  
92
%
Risk-free interest rate
  
1.43
%
  
0.72
%
  
0.87
%
  
1.05
%
Expected term
 
5.6 years
  
5.6 years
  
5.6 years
  
5.6 years
 
Stock option activity
Stock option activity for the nine months ended September 30, 2013 was as follows:

 
 
Options
  
Weighted Average Exercise Price Per Share
  
Weighted Average Remaining Contractual Life (in years)
  
Aggregate Intrinsic Value (in thousands)
 
Options outstanding at December 31, 2012
  
6,788,383
  
$
8.99
   
7.36
  
$
1,531
 
Granted
  
1,144,403
  
$
3.10
         
Exercised
  
(297,149
)
 
$
1.75
         
Forfeited, expired or cancelled
  
(129,296
)
 
$
17.88
         
Options outstanding at September 30, 2013
  
7,506,341
  
$
8.23
   
7.01
  
$
8,262
 
Options exercisable at September 30, 2013
  
4,817,306
  
$
11.13
   
6.03
  
$
4,050
 
Unvested RSU activity
 Unvested RSU activity for the nine months ended September 30, 2013 is summarized below:

 
 
  
Weighted-
 
 
   
Average
 
 
 
Number of
Shares
  
Grant-Date
 Fair Value
 
Unvested balance at December 31, 2012
  
1,459,853
  
$
2.75
 
Granted
  
958,385
  
$
2.96
 
Vested
  
(419,760
)
 
$
2.99
 
Forfeited
  
(45,434
)
 
$
2.29
 
Unvested balance at September 30, 2013
  
1,953,044
  
$
2.63
 
Stock-based compensation expense
 The following table shows total stock-based compensation expense included in the condensed consolidated statements of comprehensive loss for the three and nine months ended September 30, 2013 and 2012 (in thousands):

 
 
Three Months Ended September 30,
  
Nine Months Ended September 30,
 
 
 
2013
  
2012
  
2013
  
2012
 
Research and development
 
$
474
  
$
1,304
  
$
1,904
  
$
1,984
 
Selling, general and administrative
  
753
   
818
   
2,042
   
1,384
 
Total stock-based compensation expense
 
$
1,227
  
$
2,122
  
$
3,946
  
$
3,368