MML Conservative Allocation Fund —  Portfolio of Investments

March 31, 2022 (Unaudited)

	Number of Shares		Value
MUTUAL FUNDS — 100.1%
Equity Funds — 42.4%
Invesco Oppenheimer V.I. International Growth Fund, Series I		97,299	$	235,464
Invesco V.I. Discovery Mid Cap Growth Fund, Series I		5,720		544,517
Invesco V.I. Global Fund, Series I		2,417		116,104
Invesco V.I. Main Street Fund, Series I		33,026		1,111,646
MML Blue Chip Growth Fund, Initial Class (a)		972,982		18,292,064
MML Equity Income Fund, Initial Class (a)		1,106,661		14,043,525
MML Equity Index Fund, Class III (a)		9,212		308,863
MML Focused Equity Fund, Class II (a)		2,782,214		22,508,111
MML Foreign Fund, Initial Class (a)		691,071		7,124,941
MML Fundamental Equity Fund, Class II (a)		1,141,975		13,452,460
MML Fundamental Value Fund, Class II (a)		789,703		11,790,259
MML Global Fund, Class I (a)		470,583		6,479,926
MML Income & Growth Fund, Initial Class (a)		910,751		11,457,252
MML International Equity Fund, Class II (a)		1,401,100		14,263,198
MML Large Cap Growth Fund, Initial Class (a)		702,957		9,742,990
MML Mid Cap Growth Fund, Initial Class (a)		585,335		8,510,768
MML Mid Cap Value Fund, Initial Class (a)		576,559		7,431,848
MML Small Cap Growth Equity Fund, Initial Class (a)		64,801		905,828
MML Small Company Value Fund, Class II (a)		235,103		4,001,461
MML Small/Mid Cap Value Fund, Initial Class (a)		148,458		2,069,506
MML Strategic Emerging Markets Fund, Class II (a)		463,909		4,049,925
				158,440,656
Fixed Income Funds — 57.7%
Invesco V.I. Global Strategic Income Fund, Series I		2,344,806		9,918,530
MML Dynamic Bond Fund, Class II (a)		4,521,572		43,226,229
MML High Yield Fund, Class II (a)		314,630		3,042,469
MML Inflation-Protected and Income Fund, Initial Class (a)		1,181,104		12,685,061
MML Managed Bond Fund, Initial Class (a)		6,419,580		78,405,766
MML Short-Duration Bond Fund, Class II (a)		2,051,579		18,874,527
MML Total Return Bond Fund, Class II (a)		5,030,366		49,347,891
				215,500,473
TOTAL MUTUAL FUNDS (Cost $370,825,406)				373,941,129
TOTAL LONG-TERM INVESTMENTS (Cost $370,825,406)				373,941,129
TOTAL INVESTMENTS — 100.1% (Cost $370,825,406) (b)				373,941,129
Other Assets/(Liabilities) — (0.1)%				(288,459)
NET ASSETS — 100.0%			$	373,652,670

Notes to Portfolio of Investments

Percentages are stated as a percent of net assets.

(a) Affiliated issuer.

(b) See Note 3 for aggregate cost for federal tax purposes.

MML Balanced Allocation Fund —  Portfolio of Investments

March 31, 2022 (Unaudited)

	Number of Shares		Value
MUTUAL FUNDS — 100.1%
Equity Funds — 49.1%
Invesco Oppenheimer V.I. International Growth Fund, Series I		199,831	$	483,590
Invesco V.I. Discovery Mid Cap Growth Fund, Series I		4,848		461,498
Invesco V.I. Global Fund, Series I		3,822		183,556
Invesco V.I. Main Street Fund, Series I		16,712		562,523
MML Blue Chip Growth Fund, Initial Class (a)		1,214,978		22,841,580
MML Equity Income Fund, Initial Class (a)		1,477,250		18,746,299
MML Equity Index Fund, Class III (a)		16,739		561,274
MML Focused Equity Fund, Class II (a)		3,627,656		29,347,740
MML Foreign Fund, Initial Class (a)		937,135		9,661,867
MML Fundamental Equity Fund, Class II (a)		1,536,970		18,105,508
MML Fundamental Value Fund, Class II (a)		1,002,580		14,968,519
MML Global Fund, Class I (a)		894,513		12,317,446
MML Income & Growth Fund, Initial Class (a)		1,196,368		15,050,307
MML International Equity Fund, Class II (a)		2,038,731		20,754,281
MML Large Cap Growth Fund, Initial Class (a)		990,885		13,733,672
MML Mid Cap Growth Fund, Initial Class (a)		761,198		11,067,819
MML Mid Cap Value Fund, Initial Class (a)		845,087		10,893,175
MML Small Cap Growth Equity Fund, Initial Class (a)		101,081		1,412,959
MML Small Company Value Fund, Class II (a)		286,403		4,874,572
MML Small/Mid Cap Value Fund, Initial Class (a)		203,638		2,838,716
MML Strategic Emerging Markets Fund, Class II (a)		413,916		3,613,484
				212,480,385
Fixed Income Funds — 51.0%
Invesco V.I. Global Strategic Income Fund, Series I		2,178,596		9,215,463
MML Dynamic Bond Fund, Class II (a)		4,601,988		43,995,001
MML High Yield Fund, Class II (a)		334,555		3,235,143
MML Inflation-Protected and Income Fund, Initial Class (a)		1,383,093		14,854,417
MML Managed Bond Fund, Initial Class (a)		6,417,726		78,383,121
MML Short-Duration Bond Fund, Class II (a)		2,116,869		19,475,190
MML Total Return Bond Fund, Class II (a)		5,266,136		51,660,793
				220,819,128
TOTAL MUTUAL FUNDS (Cost $418,025,330)				433,299,513
TOTAL LONG-TERM INVESTMENTS (Cost $418,025,330)				433,299,513
TOTAL INVESTMENTS — 100.1% (Cost $418,025,330) (b)				433,299,513
Other Assets/(Liabilities) — (0.1)%				(332,426)
NET ASSETS — 100.0%			$	432,967,087

Notes to Portfolio of Investments

Percentages are stated as a percent of net assets.

(a) Affiliated issuer.

(b) See Note 3 for aggregate cost for federal tax purposes.

MML Moderate Allocation Fund —  Portfolio of Investments

March 31, 2022 (Unaudited)

	Number of Shares		Value
MUTUAL FUNDS — 100.1%
Equity Funds — 62.3%
Invesco Oppenheimer V.I. International Growth Fund, Series I		103,465	$	250,385
Invesco V.I. Discovery Mid Cap Growth Fund, Series I		5,555		528,803
Invesco V.I. Global Fund, Series I		6,317		303,382
MML Blue Chip Growth Fund, Initial Class (a)		5,734,857		107,815,309
MML Equity Income Fund, Initial Class (a)		6,614,455		83,937,431
MML Equity Index Fund, Class III (a)		73,137		2,452,268
MML Focused Equity Fund, Class II (a)		17,501,525		141,587,337
MML Foreign Fund, Initial Class (a)		4,084,579		42,112,006
MML Fundamental Equity Fund, Class II (a)		7,022,818		82,728,792
MML Fundamental Value Fund, Class II (a)		4,785,609		71,449,147
MML Global Fund, Class I (a)		4,168,508		57,400,359
MML Income & Growth Fund, Initial Class (a)		5,451,080		68,574,592
MML International Equity Fund, Class II (a)		9,832,161		100,091,399
MML Large Cap Growth Fund, Initial Class (a)		4,274,539		59,245,105
MML Mid Cap Growth Fund, Initial Class (a)		3,448,549		50,141,909
MML Mid Cap Value Fund, Initial Class (a)		3,692,955		47,602,185
MML Small Cap Growth Equity Fund, Initial Class (a)		666,798		9,320,866
MML Small Company Value Fund, Class II (a)		1,432,940		24,388,636
MML Small/Mid Cap Value Fund, Initial Class (a)		1,002,264		13,971,558
MML Strategic Emerging Markets Fund, Class II (a)		2,339,650		20,425,148
				984,326,617
Fixed Income Funds — 37.8%
Invesco V.I. Global Strategic Income Fund, Series I		4,069,674		17,214,722
MML Dynamic Bond Fund, Class II (a)		12,931,756		123,627,590
MML High Yield Fund, Class II (a)		1,182,293		11,432,772
MML Inflation-Protected and Income Fund, Initial Class (a)		3,324,045		35,700,245
MML Managed Bond Fund, Initial Class (a)		18,028,152		220,187,469
MML Short-Duration Bond Fund, Class II (a)		5,982,811		55,041,861
MML Total Return Bond Fund, Class II (a)		13,665,970		134,063,165
				597,267,824
TOTAL MUTUAL FUNDS (Cost $1,512,919,215)				1,581,594,441
TOTAL LONG-TERM INVESTMENTS (Cost $1,512,919,215)				1,581,594,441
TOTAL INVESTMENTS — 100.1% (Cost $1,512,919,215) (b)				1,581,594,441
Other Assets/(Liabilities) — (0.1)%				(1,221,590)
NET ASSETS — 100.0%			$	1,580,372,851

Notes to Portfolio of Investments

Percentages are stated as a percent of net assets.

(a) Affiliated issuer.

(b) See Note 3 for aggregate cost for federal tax purposes.

MML Growth Allocation Fund —  Portfolio of Investments

March 31, 2022 (Unaudited)

	Number of Shares		Value
MUTUAL FUNDS — 100.1%
Equity Funds — 75.4%
Invesco Oppenheimer V.I. International Growth Fund, Series I		181,923	$	440,253
Invesco V.I. Discovery Mid Cap Growth Fund, Series I		6,427		611,868
Invesco V.I. Global Fund, Series I		12,796		614,613
MML Blue Chip Growth Fund, Initial Class (a)		4,954,800		93,150,248
MML Equity Income Fund, Initial Class (a)		5,634,901		71,506,889
MML Equity Index Fund, Class III (a)		68,877		2,309,460
MML Focused Equity Fund, Class II (a)		13,569,460		109,776,930
MML Foreign Fund, Initial Class (a)		3,427,184		35,334,264
MML Fundamental Equity Fund, Class II (a)		6,223,060		73,307,644
MML Fundamental Value Fund, Class II (a)		4,052,929		60,510,225
MML Global Fund, Class I (a)		3,312,561		45,613,968
MML Income & Growth Fund, Initial Class (a)		4,728,701		59,487,065
MML International Equity Fund, Class II (a)		7,979,349		81,229,768
MML Large Cap Growth Fund, Initial Class (a)		4,173,004		57,837,833
MML Mid Cap Growth Fund, Initial Class (a)		2,960,099		43,039,846
MML Mid Cap Value Fund, Initial Class (a)		3,103,184		40,000,044
MML Small Cap Growth Equity Fund, Initial Class (a)		749,687		10,479,525
MML Small Company Value Fund, Class II (a)		1,457,252		24,802,426
MML Small/Mid Cap Value Fund, Initial Class (a)		794,573		11,076,349
MML Strategic Emerging Markets Fund, Class II (a)		1,666,804		14,551,203
				835,680,421
Fixed Income Funds — 24.7%
Invesco V.I. Global Strategic Income Fund, Series I		2,911,655		12,316,301
MML Dynamic Bond Fund, Class II (a)		5,951,159		56,893,081
MML High Yield Fund, Class II (a)		450,979		4,360,962
MML Inflation-Protected and Income Fund, Initial Class (a)		1,488,529		15,986,801
MML Managed Bond Fund, Initial Class (a)		7,705,711		94,113,971
MML Short-Duration Bond Fund, Class II (a)		2,832,013		26,054,519
MML Total Return Bond Fund, Class II (a)		6,476,690		63,536,332
				273,261,967
TOTAL MUTUAL FUNDS (Cost $1,037,330,968)				1,108,942,388
TOTAL LONG-TERM INVESTMENTS (Cost $1,037,330,968)				1,108,942,388
TOTAL INVESTMENTS — 100.1% (Cost $1,037,330,968) (b)				1,108,942,388
Other Assets/(Liabilities) — (0.1)%				(726,068)
NET ASSETS — 100.0%			$	1,108,216,320

Notes to Portfolio of Investments

Percentages are stated as a percent of net assets.

(a) Affiliated issuer.

(b) See Note 3 for aggregate cost for federal tax purposes.

MML Aggressive Allocation Fund —  Portfolio of Investments

March 31, 2022 (Unaudited)

	Number of Shares		Value
MUTUAL FUNDS — 100.1%
Equity Funds — 90.2%
Invesco Oppenheimer V.I. International Growth Fund, Series I		65,800	$	159,237
Invesco V.I. Discovery Mid Cap Growth Fund, Series I		20,004		1,904,385
Invesco V.I. Global Fund, Series I		12,710		610,463
Invesco V.I. Main Street Fund, Series I		8,277		278,599
MML Blue Chip Growth Fund, Initial Class (a)		857,795		16,126,550
MML Equity Income Fund, Initial Class (a)		903,654		11,467,371
MML Equity Index Fund, Class III (a)		9,921		332,646
MML Focused Equity Fund, Class II (a)		2,432,102		19,675,702
MML Foreign Fund, Initial Class (a)		550,471		5,675,360
MML Fundamental Equity Fund, Class II (a)		988,887		11,649,090
MML Fundamental Value Fund, Class II (a)		679,250		10,141,196
MML Global Fund, Class I (a)		212,565		2,927,027
MML Income & Growth Fund, Initial Class (a)		784,617		9,870,485
MML International Equity Fund, Class II (a)		1,322,081		13,458,786
MML Large Cap Growth Fund, Initial Class (a)		750,537		10,402,436
MML Mid Cap Growth Fund, Initial Class (a)		489,921		7,123,452
MML Mid Cap Value Fund, Initial Class (a)		523,404		6,746,681
MML Small Cap Growth Equity Fund, Initial Class (a)		146,753		2,051,393
MML Small Company Value Fund, Class II (a)		230,757		3,927,476
MML Small/Mid Cap Value Fund, Initial Class (a)		125,700		1,752,252
MML Strategic Emerging Markets Fund, Class II (a)		286,201		2,498,534
				138,779,121
Fixed Income Funds — 9.9%
Invesco V.I. Global Strategic Income Fund, Series I		269,981		1,142,020
MML Dynamic Bond Fund, Class II (a)		306,500		2,930,141
MML High Yield Fund, Class II (a)		63,439		613,452
MML Inflation-Protected and Income Fund, Initial Class (a)		33,553		360,364
MML Managed Bond Fund, Initial Class (a)		432,895		5,287,173
MML Short-Duration Bond Fund, Class II (a)		253,503		2,332,223
MML Total Return Bond Fund, Class II (a)		265,838		2,607,871
				15,273,244
TOTAL MUTUAL FUNDS (Cost $145,398,303)				154,052,365
TOTAL LONG-TERM INVESTMENTS (Cost $145,398,303)				154,052,365
TOTAL INVESTMENTS — 100.1% (Cost $145,398,303) (b)				154,052,365
Other Assets/(Liabilities) — (0.1)%				(109,334)
NET ASSETS — 100.0%			$	153,943,031

Notes to Portfolio of Investments

Percentages are stated as a percent of net assets.

(a) Affiliated issuer.

(b) See Note 3 for aggregate cost for federal tax purposes.

MML American Funds Growth Fund —  Portfolio of Investments

March 31, 2022 (Unaudited)

	Number of Shares		Value
MUTUAL FUNDS — 100.1%
Equity Funds — 100.1%
American Funds Insurance Series Growth Fund, Class 1		2,265,626	$	263,514,904
TOTAL MUTUAL FUNDS (Cost $187,177,515)				263,514,904
TOTAL INVESTMENTS — 100.1% (Cost $187,177,515) (a)				263,514,904
Other Assets/(Liabilities) — (0.1)%				(281,507)
NET ASSETS — 100.0%			$	263,233,397

Notes to Portfolio of Investments

Percentages are stated as a percent of net assets.

(a) See Note 3 for aggregate cost for federal tax purposes.

MML American Funds International Fund —  Portfolio of Investments

March 31, 2022 (Unaudited)

	Number of Shares		Value
MUTUAL FUNDS — 100.1%
Equity Funds — 100.1%
American Funds Insurance Series International Fund, Class 1		2,631,057	$	53,989,297
				53,989,297
TOTAL MUTUAL FUNDS (Cost $51,143,718)				53,989,297
TOTAL INVESTMENTS — 100.1% (Cost $51,143,718) (a)				53,989,297
Other Assets/(Liabilities) — (0.1)%				(78,922)
NET ASSETS — 100.0%			$	53,910,375

Notes to Portfolio of Investments

Percentages are stated as a percent of net assets.

(a) See Note 3 for aggregate cost for federal tax purposes.

MML American Funds Core Allocation Fund —  Portfolio of Investments

March 31, 2022 (Unaudited)

	Number of Shares		Value
MUTUAL FUNDS — 100.1%
Equity Funds — 66.5%
American Funds Insurance Series® Growth-Income Fund, Class 1		3,583,044	$	228,275,724
American Funds Insurance Series® International Fund, Class 1		4,279,886		87,823,250
American Funds Insurance Series® Washington Mutual Investors Fund, Class 1		14,166,851		251,178,276
				567,277,250
Fixed Income Funds — 33.6%
American Funds Insurance Series® The Bond Fund Of America, Class 1		27,019,459		287,216,848
TOTAL MUTUAL FUNDS (Cost $743,177,057)				854,494,098
TOTAL LONG-TERM INVESTMENTS (Cost $743,177,057)				854,494,098
TOTAL INVESTMENTS — 100.1% (Cost $743,177,057) (a)				854,494,098
Other Assets/(Liabilities) — (0.1)%				(899,390)
NET ASSETS — 100.0%			$	853,594,708

Notes to Portfolio of Investments

Percentages are stated as a percent of net assets.

(a) See Note 3 for aggregate cost for federal tax purposes.

MML Blue Chip Growth Fund —  Portfolio of Investments

March 31, 2022 (Unaudited)

	Number of Shares		Value
EQUITIES — 99.8%
COMMON STOCK — 99.6%
Basic Materials — 0.4%
Chemicals — 0.4%
Linde PLC		5,592	$	1,786,253
The Sherwin-Williams Co.		1,137		283,818
				2,070,071
				2,070,071
Communications — 32.9%
Internet — 32.3%
Alphabet, Inc. Class A (a)		3,545		9,859,886
Alphabet, Inc. Class C (a)		15,552		43,436,580
Amazon.com, Inc. (a)		17,977		58,604,121
Booking Holdings, Inc. (a)		1,439		3,379,419
DoorDash, Inc., Class A (a)		36,577		4,286,459
JD.COM, Inc., Class A, Lockup Shares (Acquired 1/20/22, Cost $101,577)) (a) (b) (c)		2,785		77,488
Meta Platforms, Inc. Class A (a)		121,380		26,990,057
Netflix, Inc. (a)		22,340		8,368,341
Opendoor Technologies, Inc. (a) (d)		95,590		826,853
Pinterest, Inc. Class A (a)		62,619		1,541,054
Sea Ltd. ADR (a)		55,657		6,667,152
Shopify, Inc. Class A (a)		5,496		3,715,076
Snap, Inc. Class A (a)		194,039		6,983,464
Spotify Technology SA (a)		9,282		1,401,768
Tencent Holdings Ltd.		58,500		2,759,761
				178,897,479
Media — 0.4%
The Walt Disney Co. (a)		16,050		2,201,418
Telecommunications — 0.2%
T-Mobile US, Inc. (a)		11,153		1,431,488
				182,530,385
Consumer, Cyclical — 10.5%
Apparel — 0.8%
NIKE, Inc. Class B		35,573		4,786,703
Auto Manufacturers — 5.1%
Rivian Automotive, Inc. Class A (a)		25,755		1,293,931
Rivian Automotive, Inc., Lockup Shares (Acquired 1/19/21-11/18/21, Cost $2,867,931) (a) (b) (c)		58,223		2,778,867

MML Blue Chip Growth Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
Tesla, Inc. (a)		22,355	$	24,089,748
				28,162,546
Retail — 4.6%
Carvana Co. (a) (d)		35,738		4,263,186
Chipotle Mexican Grill, Inc. (a)		3,106		4,913,785
Dollar General Corp.		20,823		4,635,825
Lululemon Athletica, Inc. (a)		14,228		5,196,492
Ross Stores, Inc.		46,946		4,246,735
Starbucks Corp.		13,384		1,217,543
The TJX Cos., Inc.		16,156		978,730
				25,452,296
				58,401,545
Consumer, Non-cyclical — 14.5%
Commercial Services — 2.9%
Adyen NV (a)		39,903		788,882
Affirm Holdings, Inc. (a) (d)		21,876		1,012,421
Ant International Co. Ltd., Class C (Acquired 6/07/18, Cost $775,038) (a) (b) (c) (e)		203,348		579,542
Block, Inc. (a)		25,990		3,524,244
Cintas Corp.		1,936		823,555
PayPal Holdings, Inc. (a)		34,151		3,949,563
S&P Global, Inc.		10,567		4,334,372
TransUnion		10,740		1,109,872
				16,122,451
Health Care – Products — 5.4%
Align Technology, Inc. (a)		2,774		1,209,464
Danaher Corp.		26,045		7,639,780
Intuitive Surgical, Inc. (a)		33,644		10,149,722
Stryker Corp.		19,217		5,137,665
Teleflex, Inc.		4,916		1,744,344
Thermo Fisher Scientific, Inc.		7,038		4,156,995
				30,037,970
Health Care – Services — 3.7%
HCA Healthcare, Inc.		13,525		3,389,635
Humana, Inc.		5,704		2,482,210
UnitedHealth Group, Inc.		28,238		14,400,533
				20,272,378
Pharmaceuticals — 2.5%
AstraZeneca PLC Sponsored ADR		25,228		1,673,625
Eli Lilly & Co.		29,715		8,509,485
Zoetis, Inc.		19,616		3,699,381
				13,882,491
				80,315,290

MML Blue Chip Growth Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
Financial — 7.9%
Banks — 1.1%
The Goldman Sachs Group, Inc.		18,373	$	6,064,927
Diversified Financial Services — 6.2%
The Charles Schwab Corp.		40,357		3,402,499
Mastercard, Inc. Class A		37,345		13,346,356
Visa, Inc. Class A		79,235		17,571,946
				34,320,801
Insurance — 0.6%
Chubb Ltd.		10,634		2,274,613
Marsh & McLennan Cos., Inc.		7,168		1,221,570
				3,496,183
				43,881,911
Industrial — 0.5%
Electronics — 0.2%
TE Connectivity Ltd.		7,686		1,006,712
Miscellaneous - Manufacturing — 0.3%
General Electric Co.		20,380		1,864,770
				2,871,482
Technology — 32.9%
Computers — 7.7%
Apple, Inc.		214,987		37,538,880
Crowdstrike Holdings, Inc. Class A (a)		3,172		720,298
Fortinet, Inc. (a)		13,243		4,525,663
				42,784,841
Semiconductors — 8.2%
Advanced Micro Devices, Inc. (a)		64,394		7,040,840
ASML Holding NV		7,964		5,319,394
Marvell Technology, Inc.		41,410		2,969,511
Monolithic Power Systems, Inc.		5,031		2,443,456
NVIDIA Corp.		81,974		22,367,426
Taiwan Semiconductor Manufacturing Co. Ltd. Sponsored ADR		20,203		2,106,365
Texas Instruments, Inc.		16,141		2,961,551
				45,208,543
Software — 17.0%
Atlassian Corp. PLC Class A (a)		13,294		3,906,176
Bill.com Holdings, Inc. (a)		17,872		4,053,191
Canva, Inc. (Acquired 8/16/21, Cost $582,930) (a) (b) (c) (e)		342		466,830
Confluent, Inc. Class A (a)		21,787		893,267
Datadog, Inc. Class A (a)		10,623		1,609,066
DocuSign, Inc. (a)		14,185		1,519,497
Gusto, Inc. (Acquired 10/04/21, Cost $193,055) (a) (b) (c) (e)		6,706		192,813
HashiCorp, Inc. Class A (a) (d)		4,850		261,900
Intuit, Inc.		20,695		9,950,984
Microsoft Corp.		120,955		37,291,636

MML Blue Chip Growth Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
MongoDB, Inc. (a)		9,585	$	4,251,810
MSCI, Inc.		1,430		719,118
Paycom Software, Inc. (a)		1,360		471,077
Roper Technologies, Inc.		6,896		3,256,498
ServiceNow, Inc. (a)		26,187		14,583,278
Snowflake, Inc. Class A (a)		6,690		1,532,880
Synopsys, Inc. (a)		20,681		6,892,357
Twilio, Inc. Class A (a)		3,347		551,619
Veeva Systems, Inc. Class A (a)		7,482		1,589,626
				93,993,623
				181,987,007
TOTAL COMMON STOCK (Cost $361,436,479)				552,057,691
PREFERRED STOCK — 0.2%
Technology — 0.2%
Software — 0.2%
Canva, Inc., Series A (Acquired 11/04/21-12/17/21, Cost $37,502) (a) (b) (c) (e)		22		30,030
Canva, Inc., Series A-3 (Acquired 12/17/21, Cost $3,408) (a) (b) (c) (e)		2		2,730
Databricks, Inc., Series G (Acquired 2/01/21, Cost $172,224) (a) (b) (c) (e)		971		161,069
Databricks, Inc., Series H (Acquired 8/31/21, Cost $542,974) (a) (b) (c) (e)		2,463		408,562
Gusto, Inc., Series E (Acquired 7/13/21, Cost $254,865) (a) (b) (c) (e)		8,385		254,865
				857,256
TOTAL PREFERRED STOCK (Cost $1,010,973)				857,256
TOTAL EQUITIES (Cost $362,447,452)				552,914,947
MUTUAL FUNDS — 0.2%
Diversified Financial Services — 0.2%
State Street Navigator Securities Lending Government Money Market Portfolio (f)		1,288,549		1,288,549
TOTAL MUTUAL FUNDS (Cost $1,288,549)				1,288,549
TOTAL LONG-TERM INVESTMENTS (Cost $363,736,001)				554,203,496
SHORT-TERM INVESTMENTS — 0.3%
Mutual Fund — 0.0%
T. Rowe Price Government Reserve Investment Fund		100		100
		Principal Amount
Repurchase Agreement — 0.3%
Fixed Income Clearing Corp., Repurchase Agreement, dated 3/31/22, 0.000%, due 4/01/22 (g)	$	1,773,199		1,773,199

MML Blue Chip Growth Fund —  Portfolio of Investments (Continued)

	Principal Amount	Value
TOTAL SHORT-TERM INVESTMENTS (Cost $1,773,299)		$	1,773,299
TOTAL INVESTMENTS — 100.3% (Cost $365,509,300) (h)			555,976,795
Other Assets/(Liabilities) — (0.3)%			(1,788,481)
NET ASSETS — 100.0%		$	554,188,314

Abbreviation Legend

ADR American Depositary Receipt

Notes to Portfolio of Investments

Percentages are stated as a percent of net assets.

(a) Non-income producing security.

(b) This security is fair valued in good faith in accordance with procedures approved by the Board of Trustees. At March 31, 2022, these securities amounted to a value of $4,952,796 or 0.89% of net assets.

(c) Restricted security. Certain securities are restricted as to resale. At March 31, 2022, these securities amounted to a value of $4,952,796 or 0.89% of net assets. The Fund generally bears the costs, if any, associated with the disposition of restricted securities.

(d) Denotes all or a portion of security on loan. The total value of securities on loan as of March 31, 2022, was $3,018,685 or 0.54% of net assets. Total securities on loan may be less than the amounts identified in the Portfolio of Investments. The Fund received $1,815,667 of non-cash collateral (U.S. Treasury and/or Agency obligations) related to securities lending activity. This amount is not reflected on the Portfolio of Investments. (Note 2).

(e) Investment was valued using significant unobservable inputs.

(f) Represents investment of security lending cash collateral. (Note 2).

(g) Maturity value of $1,773,199. Collateralized by U.S. Government Agency obligations with a rate of 0.500%, maturity date of 4/15/24, and an aggregate market value, including accrued interest, of $1,808,709.

(h) See Note 3 for aggregate cost for federal tax purposes.

MML Equity Income Fund —  Portfolio of Investments

March 31, 2022 (Unaudited)

	Number of Shares		Value
EQUITIES — 100.6%
COMMON STOCK — 99.1%
Basic Materials — 5.4%
Chemicals — 3.8%
Akzo Nobel NV		15,896	$	1,365,723
CF Industries Holdings, Inc.		95,814		9,874,591
DuPont de Nemours, Inc.		3,976		292,554
International Flavors & Fragrances, Inc.		34,563		4,539,159
RPM International, Inc.		9,524		775,634
				16,847,661
Forest Products & Paper — 1.6%
International Paper Co.		161,658		7,460,517
				24,308,178
Communications — 5.0%
Media — 4.1%
Comcast Corp. Class A		141,018		6,602,463
Fox Corp. Class B		50,826		1,843,967
News Corp. Class A		228,131		5,053,102
The Walt Disney Co. (a)		35,269		4,837,496
				18,337,028
Telecommunications — 0.9%
AT&T, Inc.		58,961		1,393,248
Cisco Systems, Inc.		39,227		2,187,298
Verizon Communications, Inc.		7,674		390,914
				3,971,460
				22,308,488
Consumer, Cyclical — 4.9%
Airlines — 0.3%
Southwest Airlines Co. (a)		27,804		1,273,423
Auto Manufacturers — 1.3%
PACCAR, Inc.		24,080		2,120,725
Volkswagen AG ADR		228,973		3,926,887
				6,047,612
Lodging — 0.7%
Las Vegas Sands Corp. (a)		81,537		3,169,343
Retail — 1.9%
Kohl's Corp.		52,915		3,199,241

MML Equity Income Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
The TJX Cos., Inc.		45,494	$	2,756,027
Walmart, Inc.		16,308		2,428,587
				8,383,855
Toys, Games & Hobbies — 0.7%
Mattel, Inc. (a)		140,770		3,126,502
				22,000,735
Consumer, Non-cyclical — 24.3%
Agriculture — 1.3%
Altria Group, Inc.		11,793		616,184
Philip Morris International, Inc.		54,821		5,149,885
				5,766,069
Beverages — 0.6%
The Coca-Cola Co.		42,276		2,621,112
Biotechnology — 0.3%
Biogen, Inc. (a)		4,200		884,520
Gilead Sciences, Inc.		8,536		507,465
				1,391,985
Commercial Services — 0.4%
Nielsen Holdings PLC		66,348		1,807,319
Food — 2.8%
Conagra Brands, Inc.		171,784		5,766,789
Mondelez International, Inc. Class A		4,800		301,344
Tyson Foods, Inc. Class A		72,730		6,518,790
				12,586,923
Health Care – Products — 2.1%
Medtronic PLC		55,920		6,204,324
Zimmer Biomet Holdings, Inc.		25,217		3,225,254
Zimvie, Inc.		2,521		57,580
				9,487,158
Health Care – Services — 3.7%
Anthem, Inc.		22,842		11,220,447
Centene Corp. (a)		35,373		2,978,053
UnitedHealth Group, Inc.		4,554		2,322,403
				16,520,903
Household Products & Wares — 1.4%
Kimberly-Clark Corp.		49,812		6,134,846
Pharmaceuticals — 11.7%
AbbVie, Inc.		62,078		10,063,465
AstraZeneca PLC Sponsored ADR		34,313		2,276,324
Becton Dickinson and Co.		28,294		7,526,204
Cardinal Health, Inc.		39,337		2,230,408
Cigna Corp.		20,398		4,887,565
CVS Health Corp.		60,024		6,075,029

MML Equity Income Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
GlaxoSmithKline PLC		25,169	$	542,743
Johnson & Johnson		32,113		5,691,387
Merck & Co., Inc.		42,685		3,502,304
Organon & Co.		2,021		70,594
Pfizer, Inc.		100,335		5,194,343
Sanofi		26,969		2,750,413
Sanofi Sponsored ADR		29,386		1,508,677
				52,319,456
				108,635,771
Energy — 6.8%
Oil & Gas — 5.5%
Chevron Corp.		5,786		942,134
EOG Resources, Inc.		38,606		4,602,993
Exxon Mobil Corp.		57,811		4,774,611
Hess Corp.		18,600		1,990,944
Occidental Petroleum Corp.		31,359		1,779,310
TotalEnergies SE (b)		163,037		8,272,291
TotalEnergies SE Sponsored ADR		45,415		2,295,274
				24,657,557
Pipelines — 1.3%
Enbridge, Inc.		7,500		345,675
Targa Resources Corp.		4,013		302,861
TC Energy Corp.		87,620		4,943,521
				5,592,057
				30,249,614
Financial — 26.4%
Banks — 10.9%
Bank of America Corp.		56,861		2,343,810
The Bank of New York Mellon Corp.		9,097		451,484
Citigroup, Inc.		42,340		2,260,956
Fifth Third Bancorp		127,702		5,496,294
The Goldman Sachs Group, Inc.		15,348		5,066,375
Huntington Bancshares, Inc.		349,692		5,112,497
JP Morgan Chase & Co.		18,115		2,469,437
Morgan Stanley		40,377		3,528,950
The PNC Financial Services Group, Inc.		7,730		1,425,798
State Street Corp.		60,308		5,254,033
Wells Fargo & Co.		319,236		15,470,177
				48,879,811
Diversified Financial Services — 0.7%
The Charles Schwab Corp.		31,005		2,614,031
Franklin Resources, Inc.		13,702		382,560
Raymond James Financial, Inc.		2,937		322,806
				3,319,397
Insurance — 10.2%
American International Group, Inc.		196,222		12,316,855
Chubb Ltd.		40,083		8,573,754

MML Equity Income Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
Equitable Holdings, Inc.		168,430	$	5,206,171
The Hartford Financial Services Group, Inc.		30,997		2,225,894
Loews Corp.		120,430		7,806,273
Marsh & McLennan Cos., Inc.		4,178		712,015
MetLife, Inc.		120,639		8,478,509
				45,319,471
Real Estate Investment Trusts (REITS) — 4.6%
Equity Residential		78,399		7,049,638
Rayonier, Inc.		89,064		3,662,312
Vornado Realty Trust		13,024		590,248
Welltower, Inc.		18,601		1,788,300
Weyerhaeuser Co.		196,290		7,439,391
				20,529,889
				118,048,568
Industrial — 9.9%
Aerospace & Defense — 2.6%
The Boeing Co. (a)		19,978		3,825,787
L3 Harris Technologies, Inc.		30,530		7,585,789
				11,411,576
Electronics — 0.3%
TE Connectivity Ltd.		10,077		1,319,885
Environmental Controls — 0.8%
Stericycle, Inc. (a)		63,366		3,733,525
Machinery – Diversified — 0.1%
Flowserve Corp.		12,573		451,371
Miscellaneous - Manufacturing — 3.4%
3M Co.		5,087		757,353
General Electric Co.		124,832		11,422,128
Siemens AG Sponsored ADR		44,104		3,049,350
				15,228,831
Transportation — 2.7%
United Parcel Service, Inc. Class B		55,394		11,879,797
				44,024,985
Technology — 7.6%
Semiconductors — 4.8%
Applied Materials, Inc.		30,693		4,045,337
NXP Semiconductor NV		5,781		1,069,948
QUALCOMM, Inc.		73,393		11,215,918
Texas Instruments, Inc.		26,471		4,856,899
				21,188,102
Software — 2.8%
Citrix Systems, Inc.		33,294		3,359,365
Fiserv, Inc. (a)		24,794		2,514,112

MML Equity Income Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
Microsoft Corp.		21,694	$	6,688,477
				12,561,954
				33,750,056
Utilities — 8.8%
Electric — 7.5%
Ameren Corp.		33,365		3,128,303
Dominion Energy, Inc.		58,498		4,970,575
NextEra Energy, Inc.		27,261		2,309,279
Sempra Energy		56,128		9,436,239
The Southern Co.		161,892		11,738,789
Xcel Energy, Inc.		26,900		1,941,373
				33,524,558
Gas — 1.3%
NiSource, Inc.		178,525		5,677,095
				39,201,653
TOTAL COMMON STOCK (Cost $319,297,851)				442,528,048
PREFERRED STOCK — 1.5%
Consumer, Non-cyclical — 0.4%
Pharmaceuticals — 0.4%
Becton Dickinson and Co. Convertible 6.000% (b)		31,874		1,683,585
Utilities — 1.1%
Electric — 0.8%
NextEra Energy, Inc. Convertible		26,869		1,401,218
Southern Co. Convertible 6.75%		35,696		1,949,359
				3,350,577
Gas — 0.3%
NiSource, Inc. 7.750%		13,065		1,554,474
				4,905,051
TOTAL PREFERRED STOCK (Cost $5,995,683)				6,588,636
TOTAL EQUITIES (Cost $325,293,534)				449,116,684

MML Equity Income Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
MUTUAL FUNDS — 2.1%
Diversified Financial Services — 2.1%
State Street Navigator Securities Lending Government Money Market Portfolio (c)		9,238,457	$	9,238,457
TOTAL MUTUAL FUNDS (Cost $9,238,457)				9,238,457
TOTAL LONG-TERM INVESTMENTS (Cost $334,531,991)				458,355,141
SHORT-TERM INVESTMENTS — 1.2%
Mutual Fund — 0.0%
T. Rowe Price Government Reserve Investment Fund		1,075		1,075
		Principal Amount
Repurchase Agreement — 1.2%
Fixed Income Clearing Corp., Repurchase Agreement, dated 3/31/22, 0.000%, due 4/01/22 (d)	$	5,600,854		5,600,854
TOTAL SHORT-TERM INVESTMENTS (Cost $5,601,929)				5,601,929
TOTAL INVESTMENTS — 103.9% (Cost $340,133,920) (e)				463,957,070
Other Assets/(Liabilities) — (3.9)%				(17,247,140)
NET ASSETS — 100.0%			$	446,709,930

Abbreviation Legend

ADR American Depositary Receipt

Notes to Portfolio of Investments

Percentages are stated as a percent of net assets.

(a) Non-income producing security.

(b) Denotes all or a portion of security on loan. The total value of securities on loan as of March 31, 2022, was $9,264,021 or 2.07% of net assets. Total securities on loan may be less than the amounts identified in the Portfolio of Investments. The Fund received $494,592 of non-cash collateral (U.S. Treasury and/or Agency obligations) related to securities lending activity. This amount is not reflected on the Portfolio of Investments. (Note 2).

(c) Represents investment of security lending cash collateral. (Note 2).

(d) Maturity value of $5,600,854. Collateralized by U.S. Government Agency obligations with a rate of 0.500%, maturity date of 4/15/24, and an aggregate market value, including accrued interest, of $5,712,875.

(e) See Note 3 for aggregate cost for federal tax purposes.

MML Equity Index Fund —  Portfolio of Investments

March 31, 2022 (Unaudited)

	Number of Shares		Value
EQUITIES — 99.6%
COMMON STOCK — 99.6%
Basic Materials — 2.1%
Chemicals — 1.6%
Air Products & Chemicals, Inc.		4,221	$	1,054,870
Albemarle Corp.		2,220		490,953
Celanese Corp.		2,062		294,598
CF Industries Holdings, Inc.		4,083		420,794
Dow, Inc.		14,060		895,903
DuPont de Nemours, Inc.		9,764		718,435
Eastman Chemical Co.		2,456		275,219
Ecolab, Inc.		4,744		837,601
FMC Corp.		2,413		317,478
International Flavors & Fragrances, Inc.		4,842		635,900
Linde PLC		9,757		3,116,679
LyondellBasell Industries NV Class A		4,998		513,894
The Mosaic Co.		7,097		471,951
PPG Industries, Inc.		4,516		591,912
The Sherwin-Williams Co.		4,591		1,146,005
				11,782,192
Forest Products & Paper — 0.0%
International Paper Co.		7,333		338,418
Iron & Steel — 0.1%
Nucor Corp.		5,178		769,710
Mining — 0.4%
Freeport-McMoRan, Inc.		27,919		1,388,691
Newmont Corp.		15,159		1,204,383
				2,593,074
				15,483,394
Communications — 14.3%
Advertising — 0.1%
The Interpublic Group of Cos., Inc.		7,539		267,257
Omnicom Group, Inc.		4,035		342,491
				609,748
Internet — 10.5%
Alphabet, Inc. Class A (a)		5,725		15,923,229
Alphabet, Inc. Class C (a)		5,288		14,769,331
Amazon.com, Inc. (a)		8,330		27,155,384
Booking Holdings, Inc. (a)		781		1,834,139
CDW Corp.		2,582		461,894
eBay, Inc.		11,904		681,623

MML Equity Index Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
Etsy, Inc. (a)		2,411	$	299,639
Expedia Group, Inc. (a)		2,860		559,616
F5, Inc. (a)		1,155		241,337
Match Group, Inc. (a)		5,382		585,239
Meta Platforms, Inc. Class A (a)		43,956		9,774,056
Netflix, Inc. (a)		8,451		3,165,660
NortonLifeLock, Inc.		11,138		295,380
Twitter, Inc. (a)		15,201		588,127
VeriSign, Inc. (a)		1,838		408,881
				76,743,535
Media — 1.6%
Charter Communications, Inc. Class A (a)		2,269		1,237,785
Comcast Corp. Class A		86,116		4,031,951
Discovery, Inc. Class A (a)		3,287		81,912
Discovery, Inc. Class C (a)		5,819		145,300
DISH Network Corp. Class A (a)		4,769		150,939
FactSet Research Systems, Inc.		720		312,588
Fox Corp. Class A		6,048		238,594
Fox Corp. Class B		2,839		102,999
News Corp. Class A		7,585		168,008
News Corp. Class B		2,302		51,841
Paramount Global Class B		11,502		434,891
The Walt Disney Co. (a)		34,658		4,753,691
				11,710,499
Telecommunications — 2.1%
Arista Networks, Inc. (a)		4,264		592,611
AT&T, Inc.		135,938		3,212,215
Cisco Systems, Inc.		80,287		4,476,803
Corning, Inc.		14,220		524,860
Juniper Networks, Inc.		6,229		231,470
Lumen Technologies, Inc.		17,705		199,535
Motorola Solutions, Inc.		3,210		777,462
T-Mobile US, Inc. (a)		11,177		1,434,568
Verizon Communications, Inc.		79,914		4,070,819
				15,520,343
				104,584,125
Consumer, Cyclical — 9.7%
Airlines — 0.2%
Alaska Air Group, Inc. (a)		2,429		140,906
American Airlines Group, Inc. (a) (b)		12,261		223,763
Delta Air Lines, Inc. (a)		12,183		482,082
Southwest Airlines Co. (a)		11,276		516,441
United Airlines Holdings, Inc. (a)		6,200		287,432
				1,650,624
Apparel — 0.6%
NIKE, Inc. Class B		24,296		3,269,270
PVH Corp.		1,367		104,726
Ralph Lauren Corp.		881		99,941
Tapestry, Inc.		5,025		186,679
Under Armour, Inc. Class A (a)		3,671		62,480

MML Equity Index Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
Under Armour, Inc. Class C (a)		4,132	$	64,294
VF Corp.		6,175		351,110
				4,138,500
Auto Manufacturers — 2.9%
Cummins, Inc.		2,721		558,104
Ford Motor Co.		74,877		1,266,170
General Motors Co. (a)		27,660		1,209,848
PACCAR, Inc.		6,595		580,822
Tesla, Inc. (a)		15,936		17,172,634
				20,787,578
Auto Parts & Equipment — 0.1%
Aptiv PLC (a)		5,142		615,549
BorgWarner, Inc.		4,594		178,706
				794,255
Distribution & Wholesale — 0.3%
Copart, Inc. (a)		4,058		509,157
Fastenal Co.		10,948		650,311
LKQ Corp.		5,095		231,364
Pool Corp.		759		320,943
W.W. Grainger, Inc.		821		423,464
				2,135,239
Entertainment — 0.1%
Caesars Entertainment, Inc. (a)		4,051		313,385
Live Nation Entertainment, Inc. (a)		2,554		300,453
Penn National Gaming, Inc. (a)		3,182		134,980
				748,818
Home Builders — 0.2%
D.R. Horton, Inc.		6,139		457,417
Lennar Corp. Class A		4,976		403,902
NVR, Inc. (a)		62		276,971
PulteGroup, Inc.		4,775		200,072
				1,338,362
Home Furnishing — 0.0%
Whirlpool Corp.		1,146		198,006
Housewares — 0.0%
Newell Brands, Inc.		7,253		155,287
Leisure Time — 0.1%
Carnival Corp. (a)		15,397		311,327
Norwegian Cruise Line Holdings Ltd. (a) (b)		7,936		173,640
Royal Caribbean Cruises Ltd. (a)		4,283		358,830
				843,797
Lodging — 0.4%
Hilton Worldwide Holdings, Inc. (a)		5,292		803,008
Las Vegas Sands Corp. (a)		6,587		256,037
Marriott International, Inc. Class A (a)		5,208		915,306

MML Equity Index Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
MGM Resorts International		7,170	$	300,710
Wynn Resorts Ltd. (a)		2,017		160,835
				2,435,896
Retail — 4.8%
Advance Auto Parts, Inc.		1,189		246,075
AutoZone, Inc. (a)		393		803,520
Bath & Body Works, Inc.		5,001		239,048
Best Buy Co., Inc.		4,121		374,599
CarMax, Inc. (a) (b)		3,082		297,351
Chipotle Mexican Grill, Inc. (a)		535		846,386
Costco Wholesale Corp.		8,441		4,860,750
Darden Restaurants, Inc.		2,463		327,456
Dollar General Corp.		4,411		982,021
Dollar Tree, Inc. (a)		4,277		684,962
Domino's Pizza, Inc.		690		280,837
Genuine Parts Co.		2,708		341,262
The Home Depot, Inc.		19,878		5,950,082
Lowe's Cos., Inc.		12,826		2,593,289
McDonald's Corp.		14,225		3,517,558
O'Reilly Automotive, Inc. (a)		1,282		878,119
Ross Stores, Inc.		6,759		611,419
Starbucks Corp.		21,897		1,991,970
Target Corp.		9,121		1,935,659
The TJX Cos., Inc.		22,708		1,375,651
Tractor Supply Co.		2,162		504,546
Ulta Beauty, Inc. (a)		1,034		411,759
Walgreens Boots Alliance, Inc.		13,655		611,334
Walmart, Inc.		26,930		4,010,416
Yum! Brands, Inc.		5,501		652,033
				35,328,102
Toys, Games & Hobbies — 0.0%
Hasbro, Inc.		2,449		200,622
				70,755,086
Consumer, Non-cyclical — 20.0%
Agriculture — 0.8%
Altria Group, Inc.		34,706		1,813,388
Archer-Daniels-Midland Co.		10,638		960,186
Philip Morris International, Inc.		29,503		2,771,512
				5,545,086
Beverages — 1.5%
Brown-Forman Corp. Class B		3,501		234,637
The Coca-Cola Co.		74,003		4,588,186
Constellation Brands, Inc. Class A		3,121		718,829
Molson Coors Beverage Co. Class B		3,609		192,648
Monster Beverage Corp. (a)		7,135		570,086
PepsiCo, Inc.		26,336		4,408,120
				10,712,506
Biotechnology — 1.5%
Amgen, Inc.		10,722		2,592,794

MML Equity Index Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
Bio-Rad Laboratories, Inc. Class A (a)		408	$	229,798
Biogen, Inc. (a)		2,792		587,995
Corteva, Inc.		13,857		796,500
Gilead Sciences, Inc.		23,842		1,417,407
Illumina, Inc. (a)		2,971		1,038,068
Incyte Corp. (a)		3,596		285,594
Moderna, Inc. (a)		6,707		1,155,348
Regeneron Pharmaceuticals, Inc. (a)		2,031		1,418,491
Vertex Pharmaceuticals, Inc. (a)		4,846		1,264,661
				10,786,656
Commercial Services — 1.8%
Automatic Data Processing, Inc.		8,014		1,823,506
Cintas Corp.		1,678		713,804
Equifax, Inc.		2,320		550,072
FleetCor Technologies, Inc. (a)		1,538		383,054
Gartner, Inc. (a)		1,564		465,227
Global Payments, Inc.		5,421		741,810
MarketAxess Holdings, Inc.		728		247,666
Moody's Corp.		3,076		1,037,873
Nielsen Holdings PLC		6,830		186,049
PayPal Holdings, Inc. (a)		22,177		2,564,770
Quanta Services, Inc.		2,713		357,058
Robert Half International, Inc.		2,099		239,664
Rollins, Inc.		4,290		150,364
S&P Global, Inc.		6,742		2,765,433
United Rentals, Inc. (a)		1,379		489,835
Verisk Analytics, Inc.		3,066		658,056
				13,374,241
Cosmetics & Personal Care — 1.3%
Colgate-Palmolive Co.		16,034		1,215,858
The Estee Lauder Cos., Inc. Class A		4,424		1,204,744
The Procter & Gamble Co.		45,631		6,972,417
				9,393,019
Food — 1.0%
Campbell Soup Co.		3,890		173,377
Conagra Brands, Inc.		9,132		306,561
General Mills, Inc.		11,531		780,879
The Hershey Co.		2,768		599,632
Hormel Foods Corp.		5,400		278,316
The J.M. Smucker Co.		2,076		281,111
Kellogg Co.		4,836		311,874
The Kraft Heinz Co.		13,464		530,347
The Kroger Co.		12,737		730,722
Lamb Weston Holdings, Inc.		2,780		166,550
McCormick & Co., Inc.		4,754		474,449
Mondelez International, Inc. Class A		26,429		1,659,213
Sysco Corp.		9,660		788,739
Tyson Foods, Inc. Class A		5,592		501,211
				7,582,981
Health Care – Products — 3.9%
Abbott Laboratories		33,662		3,984,234
ABIOMED, Inc. (a)		869		287,847

MML Equity Index Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
Align Technology, Inc. (a)		1,394	$	607,784
Baxter International, Inc.		9,518		738,026
Bio-Techne Corp.		745		322,615
Boston Scientific Corp. (a)		27,088		1,199,727
The Cooper Cos., Inc.		938		391,699
Danaher Corp.		12,114		3,553,400
Dentsply Sirona, Inc.		4,187		206,084
Edwards Lifesciences Corp. (a)		11,871		1,397,454
Henry Schein, Inc. (a)		2,640		230,182
Hologic, Inc. (a)		4,789		367,891
IDEXX Laboratories, Inc. (a)		1,613		882,408
Intuitive Surgical, Inc. (a)		6,810		2,054,441
Medtronic PLC		25,588		2,838,989
PerkinElmer, Inc.		2,399		418,529
ResMed, Inc.		2,784		675,148
Steris PLC		1,901		459,605
Stryker Corp.		6,383		1,706,495
Teleflex, Inc.		897		318,282
Thermo Fisher Scientific, Inc.		7,501		4,430,466
Waters Corp. (a)		1,161		360,363
West Pharmaceutical Services, Inc.		1,410		579,101
Zimmer Biomet Holdings, Inc.		3,962		506,740
				28,517,510
Health Care – Services — 2.4%
Anthem, Inc.		4,620		2,269,436
Catalent, Inc. (a)		3,409		378,058
Centene Corp. (a)		11,092		933,835
Charles River Laboratories International, Inc. (a)		966		274,315
DaVita, Inc. (a)		1,173		132,678
HCA Healthcare, Inc.		4,552		1,140,822
Humana, Inc.		2,444		1,063,556
IQVIA Holdings, Inc. (a)		3,631		839,524
Laboratory Corp. of America Holdings (a)		1,772		467,206
Molina Healthcare, Inc. (a)		1,110		370,285
Quest Diagnostics, Inc.		2,265		309,988
UnitedHealth Group, Inc.		17,929		9,143,252
Universal Health Services, Inc. Class B		1,405		203,655
				17,526,610
Household Products & Wares — 0.2%
Avery Dennison Corp.		1,564		272,089
Church & Dwight Co., Inc.		4,649		462,018
The Clorox Co.		2,343		325,747
Kimberly-Clark Corp.		6,409		789,332
				1,849,186
Pharmaceuticals — 5.6%
AbbVie, Inc.		33,654		5,455,650
AmerisourceBergen Corp.		2,866		443,399
Becton Dickinson and Co.		5,421		1,441,986
Bristol-Myers Squibb Co.		41,494		3,030,307
Cardinal Health, Inc.		5,318		301,530
Cigna Corp.		6,148		1,473,122
CVS Health Corp.		24,985		2,528,732
DexCom, Inc. (a)		1,843		942,879

MML Equity Index Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
Eli Lilly & Co.		15,114	$	4,328,196
Johnson & Johnson		50,115		8,881,881
McKesson Corp.		2,852		873,083
Merck & Co., Inc.		48,085		3,945,374
Organon & Co.		4,858		169,690
Pfizer, Inc.		106,848		5,531,521
Viatris, Inc.		23,170		252,090
Zoetis, Inc.		8,996		1,696,556
				41,295,996
				146,583,791
Energy — 4.0%
Energy – Alternate Sources — 0.1%
Enphase Energy, Inc. (a)		2,565		517,566
SolarEdge Technologies, Inc. (a)		1,000		322,370
				839,936
Oil & Gas — 3.3%
APA Corp.		6,980		288,483
Chevron Corp.		36,696		5,975,210
ConocoPhillips		24,790		2,479,000
Coterra Energy, Inc.		15,563		419,734
Devon Energy Corp.		11,953		706,781
Diamondback Energy, Inc.		3,261		447,018
EOG Resources, Inc.		11,128		1,326,791
Exxon Mobil Corp.		80,591		6,656,011
Hess Corp.		5,247		561,639
Marathon Oil Corp.		14,820		372,130
Marathon Petroleum Corp.		11,022		942,381
Occidental Petroleum Corp.		16,891		958,395
Phillips 66		8,895		768,439
Pioneer Natural Resources Co.		4,318		1,079,630
Valero Energy Corp.		7,768		788,763
				23,770,405
Oil & Gas Services — 0.3%
Baker Hughes Co.		17,241		627,745
Halliburton Co.		17,105		647,766
Schlumberger NV		26,715		1,103,597
				2,379,108
Pipelines — 0.3%
Kinder Morgan, Inc.		37,041		700,445
ONEOK, Inc.		8,456		597,247
The Williams Cos., Inc.		23,090		771,437
				2,069,129
				29,058,578
Financial — 14.8%
Banks — 4.6%
Bank of America Corp.		135,319		5,577,849
The Bank of New York Mellon Corp.		14,083		698,939

MML Equity Index Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
Citigroup, Inc.		37,776	$	2,017,238
Citizens Financial Group, Inc.		8,113		367,762
Comerica, Inc.		2,520		227,884
Fifth Third Bancorp		12,997		559,391
First Republic Bank		3,410		552,761
The Goldman Sachs Group, Inc.		6,462		2,133,106
Huntington Bancshares, Inc.		27,473		401,655
JP Morgan Chase & Co.		56,257		7,668,954
KeyCorp.		17,675		395,567
M&T Bank Corp.		2,450		415,275
Morgan Stanley		26,986		2,358,576
Northern Trust Corp.		3,947		459,628
The PNC Financial Services Group, Inc.		7,995		1,474,678
Regions Financial Corp.		18,067		402,172
Signature Bank		1,194		350,427
State Street Corp.		6,948		605,310
SVB Financial Group (a)		1,116		624,346
Truist Financial Corp.		25,375		1,438,763
US Bancorp		25,707		1,366,327
Wells Fargo & Co.		73,971		3,584,635
Zions Bancorp NA		2,886		189,206
				33,870,449
Diversified Financial Services — 3.7%
American Express Co.		11,709		2,189,583
Ameriprise Financial, Inc.		2,111		634,060
BlackRock, Inc.		2,715		2,074,722
Capital One Financial Corp.		7,879		1,034,434
Cboe Global Markets, Inc.		2,043		233,760
The Charles Schwab Corp.		28,619		2,412,868
CME Group, Inc.		6,833		1,625,297
Discover Financial Services		5,482		604,062
Franklin Resources, Inc.		5,395		150,628
Intercontinental Exchange, Inc.		10,712		1,415,269
Invesco Ltd.		6,541		150,835
Mastercard, Inc. Class A		16,429		5,871,396
Nasdaq, Inc.		2,221		395,782
Raymond James Financial, Inc.		3,557		390,950
Synchrony Financial		9,923		345,420
T. Rowe Price Group, Inc.		4,363		659,642
Visa, Inc. Class A		31,570		7,001,279
				27,189,987
Insurance — 3.8%
Aflac, Inc.		11,421		735,398
The Allstate Corp.		5,342		739,920
American International Group, Inc.		15,806		992,143
Aon PLC Class A		4,089		1,331,501
Arthur J Gallagher & Co.		3,968		692,813
Assurant, Inc.		1,095		199,104
Berkshire Hathaway, Inc. Class B (a)		34,863		12,303,501
Brown & Brown, Inc.		4,478		323,625
Chubb Ltd.		8,200		1,753,980
Cincinnati Financial Corp.		2,872		390,477
Everest Re Group Ltd.		756		227,843
Globe Life, Inc.		1,793		180,376
The Hartford Financial Services Group, Inc.		6,375		457,789

MML Equity Index Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
Lincoln National Corp.		3,171	$	207,257
Loews Corp.		3,733		241,973
Marsh & McLennan Cos., Inc.		9,600		1,636,032
MetLife, Inc.		13,357		938,730
Principal Financial Group, Inc.		4,668		342,678
The Progressive Corp.		11,121		1,267,683
Prudential Financial, Inc.		7,196		850,351
The Travelers Cos., Inc.		4,592		839,096
W.R. Berkley Corp.		3,989		265,594
Willis Towers Watson PLC		2,324		548,975
				27,466,839
Real Estate — 0.1%
CBRE Group, Inc. Class A (a)		6,360		582,067
Real Estate Investment Trusts (REITS) — 2.6%
Alexandria Real Estate Equities, Inc.		2,771		557,664
American Tower Corp.		8,669		2,177,826
AvalonBay Communities, Inc.		2,656		659,671
Boston Properties, Inc.		2,723		350,722
Crown Castle International Corp.		8,217		1,516,858
Digital Realty Trust, Inc.		5,395		765,011
Duke Realty Corp.		7,239		420,296
Equinix, Inc.		1,712		1,269,654
Equity Residential		6,486		583,221
Essex Property Trust, Inc.		1,242		429,086
Extra Space Storage, Inc.		2,541		522,430
Federal Realty Investment Trust		1,341		163,696
Healthpeak Properties, Inc.		10,266		352,432
Host Hotels & Resorts, Inc.		13,680		265,802
Iron Mountain, Inc.		5,546		307,304
Kimco Realty Corp.		11,755		290,349
Mid-America Apartment Communities, Inc.		2,189		458,486
Prologis, Inc.		14,087		2,274,769
Public Storage		2,901		1,132,202
Realty Income Corp.		10,748		744,836
Regency Centers Corp.		2,961		211,238
SBA Communications Corp.		2,068		711,599
Simon Property Group, Inc.		6,247		821,855
UDR, Inc.		5,693		326,607
Ventas, Inc.		7,569		467,461
Vornado Realty Trust		3,043		137,909
Welltower, Inc.		8,269		794,982
Weyerhaeuser Co.		14,232		539,393
				19,253,359
Savings & Loans — 0.0%
People's United Financial, Inc.		8,224		164,398
				108,527,099
Industrial — 7.6%
Aerospace & Defense — 1.6%
The Boeing Co. (a)		10,432		1,997,728
General Dynamics Corp.		4,388		1,058,298
Howmet Aerospace, Inc.		7,241		260,242
L3 Harris Technologies, Inc.		3,735		928,035

MML Equity Index Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
Lockheed Martin Corp.		4,614	$	2,036,620
Northrop Grumman Corp.		2,793		1,249,085
Raytheon Technologies Corp.		28,408		2,814,381
Teledyne Technologies, Inc. (a)		888		419,695
TransDigm Group, Inc. (a)		1,003		653,495
				11,417,579
Building Materials — 0.4%
Carrier Global Corp.		16,286		747,039
Fortune Brands Home & Security, Inc.		2,560		190,157
Johnson Controls International PLC		13,375		876,999
Martin Marietta Materials, Inc.		1,184		455,710
Masco Corp.		4,618		235,518
Mohawk Industries, Inc. (a)		1,054		130,907
Vulcan Materials Co.		2,518		462,556
				3,098,886
Electrical Components & Equipment — 0.3%
AMETEK, Inc.		4,395		585,326
Emerson Electric Co.		11,308		1,108,749
Generac Holdings, Inc. (a)		1,195		355,226
				2,049,301
Electronics — 1.0%
Agilent Technologies, Inc.		5,754		761,427
Allegion PLC		1,719		188,712
Amphenol Corp. Class A		11,402		859,140
Fortive Corp.		6,799		414,263
Garmin Ltd.		2,874		340,885
Honeywell International, Inc.		13,055		2,540,242
Keysight Technologies, Inc. (a)		3,505		553,685
Mettler-Toledo International, Inc. (a)		437		600,084
TE Connectivity Ltd.		6,203		812,469
Trimble, Inc. (a)		4,763		343,603
				7,414,510
Engineering & Construction — 0.0%
Jacobs Engineering Group, Inc.		2,461		339,150
Environmental Controls — 0.3%
Pentair PLC		3,178		172,279
Republic Services, Inc.		3,989		528,543
Waste Management, Inc.		7,329		1,161,646
				1,862,468
Hand & Machine Tools — 0.1%
Snap-on, Inc.		1,011		207,740
Stanley Black & Decker, Inc.		3,095		432,650
				640,390
Machinery – Construction & Mining — 0.3%
Caterpillar, Inc.		10,288		2,292,372

MML Equity Index Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
Machinery – Diversified — 0.7%
Deere & Co.		5,339	$	2,218,141
Dover Corp.		2,730		428,337
IDEX Corp.		1,456		279,159
Ingersoll Rand, Inc.		7,747		390,062
Nordson Corp.		1,027		233,211
Otis Worldwide Corp.		8,084		622,064
Rockwell Automation, Inc.		2,208		618,306
Westinghouse Air Brake Technologies Corp.		3,535		339,961
Xylem, Inc.		3,409		290,651
				5,419,892
Miscellaneous - Manufacturing — 1.1%
3M Co.		10,872		1,618,623
A.O. Smith Corp.		2,552		163,047
Eaton Corp. PLC		7,586		1,151,251
General Electric Co.		20,927		1,914,821
Illinois Tool Works, Inc.		5,440		1,139,136
Parker-Hannifin Corp.		2,456		696,915
Textron, Inc.		4,170		310,165
Trane Technologies PLC		4,446		678,904
				7,672,862
Packaging & Containers — 0.2%
Amcor PLC		29,121		329,941
Ball Corp.		6,167		555,030
Packaging Corp. of America		1,820		284,120
Sealed Air Corp.		2,797		187,287
WestRock Co.		5,011		235,668
				1,592,046
Shipbuilding — 0.0%
Huntington Ingalls Industries, Inc.		769		153,369
Transportation — 1.6%
C.H. Robinson Worldwide, Inc.		2,474		266,475
CSX Corp.		42,192		1,580,090
Expeditors International of Washington, Inc.		3,217		331,866
FedEx Corp.		4,646		1,075,038
J.B. Hunt Transport Services, Inc.		1,588		318,854
Norfolk Southern Corp.		4,564		1,301,744
Old Dominion Freight Line, Inc.		1,775		530,157
Union Pacific Corp.		12,124		3,312,398
United Parcel Service, Inc. Class B		13,880		2,976,705
				11,693,327
				55,646,152
Technology — 24.4%
Computers — 8.5%
Accenture PLC Class A		12,031		4,057,214
Apple, Inc.		295,128		51,532,300
Cognizant Technology Solutions Corp. Class A		9,986		895,445
DXC Technology Co. (a)		4,654		151,860
EPAM Systems, Inc. (a)		1,079		320,042

MML Equity Index Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
Fortinet, Inc. (a)		2,582	$	882,373
Hewlett Packard Enterprise Co.		24,622		411,434
HP, Inc.		20,611		748,179
International Business Machines Corp.		17,072		2,219,701
Leidos Holdings, Inc.		2,657		287,009
NetApp, Inc.		4,238		351,754
Seagate Technology Holdings PLC		3,834		344,677
Western Digital Corp. (a)		5,911		293,481
				62,495,469
Office & Business Equipment — 0.1%
Zebra Technologies Corp. Class A (a)		1,016		420,319
Semiconductors — 5.9%
Advanced Micro Devices, Inc. (a)		31,121		3,402,770
Analog Devices, Inc.		10,000		1,651,800
Applied Materials, Inc.		16,904		2,227,947
Broadcom, Inc.		7,860		4,949,285
Intel Corp.		77,516		3,841,693
IPG Photonics Corp. (a)		687		75,405
KLA Corp.		2,883		1,055,351
Lam Research Corp.		2,656		1,427,892
Microchip Technology, Inc.		10,584		795,282
Micron Technology, Inc.		21,316		1,660,303
Monolithic Power Systems, Inc.		822		399,229
NVIDIA Corp.		47,591		12,985,680
NXP Semiconductor NV		5,057		935,950
Qorvo, Inc. (a)		2,095		259,990
QUALCOMM, Inc.		21,454		3,278,600
Skyworks Solutions, Inc.		3,139		418,366
Teradyne, Inc.		3,099		366,395
Texas Instruments, Inc.		17,581		3,225,762
				42,957,700
Software — 9.9%
Activision Blizzard, Inc.		14,805		1,186,029
Adobe, Inc. (a)		8,979		4,091,012
Akamai Technologies, Inc. (a)		3,077		367,363
ANSYS, Inc. (a)		1,659		526,981
Autodesk, Inc. (a)		4,179		895,769
Broadridge Financial Solutions, Inc.		2,212		344,430
Cadence Design Systems, Inc. (a)		5,269		866,540
Ceridian HCM Holding, Inc. (a)		2,569		175,617
Cerner Corp.		5,593		523,281
Citrix Systems, Inc.		2,380		240,142
Electronic Arts, Inc.		5,375		679,991
Fidelity National Information Services, Inc.		11,573		1,162,161
Fiserv, Inc. (a)		11,297		1,145,516
Intuit, Inc.		5,385		2,589,323
Jack Henry & Associates, Inc.		1,394		274,688
Microsoft Corp.		142,712		43,999,537
MSCI, Inc.		1,547		777,955
Oracle Corp.		29,993		2,481,321
Paychex, Inc.		6,105		833,149
Paycom Software, Inc. (a)		921		319,016
PTC, Inc. (a)		2,024		218,025
Roper Technologies, Inc.		2,005		946,821

MML Equity Index Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
salesforce.com, Inc. (a)		18,751	$	3,981,212
ServiceNow, Inc. (a)		3,807		2,120,080
Synopsys, Inc. (a)		2,921		973,482
Take-Two Interactive Software, Inc. (a) (b)		2,187		336,229
Tyler Technologies, Inc. (a)		777		345,680
				72,401,350
				178,274,838
Utilities — 2.7%
Electric — 2.5%
AES Corp.		12,767		328,495
Alliant Energy Corp.		4,794		299,529
Ameren Corp.		4,927		461,956
American Electric Power Co., Inc.		9,582		955,996
CenterPoint Energy, Inc.		11,901		364,647
CMS Energy Corp.		5,516		385,789
Consolidated Edison, Inc.		6,732		637,386
Constellation Energy Corp.		6,201		348,806
Dominion Energy, Inc.		15,401		1,308,623
DTE Energy Co.		3,688		487,590
Duke Energy Corp.		14,629		1,633,474
Edison International		7,202		504,860
Entergy Corp.		3,826		446,685
Evergy, Inc.		4,393		300,218
Eversource Energy		6,528		575,704
Exelon Corp.		18,604		886,109
FirstEnergy Corp.		10,854		497,764
NextEra Energy, Inc.		37,352		3,164,088
NRG Energy, Inc.		4,690		179,908
Pinnacle West Capital Corp.		2,120		165,572
PPL Corp.		14,257		407,180
Public Service Enterprise Group, Inc.		9,611		672,770
Sempra Energy		6,072		1,020,825
The Southern Co.		20,164		1,462,092
WEC Energy Group, Inc.		5,992		598,062
Xcel Energy, Inc.		10,250		739,742
				18,833,870
Gas — 0.1%
Atmos Energy Corp.		2,578		308,045
NiSource, Inc.		7,517		239,041
				547,086
Water — 0.1%
American Water Works Co., Inc.		3,451		571,244
				19,952,200
TOTAL COMMON STOCK (Cost $353,445,005)				728,865,263
TOTAL EQUITIES (Cost $353,445,005)				728,865,263

MML Equity Index Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
MUTUAL FUNDS — 0.1%
Diversified Financial Services — 0.1%
State Street Navigator Securities Lending Government Money Market Portfolio (c)		385,515	$	385,515
TOTAL MUTUAL FUNDS (Cost $385,515)				385,515
TOTAL LONG-TERM INVESTMENTS (Cost $353,830,520)				729,250,778
	Principal Amount
SHORT-TERM INVESTMENTS — 0.3%
Repurchase Agreement — 0.2%
Fixed Income Clearing Corp., Repurchase Agreement, dated 3/31/22, 0.000%, due 4/01/22 (d)	$	2,086,609		2,086,609
U.S. Treasury Bill — 0.1%
U.S. Treasury Bill
0.315% 6/30/22 (e) (f)		860,000		858,893
TOTAL SHORT-TERM INVESTMENTS (Cost $2,945,943)				2,945,502
TOTAL INVESTMENTS — 100.0% (Cost $356,776,463) (g)				732,196,280
Other Assets/(Liabilities) — (0.0)%				(358,138)
NET ASSETS — 100.0%			$	731,838,142

Notes to Portfolio of Investments

Percentages are stated as a percent of net assets.

(a) Non-income producing security.

(b) Denotes all or a portion of security on loan. The total value of securities on loan as of March 31, 2022, was $1,022,732 or 0.14% of net assets. Total securities on loan may be less than the amounts identified in the Portfolio of Investments. The Fund received $660,747 of non-cash collateral (U.S. Treasury and/or Agency obligations) related to securities lending activity. This amount is not reflected on the Portfolio of Investments. (Note 2).

(c) Represents investment of security lending cash collateral. (Note 2).

(d) Maturity value of $2,086,609. Collateralized by U.S. Government Agency obligations with a rate of 1.750%, maturity date of 6/30/24, and an aggregate market value, including accrued interest, of $2,128,356.

(e) The rate shown represents yield-to-maturity.

(f) All or a portion of this security is pledged/held as collateral for open derivatives.

(g) See Note 3 for aggregate cost for federal tax purposes.

Futures contracts
	Expiration Date	Number of Contracts		Notional Amount		Value/ Net Unrealized Appreciation/ (Depreciation)
Long
S&P 500 E Mini Index	6/17/22		13	$	2,743,106	$	201,882

MML Focused Equity Fund —  Portfolio of Investments

March 31, 2022 (Unaudited)

	Number of Shares		Value
EQUITIES — 99.1%
COMMON STOCK — 99.1%
Basic Materials — 7.4%
Chemicals — 7.4%
Ecolab, Inc.		81,291	$	14,352,739
Linde PLC		35,045		11,194,424
				25,547,163
Consumer, Cyclical — 13.0%
Apparel — 4.2%
NIKE, Inc. Class B		106,872		14,380,696
Retail — 8.8%
McDonald's Corp.		52,832		13,064,297
The TJX Cos., Inc.		287,206		17,398,940
				30,463,237
				44,843,933
Consumer, Non-cyclical — 38.5%
Beverages — 7.8%
The Coca-Cola Co.		169,333		10,498,646
PepsiCo, Inc.		99,346		16,628,534
				27,127,180
Cosmetics & Personal Care — 8.8%
Colgate-Palmolive Co.		206,919		15,690,668
The Procter & Gamble Co.		95,994		14,667,883
				30,358,551
Health Care – Products — 12.0%
Baxter International, Inc.		132,586		10,280,719
Danaher Corp.		20,803		6,102,144
Medtronic PLC		117,275		13,011,661
Stryker Corp.		45,109		12,059,891
				41,454,415
Health Care – Services — 5.3%
UnitedHealth Group, Inc.		35,993		18,355,350
Pharmaceuticals — 4.6%
Johnson & Johnson		88,730		15,725,618

MML Focused Equity Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
			$	133,021,114
Financial — 19.7%
Diversified Financial Services — 8.6%
American Express Co.		68,545		12,817,915
Visa, Inc. Class A		76,669		17,002,884
				29,820,799
Insurance — 9.3%
Chubb Ltd.		73,380		15,695,982
Marsh & McLennan Cos., Inc.		96,935		16,519,663
				32,215,645
Real Estate Investment Trusts (REITS) — 1.8%
American Tower Corp.		24,495		6,153,634
				68,190,078
Industrial — 15.2%
Aerospace & Defense — 6.9%
Lockheed Martin Corp.		16,094		7,103,892
Northrop Grumman Corp.		37,346		16,701,878
				23,805,770
Electronics — 4.8%
Honeywell International, Inc.		84,669		16,474,894
Transportation — 3.5%
Union Pacific Corp.		44,950		12,280,789
				52,561,453
Technology — 5.3%
Computers — 2.0%
Accenture PLC Class A		20,613		6,951,322
Software — 3.3%
Microsoft Corp.		36,152		11,146,023
				18,097,345
TOTAL COMMON STOCK (Cost $282,480,226)				342,261,086
TOTAL EQUITIES (Cost $282,480,226)				342,261,086
TOTAL LONG-TERM INVESTMENTS (Cost $282,480,226)				342,261,086
	Principal Amount
SHORT-TERM INVESTMENTS — 0.8%
Repurchase Agreement — 0.8%
Fixed Income Clearing Corp., Repurchase Agreement, dated 3/31/22, 0.000%, due 4/01/22 (a)	$	2,853,574		2,853,574

MML Focused Equity Fund —  Portfolio of Investments (Continued)

	Principal Amount	Value
TOTAL SHORT-TERM INVESTMENTS (Cost $2,853,574)		$	2,853,574
TOTAL INVESTMENTS — 99.9% (Cost $285,333,800) (b)			345,114,660
Other Assets/(Liabilities) — 0.1%			406,421
NET ASSETS — 100.0%		$	345,521,081

Notes to Portfolio of Investments

Percentages are stated as a percent of net assets.

(a) Maturity value of $2,853,574. Collateralized by U.S. Government Agency obligations with a rate of 0.500%, maturity date of 4/15/24, and an aggregate market value, including accrued interest, of $2,910,659.

(b) See Note 3 for aggregate cost for federal tax purposes.

MML Foreign Fund —  Portfolio of Investments

March 31, 2022 (Unaudited)

	Number of Shares		Value
EQUITIES — 99.8%
COMMON STOCK — 99.8%
Australia — 4.1%
BHP Group Ltd. (a)		22,900	$	891,954
BHP Group Ltd. Sponsored ADR		10,700		826,575
Lendlease Corp Ltd.		165,200		1,373,002
Macquarie Group Ltd.		19,600		2,948,401
Santos Ltd.		522,500		3,004,480
				9,044,412
Austria — 0.3%
ams-OSRAM AG (b)		44,600		681,824
Belgium — 2.5%
Anheuser-Busch InBev SA		43,300		2,597,104
Groupe Bruxelles Lambert SA		14,900		1,545,201
KBC Group NV		19,100		1,373,664
				5,515,969
Canada — 0.3%
TFI International, Inc.		6,200		660,345
Cayman Islands — 2.7%
CK Asset Holdings Ltd.		346,000		2,364,813
CK Hutchison Holdings Ltd.		481,600		3,530,928
				5,895,741
Finland — 0.5%
Nokia OYJ (b)		216,100		1,189,606
France — 9.5%
Amundi SA (c)		21,400		1,460,192
Capgemini SE		11,000		2,446,875
Dassault Aviation SA		9,000		1,430,843
Engie SA (a)		295,200		3,869,602
Rexel SA		38,100		811,644
Sanofi		34,100		3,477,663
TotalEnergies SE (a)		52,200		2,648,562
Ubisoft Entertainment SA (b)		23,900		1,052,163
Veolia Environnement SA		113,512		3,622,095
				20,819,639
Germany — 11.9%
Allianz SE Registered		6,900		1,647,682
BASF SE		47,300		2,697,272
Bayer AG Registered		10,700		732,064
Deutsche Boerse AG		15,000		2,694,581

MML Foreign Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
Deutsche Post AG Registered		46,900	$	2,252,069
Fresenius SE & Co. KGaA		71,000		2,612,527
HeidelbergCement AG		43,100		2,464,941
Infineon Technologies AG		61,200		2,090,801
SAP SE		26,000		2,901,166
Siemens AG Registered		22,700		3,145,362
Siemens Energy AG (b)		87,700		2,003,826
Talanx AG (b)		20,700		918,635
				26,160,926
Ireland — 4.0%
AIB Group PLC		553,500		1,214,576
DCC PLC		31,000		2,402,652
Linde PLC		6,700		2,140,181
Smurfit Kappa Group PLC		65,900		2,920,869
				8,678,278
Israel — 1.1%
Check Point Software Technologies Ltd. (b)		17,800		2,461,028
Italy — 0.5%
Prysmian SpA		31,200		1,062,782
Japan — 22.9%
Astellas Pharma, Inc.		214,700		3,365,706
Denka Co. Ltd.		55,600		1,545,411
FANUC Corp.		12,200		2,145,473
Fujitsu Ltd.		13,500		2,012,610
Hitachi Ltd.		54,000		2,705,858
Japan Airlines Co. Ltd. (b)		52,800		982,723
Kirin Holdings Co. Ltd.		106,400		1,591,288
Kyocera Corp.		40,000		2,248,348
Nintendo Co. Ltd.		6,600		3,330,115
Olympus Corp.		98,900		1,878,941
ORIX Corp.		203,300		4,057,662
Rakuten Group, Inc.		269,600		2,125,376
SBI Holdings, Inc.		46,200		1,167,561
Sega Sammy Holdings, Inc.		75,500		1,303,163
Seven & i Holdings Co. Ltd.		81,900		3,896,820
Sony Group Corp.		56,100		5,790,136
Square Enix Holdings Co. Ltd.		27,000		1,195,211
Sumitomo Mitsui Financial Group, Inc.		92,100		2,939,300
Toshiba Corp.		77,000		2,921,819
Toyota Industries Corp.		45,500		3,131,764
				50,335,285
Luxembourg — 1.0%
ArcelorMittal SA		66,900		2,150,381
Netherlands — 5.8%
AerCap Holdings NV (b)		17,300		869,844
Airbus SE (a) (b)		4,600		556,018
ASML Holding NV		4,000		2,667,998
CNH Industrial NV		134,000		2,121,766
EXOR NV		10,400		793,229
Heineken Holding NV		35,400		2,770,643
Koninklijke Philips NV		35,779		1,094,587

MML Foreign Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
NXP Semiconductor NV		10,300	$	1,906,324
				12,780,409
Norway — 0.9%
Mowi ASA		77,100		2,078,480
Republic of Korea — 1.8%
Samsung Electronics Co. Ltd.		70,700		4,038,924
Singapore — 1.6%
DBS Group Holdings, Ltd.		134,500		3,536,051
Sweden — 2.6%
Essity AB Class B		49,700		1,171,005
Investor AB Class B		77,000		1,672,133
Lundin Energy AB (a)		29,900		1,267,597
Volvo AB Class B		84,298		1,572,285
				5,683,020
Switzerland — 9.8%
ABB Ltd. Registered		84,000		2,717,698
Alcon, Inc.		8,300		655,643
Cie Financiere Richemont SA Registered		17,800		2,257,903
Nestle SA Registered		26,200		3,401,306
Novartis AG Registered		43,900		3,850,094
Roche Holding AG		9,800		3,875,658
UBS Group AG Registered		238,300		4,655,668
				21,413,970
United Kingdom — 16.0%
Ashtead Group PLC		16,100		1,014,913
Aviva PLC		301,200		1,777,959
Barratt Developments PLC		135,100		921,750
BP PLC		394,800		1,929,400
The British Land Co. PLC		95,883		664,125
Bunzl PLC		46,100		1,792,380
Entain PLC (b)		40,700		872,954
GlaxoSmithKline PLC		97,000		2,091,702
Glencore PLC		295,400		1,919,736
Imperial Brands PLC		68,600		1,444,445
Inchcape PLC		101,900		889,528
Informa PLC (b)		215,300		1,689,727
Kingfisher PLC		179,400		598,721
Liberty Global PLC Class C (b)		35,500		919,805
Lloyds Banking Group PLC		4,554,600		2,788,315
Melrose Industries PLC		858,290		1,389,313
Persimmon PLC		61,100		1,718,516
Reckitt Benckiser Group PLC		34,900		2,667,651
Smith & Nephew PLC		144,800		2,308,054
Tesco PLC		710,805		2,569,801

MML Foreign Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
Unilever PLC		68,000	$	3,078,341
				35,047,136
TOTAL COMMON STOCK (Cost $207,614,987)				219,234,206
TOTAL EQUITIES (Cost $207,614,987)				219,234,206
MUTUAL FUNDS — 1.8%
United States — 1.8%
State Street Navigator Securities Lending Government Money Market Portfolio (d)		4,049,203		4,049,203
TOTAL MUTUAL FUNDS (Cost $4,049,203)				4,049,203
TOTAL LONG-TERM INVESTMENTS (Cost $211,664,190)				223,283,409
TOTAL INVESTMENTS — 101.6% (Cost $211,664,190) (e)				223,283,409
Other Assets/(Liabilities) — (1.6)%				(3,565,548)
NET ASSETS — 100.0%			$	219,717,861

Abbreviation Legend

ADR American Depositary Receipt

Notes to Portfolio of Investments

Percentages are stated as a percent of net assets.

(a) Denotes all or a portion of security on loan. The total value of securities on loan as of March 31, 2022, was $7,684,364 or 3.50% of net assets. Total securities on loan may be less than the amounts identified in the Portfolio of Investments. The Fund received $4,123,056 of non-cash collateral (U.S. Treasury and/or Agency obligations) related to securities lending activity. This amount is not reflected on the Portfolio of Investments. (Note 2).

(b) Non-income producing security.

(c) Security is exempt from registration under Regulation S or Rule 144A of the Securities Act of 1933. These securities are considered restricted and may be resold in transactions exempt from registration. At March 31, 2022, the aggregate market value of these securities amounted to $1,460,192 or 0.66% of net assets.

(d) Represents investment of security lending cash collateral. (Note 2).

(e) See Note 3 for aggregate cost for federal tax purposes.

MML Fundamental Equity Fund —  Portfolio of Investments

March 31, 2022 (Unaudited)

	Number of Shares		Value
EQUITIES — 99.8%
COMMON STOCK — 99.8%
Basic Materials — 0.7%
Chemicals — 0.7%
Valvoline, Inc.		51,103	$	1,612,811
Communications — 14.9%
Internet — 11.0%
Airbnb, Inc. Class A (a)		16,628		2,856,025
Alphabet, Inc. Class A (a)		2,332		6,486,108
Amazon.com, Inc. (a)		3,856		12,570,367
Netflix, Inc. (a)		6,059		2,269,641
Snap, Inc. Class A (a)		15,549		559,609
				24,741,750
Media — 1.3%
Comcast Corp. Class A		59,209		2,772,165
Telecommunications — 2.6%
Motorola Solutions, Inc.		9,821		2,378,647
Verizon Communications, Inc.		67,147		3,420,468
				5,799,115
				33,313,030
Consumer, Cyclical — 6.0%
Auto Manufacturers — 1.9%
General Motors Co. (a)		52,386		2,291,363
Tesla, Inc. (a)		1,943		2,093,777
				4,385,140
Entertainment — 0.3%
Warner Music Group Corp. Class A		15,273		578,083
Home Builders — 0.7%
D.R. Horton, Inc.		21,460		1,598,985
Retail — 3.1%
CarMax, Inc. (a) (b)		19,659		1,896,700
Floor & Decor Holdings, Inc. Class A (a)		6,929		561,249
O'Reilly Automotive, Inc. (a)		2,690		1,842,542
Target Corp.		12,662		2,687,130
				6,987,621
				13,549,829

MML Fundamental Equity Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
Consumer, Non-cyclical — 22.5%
Beverages — 1.3%
The Coca-Cola Co.		24,533	$	1,521,046
Constellation Brands, Inc. Class A		5,694		1,311,442
				2,832,488
Biotechnology — 0.9%
Seagen, Inc. (a)		13,664		1,968,299
Commercial Services — 1.0%
PayPal Holdings, Inc. (a)		11,436		1,322,573
TransUnion		9,559		987,827
				2,310,400
Cosmetics & Personal Care — 2.8%
Coty, Inc. Class A (a)		24,312		218,565
The Procter & Gamble Co.		40,000		6,112,000
				6,330,565
Food — 2.1%
Mondelez International, Inc. Class A		42,806		2,687,361
Sysco Corp.		25,073		2,047,210
				4,734,571
Health Care – Products — 2.0%
Avantor, Inc. (a)		57,198		1,934,437
The Cooper Cos., Inc.		6,258		2,613,278
				4,547,715
Health Care – Services — 5.8%
HCA Healthcare, Inc.		21,804		5,464,518
Tenet Healthcare Corp. (a)		19,096		1,641,492
UnitedHealth Group, Inc.		11,583		5,906,983
				13,012,993
Pharmaceuticals — 6.6%
AstraZeneca PLC Sponsored ADR		62,347		4,136,100
Bayer AG Registered		14,246		974,672
CVS Health Corp.		45,440		4,598,982
Eli Lilly & Co.		17,753		5,083,927
				14,793,681
				50,530,712
Energy — 3.7%
Oil & Gas — 2.3%
APA Corp.		37,001		1,529,251
Exxon Mobil Corp.		44,540		3,678,559
				5,207,810

MML Fundamental Equity Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
Oil & Gas Services — 0.3%
Nov, Inc.		29,039	$	569,455
Pipelines — 1.1%
Cheniere Energy, Inc.		10,634		1,474,404
Magellan Midstream Partners LP (c)		21,361		1,048,184
				2,522,588
				8,299,853
Financial — 13.5%
Banks — 3.9%
First Citizens BancShares, Inc. Class A		3,460		2,302,976
JP Morgan Chase & Co.		24,592		3,352,381
Signature Bank		5,389		1,581,618
SVB Financial Group (a)		2,864		1,602,265
				8,839,240
Diversified Financial Services — 3.6%
American Express Co.		17,739		3,317,193
Intercontinental Exchange, Inc.		27,306		3,607,669
Rocket Cos., Inc. Class A		103,374		1,149,519
				8,074,381
Insurance — 3.2%
The Allstate Corp.		23,433		3,245,705
Equitable Holdings, Inc.		126,537		3,911,258
				7,156,963
Real Estate Investment Trusts (REITS) — 2.8%
Prologis, Inc.		38,697		6,248,791
				30,319,375
Industrial — 11.3%
Aerospace & Defense — 1.9%
Raytheon Technologies Corp.		43,714		4,330,746
Building Materials — 1.3%
Vulcan Materials Co.		15,275		2,806,017
Environmental Controls — 0.6%
Waste Connections, Inc.		10,427		1,456,652
Machinery – Diversified — 3.3%
Deere & Co.		7,356		3,056,124
Otis Worldwide Corp.		40,699		3,131,788
Rockwell Automation, Inc.		3,958		1,108,359
				7,296,271
Transportation — 4.2%
Union Pacific Corp.		9,312		2,544,131

MML Fundamental Equity Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
United Parcel Service, Inc. Class B		32,176	$	6,900,465
				9,444,596
				25,334,282
Technology — 25.4%
Computers — 6.0%
Accenture PLC Class A		9,046		3,050,582
Amdocs Ltd.		6,708		551,465
Apple, Inc.		56,488		9,863,370
				13,465,417
Semiconductors — 5.1%
Advanced Micro Devices, Inc. (a)		25,180		2,753,181
Applied Materials, Inc.		22,169		2,921,875
QUALCOMM, Inc.		37,338		5,705,993
				11,381,049
Software — 14.3%
Doximity, Inc. Class A (a)		13,285		692,016
Fiserv, Inc. (a)		29,357		2,976,800
Manhattan Associates, Inc. (a)		4,749		658,734
Microsoft Corp.		49,149		15,153,128
salesforce.com, Inc. (a)		20,482		4,348,738
ServiceNow, Inc. (a)		2,219		1,235,739
VMware, Inc. Class A		39,220		4,465,981
Workday, Inc. Class A (a)		9,075		2,173,099
Zynga, Inc. Class A (a)		48,241		445,747
				32,149,982
				56,996,448
Utilities — 1.8%
Electric — 1.8%
FirstEnergy Corp.		88,738		4,069,525
TOTAL COMMON STOCK (Cost $192,023,108)				224,025,865
TOTAL EQUITIES (Cost $192,023,108)				224,025,865
TOTAL LONG-TERM INVESTMENTS (Cost $192,023,108)				224,025,865
	Principal Amount
SHORT-TERM INVESTMENTS — 0.3%
Repurchase Agreement — 0.3%
Fixed Income Clearing Corp., Repurchase Agreement, dated 3/31/22, 0.000%, due 4/01/22 (d)	$	648,435		648,435
TOTAL SHORT-TERM INVESTMENTS (Cost $648,435)				648,435

MML Fundamental Equity Fund —  Portfolio of Investments (Continued)

	Principal Amount	Value
TOTAL INVESTMENTS — 100.1% (Cost $192,671,543) (e)		$	224,674,300
Other Assets/(Liabilities) — (0.1)%			(333,525)
NET ASSETS — 100.0%		$	224,340,775

Abbreviation Legend

ADR American Depositary Receipt

Notes to Portfolio of Investments

Percentages are stated as a percent of net assets.

(a) Non-income producing security.

(b) Denotes all or a portion of security on loan. The total value of securities on loan as of March 31, 2022, was $1,995,978 or 0.89% of net assets. Total securities on loan may be less than the amounts identified in the Portfolio of Investments. The Fund received $2,036,381 of non-cash collateral (U.S. Treasury and/or Agency obligations) related to securities lending activity. This amount is not reflected on the Portfolio of Investments. (Note 2).

(c) Security is a Master Limited Partnership

(d) Maturity value of $648,435. Collateralized by U.S. Government Agency obligations with a rate of 0.500%, maturity date of 4/15/24, and an aggregate market value, including accrued interest, of $661,476.

(e) See Note 3 for aggregate cost for federal tax purposes.

MML Fundamental Value Fund —  Portfolio of Investments

March 31, 2022 (Unaudited)

	Number of Shares		Value
EQUITIES — 99.4%
COMMON STOCK — 99.4%
Basic Materials — 2.4%
Chemicals — 2.4%
Axalta Coating Systems Ltd. (a)		53,490	$	1,314,784
DuPont de Nemours, Inc.		43,487		3,199,774
				4,514,558
Communications — 7.7%
Internet — 4.4%
Alphabet, Inc. Class A (a)		2,001		5,565,482
Meta Platforms, Inc. Class A (a)		8,095		1,800,004
NortonLifeLock, Inc.		45,991		1,219,681
				8,585,167
Telecommunications — 3.3%
Cisco Systems, Inc.		74,933		4,178,264
T-Mobile US, Inc. (a)		16,440		2,110,074
				6,288,338
				14,873,505
Consumer, Cyclical — 5.2%
Distribution & Wholesale — 0.7%
LKQ Corp.		29,225		1,327,107
Home Furnishing — 0.6%
Sony Group Corp. Sponsored ADR (b)		11,428		1,173,770
Retail — 3.9%
AutoZone, Inc. (a)		2,270		4,641,196
Kohl's Corp.		12,254		740,877
Restaurant Brands International, Inc.		12,094		706,169
The TJX Cos., Inc.		24,467		1,482,211
				7,570,453
				10,071,330
Consumer, Non-cyclical — 28.5%
Beverages — 2.4%
Coca-Cola Europacific Partners PLC		33,636		1,635,046
Keurig Dr Pepper, Inc.		76,416		2,896,166
				4,531,212

MML Fundamental Value Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
Commercial Services — 2.9%
FleetCor Technologies, Inc. (a)		6,394	$	1,592,490
Global Payments, Inc.		13,710		1,876,076
United Rentals, Inc. (a)		6,102		2,167,492
				5,636,058
Cosmetics & Personal Care — 1.8%
The Procter & Gamble Co.		22,669		3,463,823
Food — 0.8%
US Foods Holding Corp. (a)		41,071		1,545,502
Health Care – Products — 1.0%
Avantor, Inc. (a)		56,283		1,903,491
Health Care – Services — 5.0%
Centene Corp. (a)		38,744		3,261,857
ICON PLC (a)		7,022		1,707,891
UnitedHealth Group, Inc.		9,103		4,642,257
				9,612,005
Pharmaceuticals — 14.6%
AbbVie, Inc.		17,051		2,764,138
AmerisourceBergen Corp.		11,185		1,730,431
Bristol-Myers Squibb Co.		32,677		2,386,401
Cigna Corp.		15,349		3,677,774
CVS Health Corp.		33,220		3,362,196
Johnson & Johnson		40,267		7,136,520
McKesson Corp.		5,729		1,753,819
Novartis AG Sponsored ADR		20,078		1,761,845
Sanofi Sponsored ADR		67,098		3,444,811
				28,017,935
				54,710,026
Energy — 11.3%
Oil & Gas — 10.1%
Canadian Natural Resources Ltd. (b)		44,190		2,738,896
Cenovus Energy, Inc.		68,619		1,144,565
ConocoPhillips		61,242		6,124,200
Devon Energy Corp.		17,057		1,008,580
EOG Resources, Inc.		14,599		1,740,639
Marathon Petroleum Corp.		37,533		3,209,072
Pioneer Natural Resources Co.		13,296		3,324,399
				19,290,351
Oil & Gas Services — 1.2%
Schlumberger NV		56,346		2,327,653
				21,618,004
Financial — 16.9%
Banks — 8.7%
Bank of America Corp.		70,420		2,902,713
The Goldman Sachs Group, Inc.		6,493		2,143,339

MML Fundamental Value Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
JP Morgan Chase & Co.		37,966	$	5,175,525
Truist Financial Corp.		42,371		2,402,436
Wells Fargo & Co.		85,300		4,133,638
				16,757,651
Diversified Financial Services — 2.8%
Capital One Financial Corp.		17,626		2,314,118
The Charles Schwab Corp.		35,408		2,985,248
				5,299,366
Insurance — 5.4%
Berkshire Hathaway, Inc. Class B (a)		18,415		6,498,837
Chubb Ltd.		11,171		2,389,477
Everest Re Group Ltd.		5,142		1,549,696
				10,438,010
				32,495,027
Industrial — 13.1%
Aerospace & Defense — 2.7%
General Dynamics Corp.		12,786		3,083,727
Howmet Aerospace, Inc.		59,768		2,148,062
				5,231,789
Building Materials — 1.5%
CRH PLC Sponsored ADR		26,104		1,045,727
Mohawk Industries, Inc. (a)		9,176		1,139,659
Owens Corning		7,728		707,112
				2,892,498
Electronics — 0.5%
Allegion PLC		7,767		852,661
Machinery – Construction & Mining — 0.8%
Caterpillar, Inc.		7,068		1,574,892
Machinery – Diversified — 4.2%
Deere & Co.		7,120		2,958,075
Dover Corp.		6,585		1,033,186
The Middleby Corp. (a)		5,708		935,770
Otis Worldwide Corp.		23,175		1,783,316
Westinghouse Air Brake Technologies Corp.		14,780		1,421,393
				8,131,740
Miscellaneous - Manufacturing — 1.4%
Eaton Corp. PLC		17,773		2,697,231
Transportation — 2.0%
Canadian National Railway Co.		13,887		1,862,802
Union Pacific Corp.		7,186		1,963,287
				3,826,089
				25,206,900

MML Fundamental Value Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
Technology — 12.3%
Computers — 1.9%
Cognizant Technology Solutions Corp. Class A		18,231	$	1,634,774
Leidos Holdings, Inc.		12,219		1,319,896
NetApp, Inc.		8,819		731,977
				3,686,647
Semiconductors — 7.3%
Applied Materials, Inc.		15,636		2,060,825
KLA Corp.		3,043		1,113,921
Lam Research Corp.		1,784		959,096
Microchip Technology, Inc.		13,121		985,912
Micron Technology, Inc.		38,495		2,998,376
NXP Semiconductor NV		5,553		1,027,749
Qorvo, Inc. (a)		12,135		1,505,953
QUALCOMM, Inc.		22,352		3,415,833
				14,067,665
Software — 3.1%
Activision Blizzard, Inc.		18,969		1,519,607
Fidelity National Information Services, Inc.		26,101		2,621,062
SS&C Technologies Holdings, Inc		22,819		1,711,881
				5,852,550
				23,606,862
Utilities — 2.0%
Electric — 2.0%
CenterPoint Energy, Inc.		60,897		1,865,884
Dominion Energy, Inc.		24,241		2,059,758
				3,925,642
TOTAL COMMON STOCK (Cost $143,645,976)				191,021,854
TOTAL EQUITIES (Cost $143,645,976)				191,021,854
TOTAL LONG-TERM INVESTMENTS (Cost $143,645,976)				191,021,854
	Principal Amount
SHORT-TERM INVESTMENTS — 0.6%
Repurchase Agreement — 0.6%
Fixed Income Clearing Corp., Repurchase Agreement, dated 3/31/22, 0.000%, due 4/01/22 (c)	$	1,226,980		1,226,980
TOTAL SHORT-TERM INVESTMENTS (Cost $1,226,980)				1,226,980

MML Fundamental Value Fund —  Portfolio of Investments (Continued)

	Principal Amount	Value
TOTAL INVESTMENTS — 100.0% (Cost $144,872,956) (d)		$	192,248,834
Other Assets/(Liabilities) — 0.0%			22,145
NET ASSETS — 100.0%		$	192,270,979

Abbreviation Legend

ADR American Depositary Receipt

Notes to Portfolio of Investments

Percentages are stated as a percent of net assets.

(a) Non-income producing security.

(b) Denotes all or a portion of security on loan. The total value of securities on loan as of March 31, 2022, was $3,435,920 or 1.79% of net assets. Total securities on loan may be less than the amounts identified in the Portfolio of Investments. The Fund received $3,539,844 of non-cash collateral (U.S. Treasury and/or Agency obligations) related to securities lending activity. This amount is not reflected on the Portfolio of Investments. (Note 2).

(c) Maturity value of $1,226,980. Collateralized by U.S. Government Agency obligations with a rate of 0.375%, maturity date of 9/15/24, and an aggregate market value, including accrued interest, of $1,251,551.

(d) See Note 3 for aggregate cost for federal tax purposes.

MML Global Fund —  Portfolio of Investments

March 31, 2022 (Unaudited)

	Number of Shares		Value
EQUITIES — 98.8%
COMMON STOCK — 98.8%
Austria — 0.3%
Erste Group Bank AG		15,374	$	557,589
Canada — 4.0%
Canadian National Railway Co.		22,384		3,002,590
Canadian Pacific Railway Ltd. (a)		44,284		3,655,201
				6,657,791
Denmark — 0.5%
Carlsberg A/S Class B		6,529		797,154
France — 10.8%
Air Liquide SA		8,814		1,541,914
Danone SA		42,610		2,347,294
EssilorLuxottica SA		3,587		654,789
Hermes International		410		581,537
Legrand SA		18,662		1,774,140
LVMH Moet Hennessy Louis Vuitton SE		5,175		3,684,960
Pernod Ricard SA		11,208		2,458,161
Schneider Electric SE		29,292		4,890,488
				17,933,283
Germany — 4.1%
Bayer AG Registered		30,250		2,069,620
Brenntag SE		9,559		772,931
Deutsche Boerse AG		4,673		839,452
Merck KGaA		11,353		2,377,646
MTU Aero Engines AG		3,238		752,338
				6,811,987
Ireland — 10.7%
Accenture PLC Class A		13,068		4,406,921
Aon PLC Class A		7,414		2,414,221
Linde PLC (b)		10,199		3,286,535
Linde PLC		2,690		859,267
Medtronic PLC		41,045		4,553,943
Willis Towers Watson PLC		9,509		2,246,216
				17,767,103
Israel — 1.0%
Check Point Software Technologies Ltd. (b)		11,529		1,594,000
Japan — 1.9%
Hoya Corp.		6,300		718,904
Kubota Corp.		95,800		1,796,056

MML Global Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
Olympus Corp.		30,400	$	577,551
				3,092,511
Mexico — 0.3%
Grupo Financiero Banorte SAB de CV Class O		57,770		435,148
Netherlands — 2.4%
Akzo Nobel NV		17,935		1,540,906
Heineken NV		23,263		2,217,869
Wolters Kluwer NV		2,653		282,411
				4,041,186
Republic of Korea — 0.8%
Samsung Electronics Co. Ltd.		23,639		1,350,440
Spain — 0.9%
Aena SME SA (b) (c)		8,708		1,442,535
Sweden — 1.2%
Essity AB Class B		86,762		2,044,240
Switzerland — 8.8%
Adecco Group AG Registered		10,344		468,178
Cie Financiere Richemont SA Registered		16,726		2,121,668
Julius Baer Group Ltd.		10,998		637,958
Nestle SA Registered		32,215		4,182,177
Roche Holding AG		11,475		4,538,079
Sonova Holding AG Registered		1,512		630,657
UBS Group AG Registered		96,876		1,892,667
				14,471,384
United Kingdom — 7.5%
Aptiv PLC (b)		5,799		694,198
Burberry Group PLC		40,708		888,946
Compass Group PLC		53,339		1,149,512
Diageo PLC		79,097		3,997,815
London Stock Exchange Group PLC		8,221		858,895
Reckitt Benckiser Group PLC		28,467		2,175,932
Rolls-Royce Holdings PLC (b)		608,961		800,660
Whitbread PLC (b)		17,979		669,053
WPP PLC		94,443		1,235,903
				12,470,914
United States — 43.6%
3M Co.		12,324		1,834,797
Abbott Laboratories		20,701		2,450,170
American Express Co.		12,347		2,308,889
Amphenol Corp. Class A		13,696		1,031,994
Boston Scientific Corp. (b)		57,823		2,560,981
Carrier Global Corp.		9,366		429,618
The Charles Schwab Corp.		33,647		2,836,779
Cisco Systems, Inc.		11,093		618,546
Cognizant Technology Solutions Corp. Class A		23,301		2,089,401
Colgate-Palmolive Co.		17,907		1,357,888
Comcast Corp. Class A		94,515		4,425,192
The Cooper Cos., Inc.		4,061		1,695,833

MML Global Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
eBay, Inc.		24,464	$	1,400,809
Equifax, Inc.		8,767		2,078,656
Fidelity National Information Services, Inc.		22,000		2,209,240
Fiserv, Inc. (b)		24,140		2,447,796
The Goldman Sachs Group, Inc.		8,262		2,727,286
Honeywell International, Inc.		16,833		3,275,365
International Flavors & Fragrances, Inc.		9,349		1,227,804
Liberty Broadband Corp. Class C (b)		16,932		2,291,238
Marriott International, Inc. Class A (b)		6,510		1,144,132
Microchip Technology, Inc.		11,194		841,117
Omnicom Group, Inc.		5,282		448,336
Oracle Corp.		32,516		2,690,049
Otis Worldwide Corp.		10,844		834,446
PayPal Holdings, Inc. (b)		7,579		876,511
PPG Industries, Inc.		12,261		1,607,049
Stryker Corp.		11,156		2,982,557
Thermo Fisher Scientific, Inc.		9,205		5,436,933
Union Pacific Corp.		6,506		1,777,504
United Parcel Service, Inc. Class B		8,705		1,866,874
Visa, Inc. Class A		22,935		5,086,295
The Walt Disney Co. (b)		19,642		2,694,097
Waters Corp. (b)		4,248		1,318,537
Zimmer Biomet Holdings, Inc.		9,313		1,191,132
				72,093,851
TOTAL COMMON STOCK (Cost $125,990,782)				163,561,116
TOTAL EQUITIES (Cost $125,990,782)				163,561,116
WARRANTS — 0.1%
Switzerland — 0.1%
Cie Financiere Richemont SA, Expires 11/22/23, Strike 67.00 (b)		105,484		83,333
TOTAL WARRANTS (Cost $0)				83,333
TOTAL LONG-TERM INVESTMENTS (Cost $125,990,782)				163,644,449
	Principal Amount
SHORT-TERM INVESTMENTS — 0.3%
Repurchase Agreement — 0.3%
Fixed Income Clearing Corp., Repurchase Agreement, dated 3/31/22, 0.000%, due 4/01/22 (d)	$	515,689		515,689
TOTAL SHORT-TERM INVESTMENTS (Cost $515,689)				515,689

MML Global Fund —  Portfolio of Investments (Continued)

	Principal Amount	Value
TOTAL INVESTMENTS — 99.2% (Cost $126,506,471) (e)		$	164,160,138
Other Assets/(Liabilities) — 0.8%			1,382,556
NET ASSETS — 100.0%		$	165,542,694

Notes to Portfolio of Investments

Percentages are stated as a percent of net assets.

(a) Denotes all or a portion of security on loan. The total value of securities on loan as of March 31, 2022, was $991,677 or 0.60% of net assets. Total securities on loan may be less than the amounts identified in the Portfolio of Investments. The Fund received $1,021,671 of non-cash collateral (U.S. Treasury and/or Agency obligations) related to securities lending activity. This amount is not reflected on the Portfolio of Investments. (Note 2).

(b) Non-income producing security.

(c) Security is exempt from registration under Regulation S or Rule 144A of the Securities Act of 1933. These securities are considered restricted and may be resold in transactions exempt from registration. At March 31, 2022, the aggregate market value of these securities amounted to $1,442,535 or 0.87% of net assets.

(d) Maturity value of $515,689. Collateralized by U.S. Government Agency obligations with a rate of 0.500%, maturity date of 4/15/24, and an aggregate market value, including accrued interest, of $526,099.

(e) See Note 3 for aggregate cost for federal tax purposes.

MML Growth & Income Fund —  Portfolio of Investments

March 31, 2022 (Unaudited)

	Number of Shares		Value
EQUITIES — 99.6%
COMMON STOCK — 99.6%
Basic Materials — 2.4%
Chemicals — 2.4%
DuPont de Nemours, Inc.		21,811	$	1,604,853
PPG Industries, Inc.		6,181		810,144
The Sherwin-Williams Co.		5,359		1,337,714
				3,752,711
				3,752,711
Communications — 10.4%
Internet — 8.5%
Alphabet, Inc. Class A (a)		3,169		8,814,098
Alphabet, Inc. Class C (a)		1,103		3,080,668
Amazon.com, Inc. (a)		478		1,558,256
				13,453,022
Media — 1.9%
Cable One, Inc.		469		686,728
Comcast Corp. Class A		51,141		2,394,422
				3,081,150
				16,534,172
Consumer, Cyclical — 8.7%
Apparel — 1.7%
LVMH Moet Hennessy Louis Vuitton SE		2,463		1,753,827
NIKE, Inc. Class B		6,891		927,253
				2,681,080
Retail — 7.0%
Costco Wholesale Corp.		3,964		2,282,669
Dollar General Corp.		8,474		1,886,567
The Home Depot, Inc.		7,553		2,260,840
Starbucks Corp.		5,444		495,241
Target Corp.		11,870		2,519,051
Tractor Supply Co.		7,295		1,702,434
				11,146,802
				13,827,882

MML Growth & Income Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
Consumer, Non-cyclical — 23.7%
Beverages — 2.5%
Diageo PLC		41,387	$	2,091,831
Pernod Ricard SA		8,284		1,816,864
				3,908,695
Biotechnology — 2.5%
Illumina, Inc. (a)		2,774		969,236
Maravai LifeSciences Holdings, Inc. Class A (a)		16,627		586,434
Vertex Pharmaceuticals, Inc. (a)		9,552		2,492,785
				4,048,455
Cosmetics & Personal Care — 1.5%
Colgate-Palmolive Co.		20,838		1,580,146
The Estee Lauder Cos., Inc. Class A		2,975		810,152
				2,390,298
Food — 1.0%
Danone SA		8,673		477,777
Mondelez International, Inc. Class A		18,129		1,138,139
				1,615,916
Health Care – Products — 5.3%
Danaher Corp.		8,002		2,347,226
Medtronic PLC		25,144		2,789,727
Thermo Fisher Scientific, Inc.		5,437		3,211,364
				8,348,317
Health Care – Services — 1.5%
ICON PLC (a)		9,790		2,381,124
Household Products & Wares — 0.7%
Kimberly-Clark Corp.		9,301		1,145,511
Pharmaceuticals — 8.7%
Becton Dickinson and Co.		10,356		2,754,696
Cigna Corp.		6,388		1,530,629
Eli Lilly & Co.		7,523		2,154,361
Johnson & Johnson		20,764		3,680,004
Merck & Co., Inc.		28,871		2,368,865
Zoetis, Inc.		7,276		1,372,181
				13,860,736
				37,699,052
Energy — 2.2%
Oil & Gas — 2.2%
ConocoPhillips		34,771		3,477,100
Financial — 19.2%
Banks — 7.3%
Bank of America Corp.		88,472		3,646,816
The Goldman Sachs Group, Inc.		7,715		2,546,721

MML Growth & Income Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
JP Morgan Chase & Co.		27,765	$	3,784,925
Truist Financial Corp.		28,906		1,638,970
				11,617,432
Diversified Financial Services — 6.3%
The Charles Schwab Corp.		16,885		1,423,574
Mastercard, Inc. Class A		8,094		2,892,634
Nasdaq, Inc.		13,397		2,387,345
Visa, Inc. Class A		14,963		3,318,345
				10,021,898
Insurance — 1.2%
Chubb Ltd.		8,941		1,912,480
Private Equity — 0.6%
Blackstone, Inc.		7,421		942,022
Real Estate Investment Trusts (REITS) — 3.8%
American Tower Corp.		11,647		2,925,959
Equinix, Inc.		2,587		1,918,571
Rayonier, Inc.		27,589		1,134,460
				5,978,990
				30,472,822
Industrial — 11.8%
Aerospace & Defense — 1.4%
Howmet Aerospace, Inc.		62,217		2,236,079
Building Materials — 2.4%
Johnson Controls International PLC		35,618		2,335,472
Masco Corp.		29,100		1,484,100
				3,819,572
Electrical Components & Equipment — 0.9%
AMETEK, Inc.		10,754		1,432,218
Electronics — 3.4%
Fortive Corp.		17,410		1,060,791
Honeywell International, Inc.		13,980		2,720,229
TE Connectivity Ltd.		12,685		1,661,481
				5,442,501
Packaging & Containers — 2.2%
Ball Corp.		14,416		1,297,440
Crown Holdings, Inc.		17,665		2,209,715
				3,507,155
Transportation — 1.5%
Canadian Pacific Railway Ltd.		22,952		1,894,458

MML Growth & Income Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
Old Dominion Freight Line, Inc.		1,541	$	460,266
				2,354,724
				18,792,249
Technology — 20.5%
Computers — 5.7%
Accenture PLC Class A		6,477		2,184,239
Amdocs Ltd.		26,763		2,200,186
Apple, Inc.		26,313		4,594,513
				8,978,938
Semiconductors — 2.4%
Analog Devices, Inc.		10,019		1,654,938
Texas Instruments, Inc.		12,196		2,237,722
				3,892,660
Software — 12.4%
Adobe, Inc. (a)		4,807		2,190,165
Electronic Arts, Inc.		17,780		2,249,348
Fidelity National Information Services, Inc.		21,446		2,153,607
Fiserv, Inc. (a)		16,493		1,672,390
Microsoft Corp.		31,556		9,729,031
salesforce.com, Inc. (a)		7,878		1,672,657
				19,667,198
				32,538,796
Utilities — 0.7%
Electric — 0.7%
American Electric Power Co., Inc.		10,228		1,020,448
TOTAL COMMON STOCK (Cost $86,645,371)				158,115,232
TOTAL EQUITIES (Cost $86,645,371)				158,115,232
TOTAL LONG-TERM INVESTMENTS (Cost $86,645,371)				158,115,232
	Principal Amount
SHORT-TERM INVESTMENTS — 0.5%
Repurchase Agreement — 0.5%
Fixed Income Clearing Corp., Repurchase Agreement, dated 3/31/22, 0.000%,due 4/01/22 (b)	$	835,080		835,080
TOTAL SHORT-TERM INVESTMENTS (Cost $835,080)				835,080

MML Growth & Income Fund —  Portfolio of Investments (Continued)

	Principal Amount	Value
TOTAL INVESTMENTS — 100.1% (Cost $87,480,451) (c)		$	158,950,312
Other Assets/(Liabilities) — (0.1)%			(159,833)
NET ASSETS — 100.0%		$	158,790,479

Notes to Portfolio of Investments

Percentages are stated as a percent of net assets.

(a) Non-income producing security.

(b) Maturity value of $835,080. Collateralized by U.S. Government Agency obligations with a rate of 0.500%, maturity date of 4/15/24, and an aggregate market value, including accrued interest, of $851,897.

(c) See Note 3 for aggregate cost for federal tax purposes.

MML Income & Growth Fund —  Portfolio of Investments

March 31, 2022 (Unaudited)

	Number of Shares		Value
EQUITIES — 98.1%
COMMON STOCK — 98.1%
Basic Materials — 7.1%
Chemicals — 7.1%
Air Products & Chemicals, Inc.		21,127	$	5,279,848
Axalta Coating Systems Ltd. (a)		57,691		1,418,045
DuPont de Nemours, Inc.		54,803		4,032,405
Element Solutions, Inc.		87,996		1,927,112
International Flavors & Fragrances, Inc.		38,105		5,004,330
				17,661,740
				17,661,740
Communications — 3.2%
Media — 2.4%
Altice USA, Inc. Class A (a)		63,416		791,432
Comcast Corp. Class A		110,108		5,155,256
				5,946,688
Telecommunications — 0.8%
T-Mobile US, Inc. (a)		14,853		1,906,383
				7,853,071
Consumer, Cyclical — 8.1%
Apparel — 0.5%
Ralph Lauren Corp.		10,731		1,217,325
Entertainment — 1.7%
Marriott Vacations Worldwide Corp.		11,183		1,763,559
SeaWorld Entertainment, Inc. (a)		33,175		2,469,547
				4,233,106
Food Services — 0.8%
Aramark		49,197		1,849,807
Lodging — 1.6%
Las Vegas Sands Corp. (a)		47,965		1,864,400
MGM Resorts International		51,847		2,174,463
				4,038,863
Retail — 3.5%
Advance Auto Parts, Inc.		14,872		3,077,909
Dollar General Corp.		8,412		1,872,764
Lithia Motors, Inc.		4,803		1,441,476

MML Income & Growth Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
Lowe's Cos., Inc.		11,485	$	2,322,152
				8,714,301
				20,053,402
Consumer, Non-cyclical — 26.8%
Agriculture — 2.1%
Altria Group, Inc.		44,030		2,300,567
Philip Morris International, Inc.		29,982		2,816,509
				5,117,076
Beverages — 2.9%
Coca-Cola Europacific Partners PLC		107,474		5,224,311
Molson Coors Beverage Co. Class B		38,157		2,036,821
				7,261,132
Biotechnology — 2.2%
Corteva, Inc.		94,362		5,423,928
Commercial Services — 0.2%
PROG Holdings, Inc. (a)		18,875		543,034
Cosmetics & Personal Care — 0.9%
The Procter & Gamble Co.		14,977		2,288,486
Health Care – Products — 3.7%
Envista Holdings Corp. (a)		27,132		1,321,600
Hologic, Inc. (a)		17,068		1,311,164
LivaNova PLC (a)		14,063		1,150,775
Medtronic PLC		50,153		5,564,475
				9,348,014
Health Care – Services — 4.8%
Anthem, Inc.		5,022		2,466,907
Humana, Inc.		6,070		2,641,482
UnitedHealth Group, Inc.		13,199		6,731,094
				11,839,483
Pharmaceuticals — 10.0%
Becton Dickinson and Co.		9,809		2,609,194
Cigna Corp.		19,598		4,695,877
CVS Health Corp.		53,788		5,443,883
Johnson & Johnson		10,921		1,935,529
Merck & Co., Inc.		88,042		7,223,846
Perrigo Co. PLC		80,972		3,111,754
				25,020,083
				66,841,236
Energy — 8.8%
Oil & Gas — 7.3%
Hess Corp.		65,325		6,992,388
Phillips 66		64,648		5,584,941
Pioneer Natural Resources Co.		13,970		3,492,919

MML Income & Growth Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
Valero Energy Corp.		21,488	$	2,181,891
				18,252,139
Pipelines — 1.5%
Enbridge, Inc.		80,685		3,718,772
				21,970,911
Financial — 20.9%
Banks — 9.1%
JP Morgan Chase & Co.		38,390		5,233,325
M&T Bank Corp.		12,282		2,081,799
Northern Trust Corp.		38,593		4,494,155
US Bancorp		88,896		4,724,822
Wells Fargo & Co.		124,885		6,051,927
				22,586,028
Diversified Financial Services — 2.5%
AerCap Holdings NV (a)		22,633		1,137,987
American Express Co.		27,407		5,125,109
				6,263,096
Insurance — 6.4%
The Allstate Corp.		36,821		5,100,077
American International Group, Inc.		31,718		1,990,939
Axis Capital Holdings Ltd.		21,910		1,324,898
Chubb Ltd.		26,368		5,640,115
Willis Towers Watson PLC		8,336		1,969,130
				16,025,159
Real Estate — 0.5%
The Howard Hughes Corp. (a)		12,518		1,296,990
Real Estate Investment Trusts (REITS) — 2.4%
American Campus Communities, Inc.		23,249		1,301,247
Corporate Office Properties Trust		81,347		2,321,643
MGM Growth Properties LLC Class A		30,026		1,162,006
VICI Properties, Inc.		40,408		1,150,012
				5,934,908
				52,106,181
Industrial — 12.5%
Aerospace & Defense — 3.7%
General Dynamics Corp.		12,271		2,959,520
Raytheon Technologies Corp.		64,005		6,340,975
				9,300,495
Building Materials — 0.2%
MDU Resources Group, Inc.		20,962		558,637
Electrical Components & Equipment — 0.5%
Emerson Electric Co.		12,245		1,200,622

MML Income & Growth Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
Engineering & Construction — 1.5%
AECOM		30,869	$	2,371,048
Jacobs Engineering Group, Inc.		9,419		1,298,032
				3,669,080
Hand & Machine Tools — 1.6%
Stanley Black & Decker, Inc.		28,717		4,014,349
Machinery – Construction & Mining — 1.6%
BWX Technologies, Inc.		31,517		1,697,506
Vertiv Holdings Co.		154,763		2,166,682
				3,864,188
Machinery – Diversified — 2.5%
Deere & Co.		15,258		6,339,089
Transportation — 0.9%
J.B. Hunt Transport Services, Inc.		11,274		2,263,707
				31,210,167
Technology — 6.5%
Computers — 1.8%
Cognizant Technology Solutions Corp. Class A		48,149		4,317,521
Semiconductors — 2.4%
Broadcom, Inc.		4,452		2,803,335
Microchip Technology, Inc.		23,226		1,745,201
QUALCOMM, Inc.		9,463		1,446,136
				5,994,672
Software — 2.3%
Fidelity National Information Services, Inc.		12,793		1,284,673
Oracle Corp.		54,382		4,499,023
				5,783,696
				16,095,889
Utilities — 4.2%
Electric — 4.2%
CenterPoint Energy, Inc.		46,412		1,422,063
Constellation Energy Corp.		11,756		661,275
Entergy Corp.		24,081		2,811,457
Exelon Corp.		35,270		1,679,910

MML Income & Growth Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
Pinnacle West Capital Corp.		51,058	$	3,987,630
				10,562,335
				10,562,335
TOTAL COMMON STOCK (Cost $184,476,007)				244,354,932
TOTAL EQUITIES (Cost $184,476,007)				244,354,932
TOTAL LONG-TERM INVESTMENTS (Cost $184,476,007)				244,354,932
	Principal Amount
SHORT-TERM INVESTMENTS — 1.6%
Repurchase Agreement — 1.6%
Fixed Income Clearing Corp., Repurchase Agreement, dated 3/31/22, 0.000%, due 4/01/22 (b)	$	3,816,071		3,816,071
TOTAL SHORT-TERM INVESTMENTS (Cost $3,816,071)				3,816,071
TOTAL INVESTMENTS — 99.7% (Cost $188,292,078) (c)				248,171,003
Other Assets/(Liabilities) — 0.3%				867,121
NET ASSETS — 100.0%			$	249,038,124

Notes to Portfolio of Investments

Percentages are stated as a percent of net assets.

(a) Non-income producing security.

(b) Maturity value of $3,816,071. Collateralized by U.S. Government Agency obligations with rates ranging from 0.375% - 0.500%, maturity dates ranging from 4/15/24 - 9/15/24, and an aggregate market value, including accrued interest, of $3,892,442.

(c) See Note 3 for aggregate cost for federal tax purposes.

MML International Equity Fund —  Portfolio of Investments

March 31, 2022 (Unaudited)

	Number of Shares		Value
EQUITIES — 98.1%
COMMON STOCK — 97.7%
Australia — 0.7%
Brambles Ltd.		97,100	$	716,023
Orica Ltd.		83,286		986,320
				1,702,343
Belgium — 1.1%
Anheuser-Busch InBev SA		25,300		1,517,477
KBC Group NV		15,959		1,147,764
				2,665,241
Canada — 4.0%
Canadian National Railway Co.		25,697		3,446,996
Intact Financial Corp.		7,959		1,176,008
Open Text Corp.		23,300		987,988
Restaurant Brands International, Inc.		12,500		729,875
Suncor Energy, Inc.		39,701		1,292,509
The Toronto-Dominion Bank		23,261		1,845,585
				9,478,961
Cayman Islands — 2.3%
Alibaba Group Holding Ltd. (a)		140,840		1,927,717
JD.com, Inc. Class A (a)		50		1,464
JD.COM, Inc., Class A, Lockup Shares (Acquired 1/20/22, Cost $47,415)) (a) (b) (c)		1,300		36,170
NetEase, Inc.		43,800		789,611
Tencent Holdings Ltd.		34,600		1,632,269
Trip.com Group Ltd. (a)		11,500		270,891
Vipshop Holdings Ltd. ADR (a)		88,064		792,576
				5,450,698
Denmark — 2.1%
Carlsberg A/S Class B		9,851		1,202,751
Novo Nordisk A/S Class B		34,037		3,769,485
				4,972,236
Finland — 0.2%
UPM-Kymmene OYJ (d)		15,000		489,459
France — 17.0%
Accor SA (a)		48,200		1,544,644
Air Liquide SA		26,432		4,623,993
BNP Paribas SA		42,676		2,429,854
Capgemini SE		20,094		4,469,774
Cie de Saint-Gobain		15,428		918,121

MML International Equity Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
Cie Generale des Etablissements Michelin SCA		8,905	$	1,203,002
Danone SA		19,000		1,046,670
Dassault Systemes SE		22,606		1,113,910
Edenred		4,500		222,460
Engie SA		131,084		1,718,302
EssilorLuxottica SA		12,802		2,336,942
L'Oreal SA		3,799		1,520,610
Legrand SA		16,272		1,546,930
LVMH Moet Hennessy Louis Vuitton SE		5,332		3,796,755
Pernod Ricard SA		13,495		2,959,751
Publicis Groupe SA		20,651		1,254,475
Schneider Electric SE		31,966		5,336,930
Valeo		46,110		847,171
Worldline SA (a) (e)		36,400		1,578,056
				40,468,350
Germany — 15.6%
adidas AG		3,255		760,095
Allianz SE Registered		10,610		2,533,609
Bayer AG Registered		86,142		5,893,595
Bayerische Motoren Werke AG		24,475		2,119,885
Beiersdorf AG		26,716		2,807,352
Continental AG (a)		25,220		1,819,147
Daimler Truck Holding AG (a)		39,449		1,095,934
Deutsche Boerse AG		17,552		3,153,019
Fresenius Medical Care AG & Co. KGaA		23,300		1,562,557
Fresenius SE & Co. KGaA		38,600		1,420,332
Henkel AG & Co. KGaA		16,000		1,060,534
Mercedes-Benz Group AG		31,298		2,198,266
Merck KGaA		14,731		3,085,096
MTU Aero Engines AG		4,307		1,000,716
SAP SE		40,214		4,487,211
Siemens AG Registered		5,290		732,994
thyssenkrupp AG (a)		141,374		1,217,860
Vitesco Technologies Group AG (a)		1,411		55,999
				37,004,201
Hong Kong — 1.5%
AIA Group Ltd.		344,800		3,609,861
India — 0.6%
Axis Bank Ltd. (a)		79,042		786,437
HDFC Bank Ltd. ADR		11,494		704,927
				1,491,364
Indonesia — 0.0%
Bank Mandiri Persero Tbk PT		177,700		97,505
Ireland — 1.9%
Linde PLC (a)		6,349		2,045,908
Ryanair Holdings PLC Sponsored ADR (a)		28,262		2,462,185
				4,508,093
Israel — 0.9%
Check Point Software Technologies Ltd. (a)		14,957		2,067,955

MML International Equity Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
Italy — 2.3%
Eni SpA		74,144	$	1,087,032
Intesa Sanpaolo SpA		1,868,847		4,271,638
Intesa Sanpaolo SpA		12,000		27,608
				5,386,278
Japan — 10.3%
Daikin Industries Ltd.		14,800		2,694,709
Denso Corp.		19,200		1,225,179
Hitachi Ltd.		62,700		3,141,801
Hoya Corp.		17,800		2,031,188
Koito Manufacturing Co. Ltd.		23,400		947,331
Komatsu Ltd.		25,100		602,292
Kose Corp.		10,600		1,108,639
Kubota Corp.		112,500		2,109,148
Kyocera Corp.		28,600		1,607,569
Olympus Corp.		103,700		1,970,133
Shin-Etsu Chemical Co. Ltd.		6,300		960,027
SMC Corp.		2,800		1,563,837
Sony Group Corp.		20,600		2,126,146
Terumo Corp.		53,200		1,611,202
Toyota Motor Corp.		38,600		691,461
				24,390,662
Mexico — 0.4%
Grupo Televisa SAB Sponsored ADR		81,800		957,060
Netherlands — 4.8%
CNH Industrial NV		105,653		1,672,918
EXOR NV		20,920		1,595,611
ING Groep NV		132,106		1,380,610
Iveco Group NV (a)		12,156		78,988
Koninklijke Philips NV		60,777		1,859,350
Prosus NV		37,573		1,989,015
QIAGEN NV (a)		29,825		1,463,984
Randstad NV (d)		23,501		1,410,584
				11,451,060
Portugal — 0.5%
Galp Energia SGPS SA		90,702		1,147,825
Republic of Korea — 0.5%
NAVER Corp.		4,555		1,273,684
Singapore — 0.9%
DBS Group Holdings, Ltd.		81,200		2,134,776
Spain — 1.0%
Amadeus IT Group SA (a)		37,201		2,423,124
Sweden — 1.4%
Hennes & Mauritz AB Class B (d)		94,818		1,266,267
SKF AB Class B (d)		69,100		1,123,214
Volvo AB Class B		52,170		973,049
				3,362,530

MML International Equity Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
Switzerland — 13.6%
Cie Financiere Richemont SA Registered		23,593	$	2,992,737
Credit Suisse Group AG Registered		273,919		2,161,252
Holcim Ltd.		33,659		1,643,286
Julius Baer Group Ltd.		20,350		1,180,437
Nestle SA Registered		49,573		6,435,607
Novartis AG Registered		43,602		3,823,959
Roche Holding AG		14,923		5,901,678
Sika AG Registered		4,773		1,572,497
The Swatch Group AG		2,415		686,243
UBS Group AG Registered		177,446		3,466,763
Zurich Insurance Group AG		5,068		2,498,669
				32,363,128
Taiwan — 1.1%
Taiwan Semiconductor Manufacturing Co. Ltd. Sponsored ADR		24,190		2,522,049
United Kingdom — 12.6%
Compass Group PLC		149,148		3,214,298
Diageo PLC		57,204		2,891,273
Experian PLC		56,800		2,190,492
Glencore PLC		326,144		2,119,534
Informa PLC (a)		96,300		755,786
Liberty Global PLC Class A (a)		51,600		1,316,316
Lloyds Banking Group PLC		4,154,984		2,543,671
London Stock Exchange Group PLC		9,319		973,609
Natwest Group PLC		214,600		603,845
Prudential PLC		71,700		1,059,136
Reckitt Benckiser Group PLC		30,069		2,298,384
RELX PLC		93,550		2,913,910
Rio Tinto PLC		22,399		1,776,821
Rolls-Royce Holdings PLC (a)		857,319		1,127,201
Schroders PLC		30,000		1,264,230
Smiths Group PLC		34,294		650,071
Tesco PLC		452,748		1,636,838
WPP PLC		52,592		688,231
				30,023,646
United States — 0.4%
Yum China Holdings, Inc.		23,869		991,518
TOTAL COMMON STOCK (Cost $241,251,874)				232,433,607
PREFERRED STOCK — 0.4%
Republic of Korea — 0.4%
Samsung Electronics Co. Ltd. 1.760%
		20,300		1,049,994
TOTAL PREFERRED STOCK (Cost $1,174,727)				1,049,994
TOTAL EQUITIES (Cost $242,426,601)				233,483,601

MML International Equity Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
MUTUAL FUNDS — 0.3%
United States — 0.3%
State Street Navigator Securities Lending Government Money Market Portfolio (f)		626,054	$	626,054
TOTAL MUTUAL FUNDS (Cost $626,054)				626,054
TOTAL LONG-TERM INVESTMENTS (Cost $243,052,655)				234,109,655
	Principal Amount
SHORT-TERM INVESTMENTS — 1.4%
Repurchase Agreement — 1.4%
Fixed Income Clearing Corp., Repurchase Agreement, dated 3/31/22, 0.000%, due 4/01/22 (g)	$	3,242,074		3,242,074
TOTAL SHORT-TERM INVESTMENTS (Cost $3,242,074)				3,242,074
TOTAL INVESTMENTS — 99.8% (Cost $246,294,729) (h)				237,351,729
Other Assets/(Liabilities) — 0.2%				468,420
NET ASSETS — 100.0%			$	237,820,149

Abbreviation Legend

ADR American Depositary Receipt

Notes to Portfolio of Investments

Percentages are stated as a percent of net assets.

(a) Non-income producing security.

(b) This security is fair valued in good faith in accordance with procedures approved by the Board of Trustees. At March 31, 2022, these securities amounted to a value of $36,170 or 0.02% of net assets.

(c) Restricted security. Certain securities are restricted as to resale. At March 31, 2022, these securities amounted to a value of $36,170 or 0.02% of net assets. The Fund generally bears the costs, if any, associated with the disposition of restricted securities.

(d) Denotes all or a portion of security on loan. The total value of securities on loan as of March 31, 2022, was $3,470,997 or 1.46% of net assets. Total securities on loan may be less than the amounts identified in the Portfolio of Investments. The Fund received $3,020,408 of non-cash collateral (U.S. Treasury and/or Agency obligations) related to securities lending activity. This amount is not reflected on the Portfolio of Investments. (Note 2).

(e) Security is exempt from registration under Regulation S or Rule 144A of the Securities Act of 1933. These securities are considered restricted and may be resold in transactions exempt from registration. At March 31, 2022, the aggregate market value of these securities amounted to $1,578,056 or 0.66% of net assets.

(f) Represents investment of security lending cash collateral. (Note 2).

(g) Maturity value of $3,242,074. Collateralized by U.S. Government Agency obligations with a rate of 0.500%, maturity date of 4/15/24, and an aggregate market value, including accrued interest, of $3,307,027.

(h) See Note 3 for aggregate cost for federal tax purposes.

Forward contracts
Counterparty	Settlement Date	Currency Purchased		Currency Sold		Unrealized Appreciation (Depreciation)
State Street Bank and Trust	6/15/22	CHF	166,000	USD	$180,415	$	(234)
State Street Bank and Trust Co	6/15/22	USD	1,194,465	CHF	1,097,000		3,748
						$	3,514

Currency Legend

CHF Swiss Franc

MML Large Cap Growth Fund —  Portfolio of Investments

March 31, 2022 (Unaudited)

	Number of Shares		Value
EQUITIES — 99.5%
COMMON STOCK — 99.5%
Communications — 27.2%
Internet — 22.1%
Alibaba Group Holding Ltd. Sponsored ADR (a)		28,022	$	3,048,794
Alphabet, Inc. Class A (a)		2,620		7,287,137
Alphabet, Inc. Class C (a)		2,563		7,158,433
Amazon.com, Inc. (a)		3,723		12,136,794
Meta Platforms, Inc. Class A (a)		39,728		8,833,918
Netflix, Inc. (a)		11,742		4,398,436
Shopify, Inc. Class A (a)		4,646		3,140,510
				46,004,022
Media — 5.1%
FactSet Research Systems, Inc.		8,594		3,731,085
The Walt Disney Co. (a)		51,206		7,023,415
				10,754,500
				56,758,522
Consumer, Cyclical — 7.1%
Auto Manufacturers — 3.2%
Tesla, Inc. (a)		6,153		6,630,472
Retail — 3.9%
Starbucks Corp.		46,026		4,186,985
Yum China Holdings, Inc.		30,333		1,260,033
Yum! Brands, Inc.		22,562		2,674,274
				8,121,292
				14,751,764
Consumer, Non-cyclical — 23.0%
Beverages — 2.8%
Monster Beverage Corp. (a)		73,355		5,861,065
Biotechnology — 9.0%
Illumina, Inc. (a)		14,008		4,894,395
Regeneron Pharmaceuticals, Inc. (a)		10,130		7,074,995
Vertex Pharmaceuticals, Inc. (a)		26,197		6,836,631
				18,806,021
Commercial Services — 3.1%
Block, Inc. (a)		24,167		3,277,045

MML Large Cap Growth Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
PayPal Holdings, Inc. (a)		27,635	$	3,195,988
				6,473,033
Health Care – Products — 1.3%
Intuitive Surgical, Inc. (a)		9,058		2,732,617
Pharmaceuticals — 6.8%
Novartis AG Sponsored ADR		67,136		5,891,184
Novo Nordisk A/S Sponsored ADR		24,506		2,721,391
Roche Holding AG Sponsored ADR		110,304		5,450,121
				14,062,696
				47,935,432
Financial — 7.0%
Diversified Financial Services — 7.0%
SEI Investments Co.		48,923		2,945,654
Visa, Inc. Class A		52,257		11,589,035
				14,534,689
				14,534,689
Industrial — 8.6%
Aerospace & Defense — 4.5%
The Boeing Co. (a)		49,203		9,422,374
Machinery – Diversified — 2.6%
Deere & Co.		13,275		5,515,232
Transportation — 1.5%
Expeditors International of Washington, Inc.		29,405		3,033,420
				17,971,026
Technology — 26.6%
Semiconductors — 9.0%
NVIDIA Corp.		51,295		13,996,354
QUALCOMM, Inc.		31,560		4,822,999
				18,819,353
Software — 17.6%
Autodesk, Inc. (a)		32,880		7,047,828
Microsoft Corp.		33,339		10,278,747
Oracle Corp.		111,566		9,229,855
salesforce.com, Inc. (a)		34,549		7,335,444

MML Large Cap Growth Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
Workday, Inc. Class A (a)		11,459	$	2,743,972
				36,635,846
				55,455,199
TOTAL COMMON STOCK (Cost $161,802,538)				207,406,632
TOTAL EQUITIES (Cost $161,802,538)				207,406,632
TOTAL LONG-TERM INVESTMENTS (Cost $161,802,538)				207,406,632
	Principal Amount
SHORT-TERM INVESTMENTS — 0.5%
Repurchase Agreement — 0.5%
Fixed Income Clearing Corp., Repurchase Agreement, dated 3/31/22, 0.000%, due 4/01/22 (b)	$	987,780		987,780
TOTAL SHORT-TERM INVESTMENTS (Cost $987,780)				987,780
TOTAL INVESTMENTS — 100.0% (Cost $162,790,318) (c)				208,394,412
Other Assets/(Liabilities) — (0.0)%				(95,387)
NET ASSETS — 100.0%			$	208,299,025

Abbreviation Legend

ADR American Depositary Receipt

Notes to Portfolio of Investments

Percentages are stated as a percent of net assets.

(a) Non-income producing security.

(b) Maturity value of $987,780. Collateralized by U.S. Government Agency obligations with a rate of 0.500%, maturity date of 4/15/24, and an aggregate market value, including accrued interest, of $1,007,622.

(c) See Note 3 for aggregate cost for federal tax purposes.

MML Managed Volatility Fund —  Portfolio of Investments

March 31, 2022 (Unaudited)

	Number of Shares		Value
EQUITIES — 101.5%
COMMON STOCK — 101.5%
Basic Materials — 2.2%
Chemicals — 1.6%
Air Products & Chemicals, Inc. (a)		765	$	191,181
Albemarle Corp. (a)		413		91,335
Celanese Corp. (a)		395		56,434
CF Industries Holdings, Inc. (a)		717		73,894
Dow, Inc. (a)		2,490		158,663
DuPont de Nemours, Inc. (a)		1,754		129,059
Eastman Chemical Co. (a)		421		47,177
Ecolab, Inc. (a)		845		149,193
FMC Corp. (a)		432		56,838
International Flavors & Fragrances, Inc. (a)		858		112,681
Linde PLC (a)		1,746		557,725
LyondellBasell Industries NV Class A (a)		870		89,453
The Mosaic Co. (a)		1,258		83,657
PPG Industries, Inc. (a)		830		108,788
The Sherwin-Williams Co. (a)		817		203,940
				2,110,018
Forest Products & Paper — 0.1%
International Paper Co. (a)		1,364		62,949
Iron & Steel — 0.1%
Nucor Corp. (a)		927		137,799
Mining — 0.4%
Freeport-McMoRan, Inc. (a)		4,999		248,650
Newmont Corp. (a)		2,718		215,945
				464,595
				2,775,361
Communications — 14.6%
Advertising — 0.1%
The Interpublic Group of Cos., Inc. (a)		1,413		50,091
Omnicom Group, Inc. (a)		699		59,331
				109,422
Internet — 10.7%
Alphabet, Inc. Class A (a) (b)		1,028		2,859,228
Alphabet, Inc. Class C (a) (b)		944		2,636,583
Amazon.com, Inc. (a) (b)		1,492		4,863,845
Booking Holdings, Inc. (a) (b)		139		326,435
CDW Corp. (a)		445		79,606
eBay, Inc. (a)		2,133		122,136

MML Managed Volatility Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
Etsy, Inc. (a) (b)		430	$	53,440
Expedia Group, Inc. (a) (b)		508		99,400
F5, Inc. (a) (b)		212		44,297
Match Group, Inc. (a) (b)		981		106,674
Meta Platforms, Inc. Class A (a) (b)		7,866		1,749,084
Netflix, Inc. (a) (b)		1,510		565,631
NortonLifeLock, Inc. (a)		1,956		51,873
Twitter, Inc. (a) (b)		2,707		104,734
VeriSign, Inc. (a) (b)		346		76,971
				13,739,937
Media — 1.6%
Charter Communications, Inc. Class A (a) (b)		403		219,845
Comcast Corp. Class A (a)		15,382		720,185
Discovery, Inc. Class A (a) (b)		885		22,054
Discovery, Inc. Class C (a) (b)		670		16,730
DISH Network Corp. Class A (a) (b)		884		27,979
FactSet Research Systems, Inc. (a)		130		56,439
Fox Corp. Class A (a)		778		30,692
Fox Corp. Class B (a)		818		29,677
News Corp. Class A (a)		837		18,540
News Corp. Class B (a)		1,016		22,880
Paramount Global Class B (a)		2,050		77,510
The Walt Disney Co. (a) (b)		6,194		849,569
				2,092,100
Telecommunications — 2.2%
Arista Networks, Inc. (a) (b)		760		105,625
AT&T, Inc. (a)		24,236		572,697
Cisco Systems, Inc. (a)		14,361		800,769
Corning, Inc. (a)		2,601		96,003
Juniper Networks, Inc. (a)		1,049		38,981
Lumen Technologies, Inc. (a)		3,054		34,419
Motorola Solutions, Inc. (a)		581		140,718
T-Mobile US, Inc. (a) (b)		2,030		260,550
Verizon Communications, Inc. (a)		14,404		733,740
				2,783,502
				18,724,961
Consumer, Cyclical — 9.9%
Airlines — 0.3%
Alaska Air Group, Inc. (a) (b)		399		23,146
American Airlines Group, Inc. (a) (b)		2,403		43,855
Delta Air Lines, Inc. (a) (b)		2,283		90,338
Southwest Airlines Co. (a) (b)		1,951		89,356
United Airlines Holdings, Inc. (a) (b)		1,015		47,055
				293,750
Apparel — 0.6%
NIKE, Inc. Class B (a)		4,340		583,990
PVH Corp. (a)		214		16,395
Ralph Lauren Corp. (a)		181		20,533
Tapestry, Inc. (a)		845		31,392
Under Armour, Inc. Class A (a) (b)		604		10,280

MML Managed Volatility Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
Under Armour, Inc. Class C (a) (b)		911	$	14,175
VF Corp. (a)		1,165		66,242
				743,007
Auto Manufacturers — 2.9%
Cummins, Inc. (a)		492		100,914
Ford Motor Co. (a)		13,567		229,418
General Motors Co. (a) (b)		4,909		214,720
PACCAR, Inc. (a)		1,228		108,150
Tesla, Inc. (a) (b)		2,853		3,074,393
				3,727,595
Auto Parts & Equipment — 0.1%
Aptiv PLC (a) (b)		898		107,500
BorgWarner, Inc. (a)		879		34,193
				141,693
Distribution & Wholesale — 0.3%
Copart, Inc. (a) (b)		724		90,840
Fastenal Co. (a)		2,043		121,354
LKQ Corp. (a)		980		44,502
Pool Corp. (a)		138		58,353
W.W. Grainger, Inc. (a)		145		74,790
				389,839
Entertainment — 0.1%
Caesars Entertainment, Inc. (a) (b)		750		58,020
Live Nation Entertainment, Inc. (a) (b)		465		54,703
Penn National Gaming, Inc. (a) (b)		582		24,688
				137,411
Home Builders — 0.2%
D.R. Horton, Inc. (a)		1,071		79,800
Lennar Corp. Class A (a)		926		75,163
NVR, Inc. (a) (b)		11		49,140
PulteGroup, Inc. (a)		882		36,956
				241,059
Home Furnishing — 0.0%
Whirlpool Corp. (a)		190		32,828
Housewares — 0.0%
Newell Brands, Inc. (a)		1,346		28,818
Leisure Time — 0.1%
Carnival Corp. (a) (b)		2,851		57,647
Norwegian Cruise Line Holdings Ltd. (a) (b)		1,247		27,284
Royal Caribbean Cruises Ltd. (a) (b)		788		66,019
				150,950
Lodging — 0.4%
Hilton Worldwide Holdings, Inc. (a) (b)		949		144,001
Las Vegas Sands Corp. (a) (b)		1,231		47,849
Marriott International, Inc. Class A (a) (b)		932		163,799

MML Managed Volatility Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
MGM Resorts International (a)		1,335	$	55,990
Wynn Resorts Ltd. (a) (b)		383		30,541
				442,180
Retail — 4.9%
Advance Auto Parts, Inc. (a)		220		45,531
AutoZone, Inc. (a) (b)		70		143,121
Bath & Body Works, Inc. (a)		924		44,167
Best Buy Co., Inc. (a)		729		66,266
CarMax, Inc. (a) (b)		543		52,389
Chipotle Mexican Grill, Inc. (a) (b)		96		151,875
Costco Wholesale Corp. (a)		1,510		869,533
Darden Restaurants, Inc. (a)		447		59,429
Dollar General Corp. (a)		785		174,765
Dollar Tree, Inc. (a) (b)		763		122,194
Domino's Pizza, Inc. (a)		130		52,911
Genuine Parts Co. (a)		460		57,969
The Home Depot, Inc. (a)		3,559		1,065,315
Lowe's Cos., Inc. (a)		2,294		463,824
McDonald's Corp. (a)		2,536		627,102
O'Reilly Automotive, Inc. (a) (b)		228		156,171
Ross Stores, Inc. (a)		1,220		110,361
Starbucks Corp. (a)		3,893		354,146
Target Corp. (a)		1,632		346,343
The TJX Cos., Inc. (a)		4,048		245,228
Tractor Supply Co. (a)		381		88,914
Ulta Beauty, Inc. (a) (b)		182		72,476
Walgreens Boots Alliance, Inc. (a)		2,519		112,776
Walmart, Inc. (a)		4,815		717,050
Yum! Brands, Inc. (a)		986		116,871
				6,316,727
Toys, Games & Hobbies — 0.0%
Hasbro, Inc. (a)		432		35,389
				12,681,246
Consumer, Non-cyclical — 20.4%
Agriculture — 0.8%
Altria Group, Inc. (a)		6,164		322,069
Archer-Daniels-Midland Co. (a)		1,891		170,681
Philip Morris International, Inc. (a)		5,271		495,158
				987,908
Beverages — 1.5%
Brown-Forman Corp. Class B (a)		604		40,480
The Coca-Cola Co. (a)		13,230		820,260
Constellation Brands, Inc. Class A (a)		576		132,664
Molson Coors Beverage Co. Class B (a)		613		32,722
Monster Beverage Corp. (a) (b)		1,265		101,073
PepsiCo, Inc. (a)		4,704		787,356
				1,914,555
Biotechnology — 1.5%
Amgen, Inc. (a)		1,915		463,085

MML Managed Volatility Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
Bio-Rad Laboratories, Inc. Class A (a) (b)		72	$	40,552
Biogen, Inc. (a) (b)		516		108,670
Corteva, Inc. (a)		2,466		141,746
Gilead Sciences, Inc. (a)		4,248		252,544
Illumina, Inc. (a) (b)		530		185,182
Incyte Corp. (a) (b)		626		49,717
Moderna, Inc. (a) (b)		1,208		208,090
Regeneron Pharmaceuticals, Inc. (a) (b)		365		254,923
Vertex Pharmaceuticals, Inc. (a) (b)		864		225,478
				1,929,987
Commercial Services — 1.9%
Automatic Data Processing, Inc. (a)		1,426		324,472
Cintas Corp. (a)		312		132,722
Equifax, Inc. (a)		407		96,500
FleetCor Technologies, Inc. (a) (b)		267		66,499
Gartner, Inc. (a) (b)		277		82,396
Global Payments, Inc. (a)		962		131,640
MarketAxess Holdings, Inc. (a)		139		47,288
Moody's Corp. (a)		562		189,624
Nielsen Holdings PLC (a)		1,326		36,120
PayPal Holdings, Inc. (a) (b)		3,959		457,858
Quanta Services, Inc. (a)		473		62,252
Robert Half International, Inc. (a)		357		40,762
Rollins, Inc. (a)		750		26,287
S&P Global, Inc. (a)		1,204		493,857
United Rentals, Inc. (a) (b)		243		86,316
Verisk Analytics, Inc. (a)		566		121,481
				2,396,074
Cosmetics & Personal Care — 1.3%
Colgate-Palmolive Co. (a)		2,943		223,168
The Estee Lauder Cos., Inc. Class A (a)		807		219,762
The Procter & Gamble Co. (a)		8,154		1,245,931
				1,688,861
Food — 1.1%
Campbell Soup Co. (a)		671		29,907
Conagra Brands, Inc. (a)		1,772		59,486
General Mills, Inc. (a)		2,066		139,910
The Hershey Co. (a)		487		105,499
Hormel Foods Corp. (a)		937		48,293
The J.M. Smucker Co. (a)		405		54,841
Kellogg Co. (a)		959		61,846
The Kraft Heinz Co. (a)		2,390		94,142
The Kroger Co. (a)		2,262		129,771
Lamb Weston Holdings, Inc. (a)		473		28,337
McCormick & Co., Inc. (a)		824		82,235
Mondelez International, Inc. Class A (a)		4,703		295,254
Sysco Corp. (a)		1,749		142,806
Tyson Foods, Inc. Class A (a)		984		88,196
				1,360,523
Health Care – Products — 4.0%
Abbott Laboratories (a)		6,020		712,527
ABIOMED, Inc. (a) (b)		157		52,005

MML Managed Volatility Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
Align Technology, Inc. (a) (b)		249	$	108,564
Baxter International, Inc. (a)		1,722		133,524
Bio-Techne Corp. (a)		130		56,295
Boston Scientific Corp. (a) (b)		4,937		218,660
The Cooper Cos., Inc. (a)		177		73,913
Danaher Corp. (a)		2,168		635,940
Dentsply Sirona, Inc. (a)		744		36,620
Edwards Lifesciences Corp. (a) (b)		2,114		248,860
Henry Schein, Inc. (a) (b)		480		41,851
Hologic, Inc. (a) (b)		865		66,449
IDEXX Laboratories, Inc. (a) (b)		286		156,459
Intuitive Surgical, Inc. (a) (b)		1,214		366,240
Medtronic PLC (a)		4,570		507,042
PerkinElmer, Inc. (a)		421		73,448
ResMed, Inc. (a)		504		122,225
Steris PLC (a)		346		83,652
Stryker Corp. (a)		1,137		303,977
Teleflex, Inc. (a)		159		56,418
Thermo Fisher Scientific, Inc. (a)		1,343		793,243
Waters Corp. (a) (b)		213		66,113
West Pharmaceutical Services, Inc. (a)		254		104,320
Zimmer Biomet Holdings, Inc. (a)		698		89,274
				5,107,619
Health Care – Services — 2.4%
Anthem, Inc. (a)		826		405,748
Catalent, Inc. (a) (b)		611		67,760
Centene Corp. (a) (b)		2,013		169,474
Charles River Laboratories International, Inc. (a) (b)		173		49,127
DaVita, Inc. (a) (b)		189		21,378
HCA Healthcare, Inc. (a)		812		203,503
Humana, Inc. (a)		436		189,734
IQVIA Holdings, Inc. (a) (b)		674		155,836
Laboratory Corp. of America Holdings (a) (b)		313		82,526
Molina Healthcare, Inc. (a) (b)		203		67,719
Quest Diagnostics, Inc. (a)		425		58,165
UnitedHealth Group, Inc. (a)		3,211		1,637,514
Universal Health Services, Inc. Class B (a)		276		40,006
				3,148,490
Household Products & Wares — 0.2%
Avery Dennison Corp. (a)		274		47,668
Church & Dwight Co., Inc. (a)		764		75,926
The Clorox Co. (a)		443		61,590
Kimberly-Clark Corp. (a)		1,174		144,590
				329,774
Pharmaceuticals — 5.7%
AbbVie, Inc. (a)		6,025		976,713
AmerisourceBergen Corp. (a)		501		77,510
Becton Dickinson and Co. (a)		966		256,956
Bristol-Myers Squibb Co. (a)		7,420		541,883
Cardinal Health, Inc. (a)		912		51,710
Cigna Corp. (a)		1,096		262,612
CVS Health Corp. (a)		4,472		452,611
DexCom, Inc. (a) (b)		328		167,805

MML Managed Volatility Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
Eli Lilly & Co. (a)		2,706	$	774,917
Johnson & Johnson (a)		8,977		1,590,994
McKesson Corp. (a)		509		155,820
Merck & Co., Inc. (a)		8,607		706,204
Organon & Co. (a)		877		30,634
Pfizer, Inc. (a)		19,133		990,515
Viatris, Inc. (a)		4,019		43,727
Zoetis, Inc. (a)		1,607		303,064
				7,383,675
				26,247,466
Energy — 4.0%
Energy – Alternate Sources — 0.1%
Enphase Energy, Inc. (a) (b)		458		92,415
SolarEdge Technologies, Inc. (a) (b)		185		59,639
				152,054
Oil & Gas — 3.3%
APA Corp. (a)		1,210		50,009
Chevron Corp. (a)		6,570		1,069,793
ConocoPhillips (a)		4,433		443,300
Coterra Energy, Inc. (a)		2,739		73,871
Devon Energy Corp. (a)		2,196		129,849
Diamondback Energy, Inc. (a)		572		78,410
EOG Resources, Inc. (a)		2,023		241,202
Exxon Mobil Corp. (a)		14,435		1,192,187
Hess Corp. (a)		976		104,471
Marathon Oil Corp. (a)		2,698		67,747
Marathon Petroleum Corp. (a)		1,956		167,238
Occidental Petroleum Corp. (a)		3,012		170,901
Phillips 66 (a)		1,574		135,978
Pioneer Natural Resources Co. (a)		770		192,523
Valero Energy Corp. (a)		1,388		140,938
				4,258,417
Oil & Gas Services — 0.3%
Baker Hughes Co. (a)		2,991		108,902
Halliburton Co. (a)		3,156		119,518
Schlumberger NV (a)		4,769		197,007
				425,427
Pipelines — 0.3%
Kinder Morgan, Inc. (a)		6,673		126,186
ONEOK, Inc. (a)		1,477		104,321
The Williams Cos., Inc. (a)		4,151		138,685
				369,192
				5,205,090
Financial — 15.1%
Banks — 4.7%
Bank of America Corp. (a)		24,225		998,554
The Bank of New York Mellon Corp. (a)		2,485		123,331

MML Managed Volatility Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
Citigroup, Inc. (a)		6,752	$	360,557
Citizens Financial Group, Inc. (a)		1,374		62,283
Comerica, Inc. (a)		420		37,981
Fifth Third Bancorp (a)		2,394		103,038
First Republic Bank (a)		637		103,258
The Goldman Sachs Group, Inc. (a)		1,151		379,945
Huntington Bancshares, Inc. (a)		4,977		72,764
JP Morgan Chase & Co. (a)		10,073		1,373,151
KeyCorp. (a)		3,164		70,810
M&T Bank Corp. (a)		447		75,767
Morgan Stanley (a)		4,819		421,181
Northern Trust Corp. (a)		739		86,057
The PNC Financial Services Group, Inc. (a)		1,422		262,288
Regions Financial Corp. (a)		3,148		70,074
Signature Bank (a)		210		61,633
State Street Corp. (a)		1,228		106,983
SVB Financial Group (a) (b)		198		110,771
Truist Financial Corp. (a)		4,502		255,263
US Bancorp (a)		4,674		248,423
Wells Fargo & Co. (a)		13,240		641,610
Zions Bancorp NA (a)		473		31,010
				6,056,732
Diversified Financial Services — 3.8%
American Express Co. (a)		2,092		391,204
Ameriprise Financial, Inc. (a)		369		110,833
BlackRock, Inc. (a)		482		368,330
Capital One Financial Corp. (a)		1,397		183,412
Cboe Global Markets, Inc. (a)		369		42,221
The Charles Schwab Corp. (a)		5,118		431,499
CME Group, Inc. (a)		1,249		297,087
Discover Financial Services (a)		965		106,333
Franklin Resources, Inc. (a)		962		26,859
Intercontinental Exchange, Inc. (a)		1,955		258,295
Invesco Ltd. (a)		1,134		26,150
Mastercard, Inc. Class A (a)		2,940		1,050,697
Nasdaq, Inc. (a)		416		74,131
Raymond James Financial, Inc. (a)		666		73,200
Synchrony Financial (a)		1,845		64,225
T. Rowe Price Group, Inc. (a)		754		113,997
Visa, Inc. Class A (a)		5,651		1,253,222
				4,871,695
Insurance — 3.8%
Aflac, Inc. (a)		2,002		128,909
The Allstate Corp. (a)		960		132,970
American International Group, Inc. (a)		2,814		176,635
Aon PLC Class A (a)		728		237,059
Arthur J Gallagher & Co. (a)		714		124,664
Assurant, Inc. (a)		169		30,729
Berkshire Hathaway, Inc. Class B (a) (b)		6,249		2,205,335
Brown & Brown, Inc. (a)		866		62,586
Chubb Ltd. (a)		1,465		313,363
Cincinnati Financial Corp. (a)		499		67,844
Everest Re Group Ltd. (a)		134		40,385
Globe Life, Inc. (a)		335		33,701
The Hartford Financial Services Group, Inc. (a)		1,171		84,089

MML Managed Volatility Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
Lincoln National Corp. (a)		583	$	38,105
Loews Corp. (a)		650		42,133
Marsh & McLennan Cos., Inc. (a)		1,764		300,621
MetLife, Inc. (a)		2,353		165,369
Principal Financial Group, Inc. (a)		877		64,381
The Progressive Corp. (a)		1,985		226,270
Prudential Financial, Inc. (a)		1,307		154,448
The Travelers Cos., Inc. (a)		811		148,194
W.R. Berkley Corp. (a)		650		43,250
Willis Towers Watson PLC (a)		409		96,614
				4,917,654
Real Estate — 0.1%
CBRE Group, Inc. Class A (a) (b)		1,145		104,790
Real Estate Investment Trusts (REITS) — 2.7%
Alexandria Real Estate Equities, Inc. (a)		504		101,430
American Tower Corp. (a)		1,543		387,632
AvalonBay Communities, Inc. (a)		468		116,237
Boston Properties, Inc. (a)		462		59,506
Crown Castle International Corp. (a)		1,489		274,869
Digital Realty Trust, Inc. (a)		987		139,957
Duke Realty Corp. (a)		1,292		75,014
Equinix, Inc. (a)		305		226,194
Equity Residential (a)		1,214		109,163
Essex Property Trust, Inc. (a)		209		72,205
Extra Space Storage, Inc. (a)		445		91,492
Federal Realty Investment Trust (a)		250		30,517
Healthpeak Properties, Inc. (a)		1,952		67,012
Host Hotels & Resorts, Inc. (a)		2,321		45,097
Iron Mountain, Inc. (a)		951		52,695
Kimco Realty Corp. (a)		2,138		52,809
Mid-America Apartment Communities, Inc. (a)		375		78,544
Prologis, Inc. (a)		2,515		406,122
Public Storage (a)		516		201,384
Realty Income Corp. (a)		2,002		138,739
Regency Centers Corp. (a)		572		40,806
SBA Communications Corp. (a)		372		128,005
Simon Property Group, Inc. (a)		1,105		145,374
UDR, Inc. (a)		953		54,674
Ventas, Inc. (a)		1,322		81,647
Vornado Realty Trust (a)		587		26,603
Welltower, Inc. (a)		1,534		147,479
Weyerhaeuser Co. (a)		2,519		95,470
				3,446,676
Savings & Loans — 0.0%
People's United Financial, Inc. (a)		1,555		31,084
				19,428,631
Industrial — 7.7%
Aerospace & Defense — 1.6%
The Boeing Co. (a) (b)		1,860		356,190
General Dynamics Corp. (a)		780		188,120
Howmet Aerospace, Inc. (a)		1,237		44,458
L3 Harris Technologies, Inc. (a)		665		165,233

MML Managed Volatility Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
Lockheed Martin Corp. (a)		836	$	369,011
Northrop Grumman Corp. (a)		506		226,293
Raytheon Technologies Corp. (a)		5,090		504,266
Teledyne Technologies, Inc. (a) (b)		161		76,094
TransDigm Group, Inc. (a) (b)		184		119,883
				2,049,548
Building Materials — 0.4%
Carrier Global Corp. (a)		2,885		132,335
Fortune Brands Home & Security, Inc. (a)		421		31,272
Johnson Controls International PLC (a)		2,376		155,794
Martin Marietta Materials, Inc. (a)		213		81,981
Masco Corp. (a)		845		43,095
Mohawk Industries, Inc. (a) (b)		180		22,356
Vulcan Materials Co. (a)		438		80,461
				547,294
Electrical Components & Equipment — 0.3%
AMETEK, Inc. (a)		768		102,282
Emerson Electric Co. (a)		2,057		201,689
Generac Holdings, Inc. (a) (b)		224		66,586
				370,557
Electronics — 1.0%
Agilent Technologies, Inc. (a)		1,006		133,124
Allegion PLC (a)		333		36,557
Amphenol Corp. Class A (a)		2,096		157,934
Fortive Corp. (a)		1,197		72,933
Garmin Ltd. (a)		508		60,254
Honeywell International, Inc. (a)		2,321		451,620
Keysight Technologies, Inc. (a) (b)		649		102,523
Mettler-Toledo International, Inc. (a) (b)		79		108,482
TE Connectivity Ltd. (a)		1,088		142,506
Trimble, Inc. (a) (b)		824		59,443
				1,325,376
Engineering & Construction — 0.0%
Jacobs Engineering Group, Inc. (a)		464		63,944
Environmental Controls — 0.3%
Pentair PLC (a)		591		32,038
Republic Services, Inc. (a)		721		95,533
Waste Management, Inc. (a)		1,288		204,148
				331,719
Hand & Machine Tools — 0.1%
Snap-on, Inc. (a)		193		39,658
Stanley Black & Decker, Inc. (a)		552		77,164
				116,822
Machinery – Construction & Mining — 0.3%
Caterpillar, Inc. (a)		1,840		409,989

MML Managed Volatility Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
Machinery – Diversified — 0.8%
Deere & Co. (a)		954	$	396,349
Dover Corp. (a)		516		80,960
IDEX Corp. (a)		245		46,974
Ingersoll Rand, Inc. (a)		1,450		73,008
Nordson Corp. (a)		188		42,691
Otis Worldwide Corp. (a)		1,431		110,115
Rockwell Automation, Inc. (a)		397		111,172
Westinghouse Air Brake Technologies Corp. (a)		619		59,529
Xylem, Inc. (a)		596		50,815
				971,613
Miscellaneous - Manufacturing — 1.1%
3M Co. (a)		1,921		285,999
A.O. Smith Corp. (a)		487		31,114
Eaton Corp. PLC (a)		1,348		204,572
General Electric Co. (a)		3,729		341,204
Illinois Tool Works, Inc. (a)		963		201,652
Parker-Hannifin Corp. (a)		446		126,557
Textron, Inc. (a)		723		53,777
Trane Technologies PLC (a)		783		119,564
				1,364,439
Packaging & Containers — 0.2%
Amcor PLC (a)		5,354		60,661
Ball Corp. (a)		1,092		98,280
Packaging Corp. of America (a)		329		51,360
Sealed Air Corp. (a)		527		35,288
WestRock Co. (a)		853		40,116
				285,705
Shipbuilding — 0.0%
Huntington Ingalls Industries, Inc. (a)		127		25,329
Transportation — 1.6%
C.H. Robinson Worldwide, Inc. (a)		422		45,454
CSX Corp. (a)		7,548		282,672
Expeditors International of Washington, Inc. (a)		585		60,349
FedEx Corp. (a)		825		190,897
J.B. Hunt Transport Services, Inc. (a)		273		54,816
Norfolk Southern Corp. (a)		815		232,454
Old Dominion Freight Line, Inc. (a)		319		95,279
Union Pacific Corp. (a)		2,173		593,685
United Parcel Service, Inc. Class B (a)		2,488		533,576
				2,089,182
				9,951,517
Technology — 24.8%
Computers — 8.7%
Accenture PLC Class A (a)		2,152		725,719
Apple, Inc. (a)		52,850		9,228,139
Cognizant Technology Solutions Corp. Class A (a)		1,807		162,034
DXC Technology Co. (a) (b)		867		28,290
EPAM Systems, Inc. (a) (b)		199		59,025

MML Managed Volatility Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
Fortinet, Inc. (a) (b)		460	$	157,200
Hewlett Packard Enterprise Co. (a)		4,617		77,150
HP, Inc. (a)		3,637		132,023
International Business Machines Corp. (a)		3,049		396,431
Leidos Holdings, Inc. (a)		465		50,229
NetApp, Inc. (a)		764		63,412
Seagate Technology Holdings PLC (a)		674		60,593
Western Digital Corp. (a) (b)		1,035		51,388
				11,191,633
Office & Business Equipment — 0.1%
Zebra Technologies Corp. Class A (a) (b)		183		75,707
Semiconductors — 5.9%
Advanced Micro Devices, Inc. (a) (b)		5,569		608,915
Analog Devices, Inc. (a)		1,781		294,186
Applied Materials, Inc. (a)		3,018		397,772
Broadcom, Inc. (a)		1,407		885,960
Intel Corp. (a)		13,872		687,496
IPG Photonics Corp. (a) (b)		124		13,610
KLA Corp. (a)		509		186,325
Lam Research Corp. (a)		473		254,290
Microchip Technology, Inc. (a)		1,867		140,286
Micron Technology, Inc. (a)		3,811		296,839
Monolithic Power Systems, Inc. (a)		145		70,424
NVIDIA Corp. (a)		8,521		2,325,040
NXP Semiconductor NV (a)		918		169,903
Qorvo, Inc. (a) (b)		382		47,406
QUALCOMM, Inc. (a)		3,840		586,829
Skyworks Solutions, Inc. (a)		580		77,302
Teradyne, Inc. (a)		566		66,918
Texas Instruments, Inc. (a)		3,146		577,228
				7,686,729
Software — 10.1%
Activision Blizzard, Inc. (a)		2,649		212,211
Adobe, Inc. (a) (b)		1,606		731,726
Akamai Technologies, Inc. (a) (b)		541		64,590
ANSYS, Inc. (a) (b)		299		94,977
Autodesk, Inc. (a) (b)		766		164,192
Broadridge Financial Solutions, Inc. (a)		422		65,710
Cadence Design Systems, Inc. (a) (b)		939		154,428
Ceridian HCM Holding, Inc. (a) (b)		474		32,403
Cerner Corp. (a)		995		93,092
Citrix Systems, Inc. (a)		439		44,295
Electronic Arts, Inc. (a)		952		120,438
Fidelity National Information Services, Inc. (a)		2,065		207,367
Fiserv, Inc. (a) (b)		2,065		209,391
Intuit, Inc. (a)		963		463,049
Jack Henry & Associates, Inc. (a)		242		47,686
Microsoft Corp. (a)		25,559		7,880,095
MSCI, Inc. (a)		278		139,801
Oracle Corp. (a)		5,365		443,846
Paychex, Inc. (a)		1,101		150,253
Paycom Software, Inc. (a) (b)		173		59,924
PTC, Inc. (a) (b)		360		38,779
Roper Technologies, Inc. (a)		360		170,003

MML Managed Volatility Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
salesforce.com, Inc. (a) (b)		3,352	$	711,697
ServiceNow, Inc. (a) (b)		680		378,685
Synopsys, Inc. (a) (b)		517		172,301
Take-Two Interactive Software, Inc. (a) (b)		386		59,344
Tyler Technologies, Inc. (a) (b)		142		63,174
				12,973,457
				31,927,526
Utilities — 2.8%
Electric — 2.6%
AES Corp. (a)		2,230		57,378
Alliant Energy Corp. (a)		782		48,859
Ameren Corp. (a)		892		83,634
American Electric Power Co., Inc. (a)		1,729		172,502
CenterPoint Energy, Inc. (a)		2,283		69,951
CMS Energy Corp. (a)		919		64,275
Consolidated Edison, Inc. (a)		1,146		108,503
Constellation Energy Corp. (a)		1,140		64,125
Dominion Energy, Inc. (a)		2,764		234,857
DTE Energy Co. (a)		631		83,425
Duke Energy Corp. (a)		2,712		302,822
Edison International (a)		1,290		90,429
Entergy Corp. (a)		715		83,476
Evergy, Inc. (a)		861		58,841
Eversource Energy (a)		1,123		99,037
Exelon Corp. (a)		3,420		162,895
FirstEnergy Corp. (a)		1,783		81,768
NextEra Energy, Inc. (a)		6,684		566,202
NRG Energy, Inc. (a)		802		30,765
Pinnacle West Capital Corp. (a)		364		28,428
PPL Corp. (a)		2,650		75,684
Public Service Enterprise Group, Inc. (a)		1,781		124,670
Sempra Energy (a)		1,085		182,410
The Southern Co. (a)		3,533		256,178
WEC Energy Group, Inc. (a)		1,083		108,094
Xcel Energy, Inc. (a)		1,797		129,690
				3,368,898
Gas — 0.1%
Atmos Energy Corp. (a)		483		57,713
NiSource, Inc. (a)		1,431		45,506
				103,219
Water — 0.1%
American Water Works Co., Inc. (a)		628		103,953
				3,576,070
TOTAL COMMON STOCK (Cost $70,709,343)				130,517,868
TOTAL EQUITIES (Cost $70,709,343)				130,517,868

MML Managed Volatility Fund —  Portfolio of Investments (Continued)

	Principal Amount		Value
BONDS & NOTES — 0.0%
CORPORATE DEBT — 0.0%
	Number of Shares
Forest Products & Paper — 0.0%
Sino Forest Corp. 5.000% 8/01/49 (a) (c) (d)	$	359,000	$	—
TOTAL CORPORATE DEBT (Cost $0)				—
TOTAL BONDS & NOTES (Cost $0)				—
TOTAL PURCHASED OPTIONS (#) — 0.9% (Cost $2,497,565)				1,180,794
TOTAL LONG-TERM INVESTMENTS (Cost $73,206,908)				131,698,662
	Principal Amount
SHORT-TERM INVESTMENTS — 1.4%
Repurchase Agreement — 1.4%
Fixed Income Clearing Corp., Repurchase Agreement, dated 3/31/22, 0.000%, due 4/01/22 (e)	$	1,802,855		1,802,855
TOTAL SHORT-TERM INVESTMENTS (Cost $1,802,855)				1,802,855
TOTAL INVESTMENTS — 103.8% (Cost $75,009,763) (f)				133,501,517
Other Assets/(Liabilities) — (3.8)%				(4,855,669)
NET ASSETS — 100.0%			$	128,645,848

Notes to Portfolio of Investments

Percentages are stated as a percent of net assets.

(a) All or a portion of this security is pledged/held as collateral for open derivatives.

(b) Non-income producing security.

(c) Investment was valued using significant unobservable inputs.

(d) This security is fair valued in good faith in accordance with procedures approved by the Board of Trustees. At March 31, 2022, these securities amounted to a value of $0 or 0.00% of net assets.

(e) Maturity value of $1,802,855. Collateralized by U.S. Government Agency obligations with a rate of 0.500%, maturity date of 4/15/24, and an aggregate market value, including accrued interest, of $1,838,936.

(f) See Note 3 for aggregate cost for federal tax purposes.

(#) Exchange-Traded Options Purchased
Description	Expiration Date	Exercise Price		No. of Contracts		Notional Amount		Value		Premiums Paid (Received)		Unrealized Appreciation/ (Depreciation)
Put
S&P 500 Index	5/20/22		4,000.00		42	USD	16,800,000	$	93,870	$	245,524	$	(151,654)
S&P 500 Index	5/20/22		4,025.00		42	USD	16,905,000		106,218		381,381		(275,163)
S&P 500 Index	5/20/22		4,050.00		41	USD	16,605,000		111,930		338,858		(226,928)
S&P 500 Index	6/17/22		3,950.00		42	USD	16,590,000		171,360		226,905		(55,545)
S&P 500 Index	6/17/22		4,000.00		42	USD	16,800,000		184,800		261,219		(76,419)
S&P 500 Index	6/17/22		4,025.00		36	USD	14,490,000		170,568		195,929		(25,361)
S&P 500 Index	9/16/22		3,800.00		42	USD	15,960,000		342,048		847,749		(505,701)
								$	1,180,794	$	2,497,565	$	(1,316,771)

Exchange-Traded Options Written
Description	Expiration Date	Exercise Price		No. of Contracts		Notional Amount		Value		Premiums Paid (Received)		Unrealized Appreciation/ (Depreciation)
Call
S&P 500 Index	4/29/22		4,550.00		32	USD	14,560,000	$	(270,176)	$	(253,871)	$	(16,305)
S&P 500 Index	4/29/22		4,600.00		31	USD	14,260,000		(166,160)		(185,179)		19,019
S&P 500 Index	5/20/22		4,500.00		32	USD	14,400,000		(472,224)		(296,784)		(175,440)
S&P 500 Index	5/20/22		4,550.00		33	USD	15,015,000		(376,596)		(309,787)		(66,809)
S&P 500 Index	5/20/22		4,600.00		32	USD	14,720,000		(275,840)		(250,992)		(24,848)
S&P 500 Index	5/20/22		4,700.00		32	USD	15,040,000		(145,984)		(153,520)		7,536
S&P 500 Index	5/20/22		4,750.00		32	USD	15,200,000		(87,296)		(143,920)		56,624
S&P 500 Index	6/17/22		4,650.00		31	USD	14,415,000		(285,727)		(267,917)		(17,810)
S&P 500 Index	6/17/22		4,700.00		32	USD	15,040,000		(222,048)		(330,348)		108,300
								$	(2,302,051)	$	(2,192,318)	$	(109,733)

MML Mid Cap Growth Fund —  Portfolio of Investments

March 31, 2022 (Unaudited)

	Number of Shares		Value
EQUITIES — 96.6%
COMMON STOCK — 96.2%
Basic Materials — 0.5%
Chemicals — 0.5%
RPM International, Inc.		22,500	$	1,832,400
Communications — 6.2%
Advertising — 0.5%
The Trade Desk, Inc. Class A (a)		27,200		1,883,600
Internet — 4.5%
CDW Corp.		22,441		4,014,470
DoorDash, Inc., Class A (a)		4,800		562,512
Etsy, Inc. (a) (b)		8,100		1,006,668
Farfetch Ltd. Class A (a)		21,424		323,931
IAC/InterActiveCorp (a)		8,500		852,380
Match Group, Inc. (a)		43,111		4,687,890
Palo Alto Networks, Inc. (a)		6,367		3,963,521
Spotify Technology SA (a)		12,300		1,857,546
Upwork, Inc. (a)		9,900		230,076
Vimeo, Inc. (a)		29,429		349,617
				17,848,611
Media — 0.7%
Liberty Media Corp-Liberty Formula One Class C (a)		39,100		2,730,744
Telecommunications — 0.5%
Corning, Inc.		49,400		1,823,354
				24,286,309
Consumer, Cyclical — 14.6%
Airlines — 0.9%
Southwest Airlines Co. (a)		73,000		3,343,400
Apparel — 0.2%
Deckers Outdoor Corp. (a)		3,300		903,441
Auto Manufacturers — 1.3%
Rivian Automotive, Inc., Lockup Shares (Acquired 12/23/19 - 7/10/20, Cost $1,280,667) (a) (d) (e)		108,363		5,171,949
Auto Parts & Equipment — 0.2%
Aptiv PLC (a)		4,600		550,666
Entertainment — 0.6%
DraftKings, Inc. Class A (a) (b)		36,600		712,602

MML Mid Cap Growth Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
Vail Resorts, Inc.		6,700	$	1,743,809
				2,456,411
Leisure Time — 0.1%
Peloton Interactive, Inc. Class A (a)		18,354		484,913
Lodging — 2.6%
Hilton Worldwide Holdings, Inc. (a)		50,357		7,641,171
MGM Resorts International		58,400		2,449,296
				10,090,467
Retail — 8.7%
Bath & Body Works, Inc.		24,400		1,166,320
Burlington Stores, Inc. (a)		21,500		3,916,655
Casey's General Stores, Inc.		17,600		3,487,792
Chipotle Mexican Grill, Inc. (a)		2,050		3,243,161
Dollar General Corp.		18,000		4,007,340
Dollar Tree, Inc. (a)		18,700		2,994,805
Domino's Pizza, Inc.		5,000		2,035,050
Five Below, Inc. (a)		6,900		1,092,753
Freshpet, Inc. (a)		23,597		2,421,996
Lululemon Athletica, Inc. (a)		8,308		3,034,331
O'Reilly Automotive, Inc. (a)		3,900		2,671,344
Olaplex Holdings, Inc. (a)		18,148		283,653
Ross Stores, Inc.		12,700		1,148,842
Warby Parker, Inc. Class A (a)		4,843		163,742
Wingstop, Inc.		23,771		2,789,527
				34,457,311
				57,458,558
Consumer, Non-cyclical — 33.9%
Beverages — 0.3%
The Boston Beer Co., Inc. Class A (a)		3,294		1,279,620
Biotechnology — 3.1%
Alnylam Pharmaceuticals, Inc. (a)		14,000		2,286,060
Apellis Pharmaceuticals, Inc. (a)		3,182		161,677
Argenx SE ADR (a)		5,809		1,831,636
Exelixis, Inc. (a)		18,000		408,060
Ionis Pharmaceuticals, Inc. (a)		38,500		1,426,040
Seagen, Inc. (a)		38,633		5,565,084
Ultragenyx Pharmaceutical, Inc. (a)		8,100		588,222
				12,266,779
Commercial Services — 11.4%
Bright Horizons Family Solutions, Inc. (a)		11,200		1,486,128
Chegg, Inc. (a) (b)		100,191		3,634,929
Clarivate PLC (a) (b)		102,082		1,710,894
CoStar Group, Inc. (a)		36,554		2,434,862
Equifax, Inc.		10,800		2,560,680
FleetCor Technologies, Inc. (a)		15,300		3,810,618
Gartner, Inc. (a)		21,577		6,418,294
MarketAxess Holdings, Inc.		11,063		3,763,633
Multiplan Corp. (a) (b)		129,800		607,464

MML Mid Cap Growth Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
Paylocity Holding Corp. (a)		1,400	$	288,078
Shift4 Payments, Inc. Class A (a)		60,003		3,715,986
Terminix Global Holdings, Inc. (a)		53,000		2,418,390
TransUnion		92,523		9,561,327
Verisk Analytics, Inc.		10,900		2,339,467
				44,750,750
Food — 0.2%
TreeHouse Foods, Inc. (a)		28,800		929,088
Health Care – Products — 12.0%
Alcon, Inc. (b)		27,900		2,213,307
Align Technology, Inc. (a)		6,799		2,964,364
Avantor, Inc. (a)		128,800		4,356,016
Bruker Corp.		67,400		4,333,820
The Cooper Cos., Inc.		10,900		4,551,731
Dentsply Sirona, Inc.		14,100		694,002
Exact Sciences Corp. (a) (b)		12,800		894,976
Hologic, Inc. (a)		117,900		9,057,078
ICU Medical, Inc. (a)		8,000		1,781,120
Omnicell, Inc. (a)		31,559		4,086,575
Ortho Clinical Diagnostics Holdings PLC Class H (a)		52,453		978,773
Quidel Corp. (a)		15,500		1,743,130
Teleflex, Inc.		19,240		6,826,929
West Pharmaceutical Services, Inc.		7,235		2,971,487
				47,453,308
Health Care – Services — 4.5%
Acadia Healthcare Co., Inc. (a)		45,700		2,994,721
agilon health, Inc. (a) (b)		8,756		221,965
Catalent, Inc. (a)		64,600		7,164,140
ICON PLC (a)		23,956		5,826,578
Molina Healthcare, Inc. (a)		4,500		1,501,155
				17,708,559
Household Products & Wares — 1.0%
Avery Dennison Corp.		17,200		2,992,284
Reynolds Consumer Products, Inc.		35,600		1,044,504
				4,036,788
Pharmaceuticals — 1.4%
Alkermes PLC (a)		59,100		1,554,921
Elanco Animal Health, Inc. (a)		51,400		1,341,026
Neurocrine Biosciences, Inc. (a)		12,900		1,209,375
Perrigo Co. PLC		37,000		1,421,910
				5,527,232
				133,952,124
Energy — 0.3%
Energy – Alternate Sources — 0.1%
Shoals Technologies Group, Inc. Class A (a) (b)		23,000		391,920

MML Mid Cap Growth Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
Oil & Gas — 0.2%
Venture Global LNG, Inc., Series B (Acquired 3/08/18, Cost $63,420) (a) (c) (d) (e)		21	$	124,954
Venture Global LNG, Inc., Series C (Acquired 10/16/17-3/08/18, Cost $530,257) (a) (c) (d) (e)		144		856,824
				981,778
				1,373,698
Financial — 5.0%
Diversified Financial Services — 2.6%
Cboe Global Markets, Inc.		17,800		2,036,676
Hamilton Lane, Inc. Class A		45,981		3,553,872
Raymond James Financial, Inc.		8,900		978,199
SoFi Technologies, Inc. (a) (b)		36,561		345,501
Tradeweb Markets, Inc. Class A		38,500		3,382,995
				10,297,243
Insurance — 1.5%
Assurant, Inc.		17,800		3,236,574
Axis Capital Holdings Ltd.		32,500		1,965,275
Kemper Corp.		10,200		576,708
				5,778,557
Private Equity — 0.9%
KKR & Co., Inc.		58,900		3,443,883
Real Estate — 0.0%
eXp World Holdings, Inc. (b)		7,300		154,541
				19,674,224
Industrial — 17.2%
Aerospace & Defense — 0.8%
Hexcel Corp.		54,215		3,224,166
Building Materials — 1.4%
Fortune Brands Home & Security, Inc.		40,429		3,003,066
Martin Marietta Materials, Inc.		6,760		2,601,857
				5,604,923
Electrical Components & Equipment — 0.1%
Littelfuse, Inc.		1,900		473,879
Electronics — 3.9%
Agilent Technologies, Inc.		39,900		5,279,967
Amphenol Corp. Class A		24,100		1,815,935
Fortive Corp.		41,200		2,510,316
Keysight Technologies, Inc. (a)		23,400		3,696,498
National Instruments Corp.		54,500		2,212,155
				15,514,871
Environmental Controls — 0.5%
Waste Connections, Inc.		13,900		1,941,830

MML Mid Cap Growth Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
Machinery – Construction & Mining — 1.3%
BWX Technologies, Inc.		90,773	$	4,889,034
Machinery – Diversified — 3.3%
Cognex Corp.		18,100		1,396,415
Colfax Corp. (a)		80,636		3,208,506
IDEX Corp.		11,800		2,262,414
Ingersoll Rand, Inc.		121,700		6,127,595
				12,994,930
Miscellaneous - Manufacturing — 2.0%
Textron, Inc.		104,000		7,735,520
Packaging & Containers — 2.7%
Ardagh Metal Packaging SA (a)		47,137		383,224
Ball Corp.		80,047		7,204,230
Sealed Air Corp.		48,100		3,220,776
				10,808,230
Transportation — 1.2%
J.B. Hunt Transport Services, Inc.		23,100		4,638,249
				67,825,632
Technology — 18.5%
Computers — 3.1%
Crowdstrike Holdings, Inc. Class A (a)		7,800		1,771,224
Fortinet, Inc. (a)		14,400		4,921,056
Leidos Holdings, Inc.		9,600		1,036,992
Thoughtworks Holding, Inc. (a)		9,956		207,185
Varonis Systems, Inc. (a)		88,189		4,192,505
				12,128,962
Semiconductors — 7.2%
Entegris, Inc.		7,200		945,072
KLA Corp.		20,178		7,386,359
Lattice Semiconductor Corp. (a)		9,300		566,835
Marvell Technology, Inc.		90,900		6,518,439
Microchip Technology, Inc.		100,700		7,566,598
Skyworks Solutions, Inc.		15,200		2,025,856
Synaptics, Inc. (a)		14,067		2,806,366
Wolfspeed, Inc. (a)		4,300		489,598
				28,305,123
Software — 8.2%
Atlassian Corp. PLC Class A (a)		4,400		1,292,852
Avalara, Inc. (a)		39,242		3,904,971
Bill.com Holdings, Inc. (a) (b)		6,600		1,496,814
Black Knight, Inc. (a)		32,900		1,907,871
Broadridge Financial Solutions, Inc.		5,000		778,550
CCC Intelligent Solutions Holdings, Inc. (a)		75,271		830,992
Ceridian HCM Holding, Inc. (a)		86,441		5,909,107
Clear Secure Inc. Class A (a) (b)		5,493		147,652
DocuSign, Inc. (a)		13,300		1,424,696
Doximity, Inc. Class A (a) (b)		11,400		593,826

MML Mid Cap Growth Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
Electronic Arts, Inc.		3,900	$	493,389
HashiCorp, Inc. Class A (a) (b)		519		28,026
nCino, Inc. (a)		6,900		282,762
Playtika Holding Corp. (a)		23,537		454,970
Procore Technologies, Inc. (a)		3,703		214,626
PTC, Inc. (a)		15,400		1,658,888
Roper Technologies, Inc.		5,800		2,738,934
SentinelOne Inc. Class A (a)		10,076		390,344
Synopsys, Inc. (a)		3,900		1,299,753
Veeva Systems, Inc. Class A (a)		16,000		3,399,360
ZoomInfo Technologies, Inc. (a)		56,128		3,353,088
				32,601,471
				73,035,556
TOTAL COMMON STOCK (Cost $297,371,097)				379,438,501
PREFERRED STOCK — 0.4%
Consumer, Cyclical — 0.0%
Auto Manufacturers — 0.0%
Nuro, Inc., Series D (Acquired 10/29/21, Cost $1,59,220)) (a) (c) (d) (e)		7,638		159,220
Consumer, Non-cyclical — 0.1%
Commercial Services — 0.0%
Redwood Materials, Inc., Series C (Acquired 5/28/21, Cost $200,090) (a) (c) (d) (e)		4,221		220,125
Health Care – Services — 0.1%
Caris Life Sciences, Inc., Series D (Acquired 5/11/21, Cost $271,366) (a) (c) (d) (e)		33,502		271,366
				491,491
Financial — 0.1%
Investment Companies — 0.1%
Maplebear, Inc., Series E (Acquired 11/19/21 - 12/01/21, Cost $528,790) (a) (c) (d) (e)		4,398		422,208
Maplebear, Inc., Series I (Acquired 2/26/21, Cost $95,750) (a) (c) (d) (e)		766		73,536
				495,744
Industrial — 0.1%
Electrical Components & Equipment — 0.1%
Sila Nanotechnologies, Inc., Series F (Acquired 1/07/21, Cost $407,982) (a) (c) (d) (e)		9,885		308,115
Technology — 0.1%
Software — 0.1%
Databricks, Inc., Series G (Acquired 2/01/21, Cost $133,381) (a) (c) (d) (e)		752		124,742

MML Mid Cap Growth Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
Databricks, Inc., Series H (Acquired 8/31/21, Cost $186,503) (a) (c) (d) (e)		846	$	140,334
				265,076
TOTAL PREFERRED STOCK (Cost $1,983,082)				1,719,646
TOTAL EQUITIES (Cost $299,354,179)				381,158,147
MUTUAL FUNDS — 1.0%
Diversified Financial Services — 1.0%
State Street Navigator Securities Lending Government Money Market Portfolio (f)		4,007,609		4,007,609
TOTAL MUTUAL FUNDS (Cost $4,007,609)				4,007,609
TOTAL LONG-TERM INVESTMENTS (Cost $303,361,788)				385,165,756
	Number of Shares
SHORT-TERM INVESTMENTS — 3.0%
Mutual Fund — 0.0%
T. Rowe Price Treasury Reserve Fund		102	$	102
	Principal Amount
Repurchase Agreement — 3.0%
Fixed Income Clearing Corp., Repurchase Agreement, dated 3/31/22, 0.000%, due 4/01/22 (g)	$	11,692,289		11,692,289
TOTAL SHORT-TERM INVESTMENTS (Cost $11,692,391)				11,692,391
TOTAL INVESTMENTS — 100.6% (Cost $315,054,179) (h)				396,858,147
Other Assets/(Liabilities) — (0.6)%				(2,216,152)
NET ASSETS — 100.0%			$	394,641,995

Abbreviation Legend

ADR American Depositary Receipt

Notes to Portfolio of Investments

Percentages are stated as a percent of net assets.

(a) Non-income producing security.

(b) Denotes all or a portion of security on loan. The total value of securities on loan as of March 31, 2022, was $13,636,388 or 3.46% of net assets. Total securities on loan may be less than the amounts identified in the Portfolio of Investments. The Fund received $9,990,330 of non-cash collateral (U.S. Treasury and/or Agency obligations) related to securities lending activity. This amount is not reflected on the Portfolio of Investments. (Note 2).

(c) Investment was valued using significant unobservable inputs.

(d) Restricted security. Certain securities are restricted as to resale. At March 31, 2022, these securities amounted to a value of $7,873,373 or 2.00% of net assets. The Fund generally bears the costs, if any, associated with the disposition of restricted securities.

(e) This security is fair valued in good faith in accordance with procedures approved by the Board of Trustees. At March 31, 2022, these securities amounted to a value of $7,873,373 or 2.00% of net assets.

(f) Represents investment of security lending cash collateral. (Note 2).

(g) Maturity value of $11,692,289. Collateralized by U.S. Government Agency obligations with rates ranging from 0.375% - 0.500%, maturity dates ranging from 4/15/24 - 9/15/24, and an aggregate market value, including accrued interest, of $11,926,154.

(h) See Note 3 for aggregate cost for federal tax purposes.

MML Mid Cap Value Fund —  Portfolio of Investments

March 31, 2022 (Unaudited)

	Number of Shares		Value
EQUITIES — 98.0%
COMMON STOCK — 98.0%
Basic Materials — 2.0%
Chemicals — 0.9%
Axalta Coating Systems Ltd. (a)		118,854	$	2,921,431
Forest Products & Paper — 1.1%
Mondi PLC		201,670		3,909,615
				6,831,046
Communications — 2.9%
Internet — 1.1%
F5, Inc. (a)		18,034		3,768,204
Media — 1.5%
Fox Corp. Class B		141,752		5,142,763
Telecommunications — 0.3%
Juniper Networks, Inc.		25,966		964,897
				9,875,864
Consumer, Cyclical — 14.2%
Airlines — 2.1%
Southwest Airlines Co. (a)		158,823		7,274,093
Auto Manufacturers — 1.2%
Cummins, Inc.		8,010		1,642,931
PACCAR, Inc.		28,341		2,495,992
				4,138,923
Auto Parts & Equipment — 2.0%
Aptiv PLC (a)		2,820		337,582
BorgWarner, Inc.		125,614		4,886,385
Bridgestone Corp.		39,800		1,540,318
				6,764,285
Food Services — 1.0%
Sodexo SA		41,783		3,392,068
Leisure Time — 0.8%
Polaris, Inc. (b)		25,359		2,670,810
Retail — 7.1%
Advance Auto Parts, Inc.		33,393		6,911,015
Beacon Roofing Supply, Inc. (a)		42,948		2,545,957
Cracker Barrel Old Country Store, Inc.		16,092		1,910,603

MML Mid Cap Value Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
Dollar Tree, Inc. (a)		43,397	$	6,950,030
MSC Industrial Direct Co., Inc. Class A		69,404		5,913,915
				24,231,520
				48,471,699
Consumer, Non-cyclical — 22.8%
Commercial Services — 0.3%
Euronet Worldwide, Inc. (a)		8,322		1,083,108
Food — 7.0%
Conagra Brands, Inc.		201,787		6,773,990
General Mills, Inc.		35,887		2,430,268
The J.M. Smucker Co.		17,896		2,423,297
Kellogg Co.		34,773		2,242,511
Koninklijke Ahold Delhaize NV		179,138		5,758,612
Orkla ASA		247,867		2,204,364
Sysco Corp.		23,367		1,907,915
				23,740,957
Health Care – Products — 6.3%
Dentsply Sirona, Inc.		32,213		1,585,524
Envista Holdings Corp. (a)		22,894		1,115,167
Henry Schein, Inc. (a)		59,175		5,159,468
Koninklijke Philips NV		60,846		1,857,628
Zimmer Biomet Holdings, Inc.		90,844		11,618,948
Zimvie, Inc. (a)		8,918		203,687
				21,540,422
Health Care – Services — 4.0%
Humana, Inc.		2,414		1,050,500
Quest Diagnostics, Inc.		45,342		6,205,506
Universal Health Services, Inc. Class B		44,044		6,384,178
				13,640,184
Household Products & Wares — 1.4%
Kimberly-Clark Corp.		39,718		4,891,669
Pharmaceuticals — 3.8%
AmerisourceBergen Corp.		16,955		2,623,108
Becton Dickinson and Co.		12,910		3,434,060
Cardinal Health, Inc.		71,756		4,068,565
McKesson Corp.		9,257		2,833,846
				12,959,579
				77,855,919
Energy — 5.9%
Oil & Gas — 3.4%
Devon Energy Corp.		101,291		5,989,337
Pioneer Natural Resources Co.		22,376		5,594,671
				11,584,008

MML Mid Cap Value Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
Oil & Gas Services — 1.4%
Baker Hughes Co.		135,208	$	4,922,924
Pipelines — 1.1%
Enterprise Products Partners LP		142,599		3,680,480
				20,187,412
Financial — 25.7%
Banks — 10.2%
The Bank of New York Mellon Corp.		141,280		7,011,726
Commerce Bancshares, Inc.		7,903		565,776
First Hawaiian, Inc.		151,876		4,235,822
M&T Bank Corp.		7,073		1,198,873
Northern Trust Corp.		71,300		8,302,885
Prosperity Bancshares, Inc.		66,360		4,604,057
State Street Corp.		18,730		1,631,758
Truist Financial Corp.		87,651		4,969,812
Westamerica Bancorp.		35,827		2,167,533
				34,688,242
Diversified Financial Services — 2.1%
Ameriprise Financial, Inc.		6,901		2,072,784
T. Rowe Price Group, Inc.		34,942		5,282,881
				7,355,665
Insurance — 6.1%
Aflac, Inc.		68,572		4,415,351
The Allstate Corp.		49,430		6,846,550
Chubb Ltd.		16,139		3,452,132
Reinsurance Group of America, Inc.		54,478		5,963,162
				20,677,195
Real Estate Investment Trusts (REITS) — 6.9%
Equinix, Inc.		5,447		4,039,604
Essex Property Trust, Inc.		7,214		2,492,293
Healthcare Trust of America, Inc. Class A		58,501		1,833,421
Healthpeak Properties, Inc.		170,971		5,869,434
MGM Growth Properties LLC Class A		90,961		3,520,191
Regency Centers Corp.		60,632		4,325,487
Weyerhaeuser Co.		36,624		1,388,050
				23,468,480
Savings & Loans — 0.4%
Capitol Federal Financial, Inc.		126,975		1,381,488
				87,571,070
Industrial — 12.4%
Aerospace & Defense — 1.0%
BAE Systems PLC		186,168		1,754,905
General Dynamics Corp.		6,674		1,609,635
				3,364,540

MML Mid Cap Value Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
Building Materials — 0.7%
Cie de Saint-Gobain		42,858	$	2,550,483
Electrical Components & Equipment — 2.1%
Emerson Electric Co.		71,848		7,044,696
Electronics — 2.4%
Hubbell, Inc.		7,947		1,460,420
nVent Electric PLC		152,296		5,296,855
TE Connectivity Ltd.		11,752		1,539,277
				8,296,552
Environmental Controls — 0.5%
Republic Services, Inc.		12,540		1,661,550
Hand & Machine Tools — 0.3%
Stanley Black & Decker, Inc.		7,911		1,105,879
Machinery – Construction & Mining — 1.2%
Oshkosh Corp.		39,937		4,019,659
Machinery – Diversified — 0.2%
IMI PLC		48,245		859,835
Packaging & Containers — 3.4%
Amcor PLC		197,145		2,233,653
Packaging Corp. of America		25,358		3,958,637
Sonoco Products Co.		86,077		5,384,977
				11,577,267
Transportation — 0.6%
Heartland Express, Inc.		138,412		1,947,457
				42,427,918
Technology — 5.0%
Computers — 2.4%
Amdocs Ltd.		37,257		3,062,898
HP, Inc.		136,340		4,949,142
				8,012,040
Software — 2.6%
CDK Global, Inc.		63,637		3,097,849
Cerner Corp.		22,814		2,134,478
Open Text Corp.		86,161		3,653,226
				8,885,553
				16,897,593
Utilities — 7.1%
Electric — 5.7%
Edison International		92,343		6,473,244
Evergy, Inc.		24,084		1,645,901
Eversource Energy		21,317		1,879,946
NorthWestern Corp. (b)		72,805		4,403,974
Pinnacle West Capital Corp.		48,034		3,751,455

MML Mid Cap Value Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
Xcel Energy, Inc.		16,774	$	1,210,580
				19,365,100
Gas — 1.4%
Atmos Energy Corp.		18,410		2,199,811
Spire, Inc.		37,459		2,688,058
				4,887,869
				24,252,969
TOTAL COMMON STOCK (Cost $291,231,177)				334,371,490
TOTAL EQUITIES (Cost $291,231,177)				334,371,490
MUTUAL FUNDS — 2.1%
Diversified Financial Services — 2.1%
iShares Russell Mid-Cap Value ETF (b)		58,477		6,996,188
State Street Navigator Securities Lending Government Money Market Portfolio (c)		2,365		2,365
				6,998,553
TOTAL MUTUAL FUNDS (Cost $6,816,952)				6,998,553
TOTAL LONG-TERM INVESTMENTS (Cost $298,048,129)				341,370,043
	Principal Amount
SHORT-TERM INVESTMENTS — 2.0%
Repurchase Agreement — 2.0%
Fixed Income Clearing Corp., Repurchase Agreement, dated 3/31/22, 0.000%, due 4/01/22 (d)	$	6,728,021		6,728,021
TOTAL SHORT-TERM INVESTMENTS (Cost $6,728,021)				6,728,021
TOTAL INVESTMENTS — 102.1% (Cost $304,776,150) (e)				348,098,064
Other Assets/(Liabilities) — (2.1)%				(7,035,457)
NET ASSETS — 100.0%			$	341,062,607

Abbreviation Legend

ETF Exchange-Traded Fund

Notes to Portfolio of Investments

Percentages are stated as a percent of net assets.

(a) Non-income producing security.

(b) Denotes all or a portion of security on loan. The total value of securities on loan as of March 31, 2022, was $5,941,045 or 1.74% of net assets. Total securities on loan may be less than the amounts identified in the Portfolio of Investments. The Fund received $6,059,322 of non-cash collateral (U.S. Treasury and/or Agency obligations) related to securities lending activity. This amount is not reflected on the Portfolio of Investments. (Note 2).

(c) Represents investment of security lending cash collateral. (Note 2).

(d) Maturity value of $6,728,021. Collateralized by U.S. Government Agency obligations with a rate of 0.500%, maturity date of 4/15/24, and an aggregate market value, including accrued interest, of $6,862,695.

(e) See Note 3 for aggregate cost for federal tax purposes.

Forward contracts
Counterparty	Settlement Date	Currency Purchased		Currency Sold		Unrealized Appreciation (Depreciation)
Bank of America N.A.	6/30/22	GBP	472,159	USD	$621,479	$	(1,400)
Bank of America N.A.	6/30/22	USD	6,381,263	GBP	4,818,775		52,840
Bank of America N.A.	6/30/22	USD	1,301,499	JPY	156,666,730		11,458
JP Morgan Chase Bank NA	6/30/22	USD	11,643,164	EUR	10,532,082		(48,224)
UBS AG	6/30/22	USD	1,882,024	NOK	16,543,139		3,831
						$	18,505

Currency Legend

EUR Euro

GBP British Pound

JPY Japanese Yen

NOK Norwegian Krone

USD U.S. Dollar

MML Small Cap Growth Equity Fund —  Portfolio of Investments

March 31, 2022 (Unaudited)

	Number of Shares		Value
EQUITIES — 95.2%
COMMON STOCK — 95.2%
Basic Materials — 1.4%
Chemicals — 1.4%
Cabot Corp.		21,125	$	1,445,161
Methanex Corp.		19,767		1,078,488
				2,523,649
				2,523,649
Communications — 7.2%
Internet — 5.1%
Cargurus, Inc. (a)		68,108		2,891,866
Criteo SA Sponsored ADR (a)		20,605		561,280
Perficient, Inc. (a)		20,381		2,243,744
The RealReal, Inc. (a)		9,864		71,613
Revolve Group, Inc. (a)		24,080		1,292,855
Shutterstock, Inc.		21,742		2,023,746
				9,085,104
Media — 0.6%
The New York Times Co. Class A		21,944		1,005,913
Telecommunications — 1.5%
Calix, Inc. (a)		40,717		1,747,166
Viavi Solutions, Inc. (a)		58,234		936,403
				2,683,569
				12,774,586
Consumer, Cyclical — 9.2%
Apparel — 1.4%
Carter's, Inc.		4,784		440,080
Crocs, Inc. (a)		8,781		670,868
Deckers Outdoor Corp. (a)		1,205		329,893
Steven Madden Ltd.		24,889		961,711
				2,402,552
Distribution & Wholesale — 0.6%
WESCO International, Inc. (a)		7,676		998,955
Entertainment — 0.4%
Cinemark Holdings, Inc. (a)		45,588		787,761
Food Services — 0.5%
Sovos Brands, Inc. (a)		63,286		897,395

MML Small Cap Growth Equity Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
Home Builders — 1.3%
Cavco Industries, Inc. (a)		971	$	233,865
Skyline Champion Corp. (a)		38,533		2,114,691
				2,348,556
Leisure Time — 0.5%
Acushnet Holdings Corp.		9,487		381,947
Planet Fitness, Inc. Class A (a)		6,036		509,921
				891,868
Lodging — 0.5%
Boyd Gaming Corp.		13,887		913,487
Retail — 4.0%
Denny's Corp. (a)		46,478		665,100
Floor & Decor Holdings, Inc. Class A (a)		5,741		465,021
Freshpet, Inc. (a)		7,154		734,287
La-Z-Boy, Inc.		1,600		42,192
National Vision Holdings, Inc. (a)		27,195		1,184,886
Nu Skin Enterprises, Inc. Class A		19,372		927,531
Patrick Industries, Inc.		12,181		734,514
Rush Enterprises, Inc. Class A		18,503		941,988
Victoria's Secret & Co. (a)		16,090		826,383
Wingstop, Inc.		5,284		620,077
				7,141,979
				16,382,553
Consumer, Non-cyclical — 22.0%
Biotechnology — 4.9%
Akero Therapeutics, Inc. (a)		13,017		184,711
Amicus Therapeutics, Inc. (a)		80,285		760,299
Apellis Pharmaceuticals, Inc. (a)		17,260		876,981
BioAtla, Inc. (a)		5,675		28,375
Blueprint Medicines Corp. (a)		7,271		464,471
Celldex Therapeutics, Inc. (a)		13,567		462,092
Crinetics Pharmaceuticals, Inc. (a)		7,350		161,332
Fate Therapeutics, Inc. (a)		7,249		281,044
ImmunoGen, Inc. (a)		46,857		223,039
Intra-Cellular Therapies, Inc. (a)		7,129		436,224
Karuna Therapeutics, Inc. (a)		3,715		471,025
Kodiak Sciences, Inc. (a)		3,273		25,268
Kymera Therapeutics, Inc. (a)		13,709		580,165
Myriad Genetics, Inc. (a)		25,390		639,828
NeoGenomics, Inc. (a)		19,086		231,895
PTC Therapeutics, Inc. (a)		8,716		325,194
RAPT Therapeutics, Inc. (a)		7,796		171,434
Relay Therapeutics, Inc. (a)		12,825		383,852
Revolution Medicines, Inc. (a)		19,020		485,200
Rocket Pharmaceuticals, Inc. (a)		23,436		371,695
Sage Therapeutics, Inc. (a)		5,800		191,980
Turning Point Therapeutics, Inc. (a)		14,030		376,705
Veracyte, Inc. (a)		21,538		593,803
				8,726,612

MML Small Cap Growth Equity Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
Commercial Services — 3.9%
Chegg, Inc. (a) (b)		22,310	$	809,407
Cross Country Healthcare, Inc. (a)		30,121		652,722
LiveRamp Holdings, Inc. (a)		17,169		641,949
Mister Car Wash, Inc. (a)		50,286		743,730
R1 RCM, Inc. (a)		56,625		1,515,285
Repay Holdings Corp. (a)		36,295		536,077
TriNet Group, Inc. (a)		20,619		2,028,085
				6,927,255
Cosmetics & Personal Care — 0.5%
elf Beauty, Inc. (a)		33,160		856,523
Food — 0.6%
Performance Food Group Co. (a)		21,147		1,076,594
Health Care – Products — 6.1%
Glaukos Corp. (a)		15,039		869,555
Globus Medical, Inc. Class A (a)		24,369		1,797,945
Haemonetics Corp. (a)		15,565		984,019
ICU Medical, Inc. (a)		3,226		718,237
Inari Medical, Inc. (a)		10,300		933,592
Inspire Medical Systems, Inc. (a)		7,025		1,803,247
Integra LifeSciences Holdings Corp. (a)		16,889		1,085,287
NanoString Technologies, Inc. (a)		21,294		739,966
Omnicell, Inc. (a)		9,038		1,170,331
Tandem Diabetes Care, Inc. (a)		6,736		783,329
				10,885,508
Health Care – Services — 2.4%
Acadia Healthcare Co., Inc. (a)		14,404		943,894
Amedisys, Inc. (a)		5,890		1,014,788
LHC Group, Inc. (a)		13,751		2,318,419
				4,277,101
Household Products & Wares — 0.5%
Spectrum Brands Holdings, Inc.		9,559		848,074
Pharmaceuticals — 3.1%
Aclaris Therapeutics, Inc. (a)		24,120		415,829
Ascendis Pharma A/S ADR (a)		4,415		518,144
BellRing Brands, Inc. (a)		26,465		610,812
Covetrus, Inc. (a)		42,735		717,521
Intellia Therapeutics, Inc. (a)		7,283		529,256
Madrigal Pharmaceuticals, Inc. (a)		3,302		323,992
Myovant Sciences Ltd. (a)		18,146		241,705
Owens & Minor, Inc.		39,118		1,721,974
Pacira BioSciences, Inc. (a)		5,357		408,846
Y-mAbs Therapeutics, Inc. (a)		14,796		175,777
				5,663,856
				39,261,523

MML Small Cap Growth Equity Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
Energy — 4.9%
Energy – Alternate Sources — 1.3%
Atlantica Sustainable Infrastructure PLC		27,830	$	975,998
First Solar, Inc. (a)		9,760		817,303
Maxeon Solar Technologies Ltd. (a) (b)		31,738		459,566
				2,252,867
Oil & Gas — 3.6%
Brigham Minerals, Inc. Class A		35,089		896,524
Chesapeake Energy Corp. (b)		20,173		1,755,051
Magnolia Oil & Gas Corp. Class A		65,525		1,549,666
Viper Energy Partners LP (c)		41,473		1,226,356
Whiting Petroleum Corp.		13,490		1,099,570
				6,527,167
				8,780,034
Financial — 18.1%
Banks — 5.3%
Ameris Bancorp		46,638		2,046,475
Atlantic Union Bankshares Corp.		26,519		972,982
Cadence Bank		32,045		937,637
First Interstate BancSystem, Inc. Class A		24,036		883,804
National Bank Holdings Corp. Class A		22,852		920,478
Seacoast Banking Corp. of Florida		32,283		1,130,551
Simmons First National Corp. Class A		31,646		829,758
Synovus Financial Corp.		19,142		937,958
Western Alliance Bancorp		9,780		809,980
				9,469,623
Diversified Financial Services — 2.6%
Air Lease Corp.		26,954		1,203,496
Hannon Armstrong Sustainable Infrastructure Capital, Inc. (b)		37,549		1,780,949
PRA Group, Inc. (a)		26,099		1,176,543
StepStone Group, Inc. Class A		16,710		552,433
				4,713,421
Insurance — 2.5%
Assured Guaranty Ltd.		22,604		1,438,971
Kemper Corp.		14,258		806,147
MGIC Investment Corp.		33,935		459,819
Selective Insurance Group, Inc.		12,612		1,127,009
SiriusPoint Ltd. (a)		76,696		573,686
				4,405,632
Real Estate — 0.6%
McGrath RentCorp		13,357		1,135,078
Real Estate Investment Trusts (REITS) — 7.1%
Centerspace		9,888		970,211
Essential Properties Realty Trust, Inc.		68,908		1,743,372
Hudson Pacific Properties, Inc.		37,368		1,036,962
Independence Realty Trust, Inc.		41,217		1,089,777

MML Small Cap Growth Equity Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
Life Storage, Inc.		6,493	$	911,812
Pebblebrook Hotel Trust		39,497		966,887
Phillips Edison & Co., Inc.		50,496		1,736,557
PotlatchDeltic Corp.		18,187		959,001
PS Business Parks, Inc.		7,004		1,177,232
Ryman Hospitality Properties, Inc. (a)		11,016		1,021,954
Xenia Hotels & Resorts, Inc. (a)		51,065		985,044
				12,598,809
				32,322,563
Industrial — 18.5%
Aerospace & Defense — 2.1%
Curtiss-Wright Corp.		8,752		1,314,200
Mercury Systems, Inc. (a)		19,515		1,257,742
Spirit AeroSystems Holdings, Inc. Class A		25,489		1,246,157
				3,818,099
Building Materials — 0.5%
Louisiana-Pacific Corp.		12,911		802,031
Electrical Components & Equipment — 1.1%
Acuity Brands, Inc.		5,854		1,108,162
Novanta, Inc. (a)		5,943		845,630
				1,953,792
Electronics — 0.9%
II-VI, Inc. (a) (b)		22,550		1,634,649
Engineering & Construction — 1.6%
Fluor Corp. (a) (b)		97,262		2,790,447
Environmental Controls — 0.7%
Clean Harbors, Inc. (a)		11,439		1,277,050
Hand & Machine Tools — 1.3%
Enerpac Tool Group Corp.		37,041		810,827
Kennametal, Inc.		29,637		847,915
Regal Rexnord Corp.		5,022		747,173
				2,405,915
Machinery – Diversified — 6.9%
Albany International Corp. Class A		11,380		959,562
Applied Industrial Technologies, Inc.		21,019		2,157,810
Cactus, Inc. Class A		17,307		981,999
Chart Industries, Inc. (a) (b)		7,190		1,235,026
Colfax Corp. (a)		31,429		1,250,560
Flowserve Corp.		26,464		950,058
Kornit Digital Ltd. (a)		11,677		965,571
The Middleby Corp. (a)		8,205		1,345,128
SPX FLOW, Inc.		9,910		854,440
Zurn Water Solutions Corp.		47,724		1,689,430
				12,389,584

MML Small Cap Growth Equity Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
Metal Fabricate & Hardware — 1.0%
Helios Technologies, Inc.		22,331	$	1,792,063
Miscellaneous - Manufacturing — 0.4%
John Bean Technologies Corp.		5,649		669,237
Packaging & Containers — 0.7%
Graphic Packaging Holding Co.		57,914		1,160,596
Transportation — 0.6%
Kirby Corp. (a)		14,062		1,015,136
Trucking & Leasing — 0.7%
GATX Corp.		10,648		1,313,218
				33,021,817
Technology — 12.8%
Computers — 2.4%
Rapid7, Inc. (a)		25,624		2,850,414
Varonis Systems, Inc. (a)		29,110		1,383,889
				4,234,303
Semiconductors — 3.7%
Ambarella, Inc. (a)		4,697		492,809
Cirrus Logic, Inc. (a)		3,800		322,202
FormFactor, Inc. (a)		25,320		1,064,200
MKS Instruments, Inc.		4,691		703,650
Power Integrations, Inc.		10,617		983,983
Synaptics, Inc. (a)		10,646		2,123,877
Tower Semiconductor Ltd. (a) (b)		21,309		1,031,356
				6,722,077
Software — 6.7%
Cardlytics, Inc. (a) (b)		17,814		979,414
Concentrix Corp.		6,322		1,052,992
Digital Turbine, Inc. (a)		24,529		1,074,616
Five9, Inc. (a)		10,206		1,126,742
Health Catalyst, Inc. (a)		18,284		477,761
Jamf Holding Corp. (a) (b)		71,366		2,484,250
Manhattan Associates, Inc. (a)		5,360		743,486
New Relic, Inc. (a)		22,037		1,473,835
Olo, Inc. Class A (a)		9,783		129,625
Verra Mobility Corp. (a)		85,960		1,399,429
Ziff Davis, Inc. (a)		10,216		988,704
				11,930,854
				22,887,234
Utilities — 1.1%
Electric — 0.7%
Portland General Electric Co.		21,290		1,174,144
Gas — 0.4%
ONE Gas, Inc.		9,100		802,984

MML Small Cap Growth Equity Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
			$	1,977,128
TOTAL COMMON STOCK (Cost $152,730,204)				169,931,087
TOTAL EQUITIES (Cost $152,730,204)				169,931,087
MUTUAL FUNDS — 2.5%
Diversified Financial Services — 2.5%
iShares Russell 2000 Growth ETF		8,691		2,222,723
State Street Navigator Securities Lending Government Money Market Portfolio (d)		2,243,726		2,243,726
				4,466,449
TOTAL MUTUAL FUNDS (Cost $4,354,190)				4,466,449
TOTAL LONG-TERM INVESTMENTS (Cost $157,084,394)				174,397,536
	Principal Amount
SHORT-TERM INVESTMENTS — 2.8%
Repurchase Agreement — 2.8%
Fixed Income Clearing Corp., Repurchase Agreement, dated 3/31/22, 0.000%, due 4/01/22 (e)	$	5,058,325		5,058,325
TOTAL SHORT-TERM INVESTMENTS (Cost $5,058,325)				5,058,325
TOTAL INVESTMENTS — 100.5% (Cost $162,142,719) (f)				179,455,861
Other Assets/(Liabilities) — (0.5)%				(979,565)
NET ASSETS — 100.0%			$	178,476,296

Abbreviation Legend

ADR	American Depositary Receipt
ETF	Exchange-Traded Fund

Notes to Portfolio of Investments

Percentages are stated as a percent of net assets.

(a) Non-income producing security.

(b) Denotes all or a portion of security on loan. The total value of securities on loan as of March 31, 2022, was $13,106,873 or 7.34% of net assets. Total securities on loan may be less than the amounts identified in the Portfolio of Investments. The Fund received $11,221,198 of non-cash collateral (U.S. Treasury and/or Agency obligations) related to securities lending activity. This amount is not reflected on the Portfolio of Investments. (Note 2).

(c) Security is a Master Limited Partnership.

(d) Represents investment of security lending cash collateral. (Note 2).

(e) Maturity value of $5,058,325. Collateralized by U.S. Government Agency obligations with a rate of 0.500%, maturity date of 4/15/24, and an aggregate market value, including accrued interest, of $5,159,607.

(f) See Note 3 for aggregate cost for federal tax purposes.

MML Small Company Value Fund —  Portfolio of Investments

March 31, 2022 (Unaudited)

	Number of Shares		Value
EQUITIES — 99.3%
COMMON STOCK — 99.3%
Basic Materials — 1.0%
Chemicals — 1.0%
Minerals Technologies, Inc.		14,114	$	933,641
Communications — 1.3%
Advertising — 0.8%
Entravision Communications Corp. Class A		115,620		741,124
Internet — 0.3%
Solo Brands, Inc. Class A (a) (b)		26,760		228,263
Media — 0.2%
Townsquare Media, Inc. Class A (a)		12,543		160,425
				1,129,812
Consumer, Cyclical — 17.9%
Apparel — 2.0%
Tapestry, Inc.		48,313		1,794,828
Distribution & Wholesale — 1.3%
A-Mark Precious Metals, Inc.		6,509		503,406
KAR Auction Services, Inc. (a)		40,262		726,729
				1,230,135
Entertainment — 1.1%
Accel Entertainment, Inc. (a)		57,023		694,540
Penn National Gaming, Inc. (a)		8,457		358,746
				1,053,286
Food Services — 0.5%
Healthcare Services Group, Inc.		23,068		428,373
Home Builders — 1.4%
Skyline Champion Corp. (a)		22,821		1,252,416
Leisure Time — 3.4%
Brunswick Corp.		19,072		1,542,734
Malibu Boats, Inc. Class A (a)		15,258		885,117
OneWater Marine, Inc. Class A		20,731		714,183
				3,142,034
Lodging — 0.4%
Boyd Gaming Corp.		5,401		355,278

MML Small Company Value Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
Retail — 7.3%
Beacon Roofing Supply, Inc. (a)		24,057	$	1,426,099
Dave & Buster's Entertainment, Inc. (a)		21,493		1,055,307
GMS, Inc. (a)		18,590		925,224
MarineMax, Inc. (a)		23,178		933,146
Penske Automotive Group, Inc.		20,508		1,922,010
Red Robin Gourmet Burgers, Inc. (a)		25,507		430,048
				6,691,834
Textiles — 0.5%
UniFirst Corp.		2,298		423,475
				16,371,659
Consumer, Non-cyclical — 15.8%
Commercial Services — 9.2%
AMN Healthcare Services, Inc. (a)		5,615		585,813
Barrett Business Services, Inc.		6,103		472,800
The Brink's Co.		28,095		1,910,460
Deluxe Corp.		34,068		1,030,216
Euronet Worldwide, Inc. (a)		3,158		411,014
EVERTEC, Inc.		43,966		1,799,528
Korn Ferry		22,211		1,442,382
Loomis AB		26,397		721,746
				8,373,959
Cosmetics & Personal Care — 1.5%
Edgewell Personal Care Co.		38,595		1,415,279
Health Care – Products — 2.0%
Patterson Cos., Inc.		36,866		1,193,352
Varex Imaging Corp. (a)		21,217		451,710
Zimvie, Inc.		8,405		191,970
				1,837,032
Health Care – Services — 0.4%
National HealthCare Corp.		4,934		346,515
Household Products & Wares — 2.7%
Spectrum Brands Holdings, Inc.		27,555		2,444,679
				14,417,464
Energy — 2.2%
Energy – Alternate Sources — 0.8%
Enviva, Inc.		9,077		718,444
Oil & Gas — 0.2%
Earthstone Energy, Inc. Class A (a) (b)		13,152		166,110
Oil & Gas Services — 1.2%
ChampionX Corp. (a)		45,954		1,124,954

MML Small Company Value Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
NCS Multistage Holdings, Inc. (a)		479	$	23,979
				1,148,933
				2,033,487
Financial — 32.2%
Banks — 18.9%
Ameris Bancorp		24,173		1,060,711
BankUnited, Inc.		32,764		1,440,305
ConnectOne Bancorp, Inc.		13,278		425,029
CVB Financial Corp.		9,989		231,845
F.N.B. Corp.		119,177		1,483,754
First BanCorp		96,491		1,265,962
First Hawaiian, Inc.		2,000		55,780
First Mid Bancshares, Inc.		4,750		182,828
Home BancShares, Inc.		48,171		1,088,665
Independent Bank Corp.		7,709		629,748
Independent Bank Group, Inc.		12,165		865,661
Old National Bancorp		88,693		1,452,791
Origin Bancorp, Inc.		9,679		409,325
Premier Financial Corp.		11,149		338,149
QCR Holdings, Inc.		4,537		256,749
SouthState Corp.		18,730		1,528,181
Towne Bank		13,495		404,040
UMB Financial Corp.		14,442		1,403,185
Valley National Bancorp		141,169		1,838,020
Veritex Holdings, Inc.		24,493		934,898
				17,295,626
Diversified Financial Services — 0.4%
Enact Holdings, Inc.		16,098		358,180
Insurance — 2.9%
Axis Capital Holdings Ltd.		30,565		1,848,265
James River Group Holdings Ltd.		8,478		209,746
ProAssurance Corp.		23,986		644,744
				2,702,755
Investment Companies — 1.9%
Compass Diversified Holdings (b)		73,522		1,747,618
Private Equity — 0.3%
Patria Investments Ltd.		14,611		260,222
Real Estate Investment Trusts (REITS) — 6.1%
Brandywine Realty Trust		61,986		876,482
CareTrust REIT, Inc.		30,106		581,046
Cousins Properties, Inc.		9,967		401,570
Easterly Government Properties, Inc.		18,296		386,777
Four Corners Property Trust, Inc.		27,797		751,631
Getty Realty Corp.		13,614		389,633
Healthcare Realty Trust, Inc.		5,162		141,852
Highwoods Properties, Inc.		8,629		394,690
Kite Realty Group Trust		16,692		380,077
National Health Investors, Inc. (b)		8,711		514,036
Physicians Realty Trust		14,530		254,856

MML Small Company Value Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
Sabra Health Care REIT, Inc.		16,138	$	240,295
Summit Hotel Properties, Inc. (a)		23,537		234,429
				5,547,374
Savings & Loans — 1.7%
Pacific Premier Bancorp, Inc.		32,382		1,144,704
Provident Financial Services, Inc.		17,459		408,540
				1,553,244
				29,465,019
Industrial — 21.3%
Building Materials — 2.3%
Cornerstone Building Brands, Inc. (a)		30,243		735,510
Mohawk Industries, Inc. (a)		4,627		574,673
Tecnoglass, Inc.		32,699		825,323
				2,135,506
Electrical Components & Equipment — 0.9%
Belden, Inc.		13,962		773,495
Graham Corp.		10,695		82,458
				855,953
Electronics — 6.9%
Advanced Energy Industries, Inc.		3,640		313,331
Avnet, Inc.		38,380		1,557,844
Coherent, Inc. (a)		4,550		1,243,788
II-VI, Inc. (a) (b)		32,433		2,351,068
Vontier Corp.		31,929		810,678
				6,276,709
Engineering & Construction — 1.6%
Arcosa, Inc.		4,762		272,625
Dycom Industries, Inc. (a)		12,150		1,157,409
				1,430,034
Environmental Controls — 0.4%
CECO Environmental Corp. (a)		30,698		168,532
Charah Solutions, Inc. (a) (b)		32,498		162,165
				330,697
Hand & Machine Tools — 0.2%
Luxfer Holdings PLC		10,888		182,918
Machinery – Diversified — 3.5%
Albany International Corp. Class A		859		72,431
Colfax Corp. (a)		39,231		1,561,001
DXP Enterprises, Inc. (a)		17,396		471,258
Gates Industrial Corp. PLC (a)		76,265		1,148,551
				3,253,241

MML Small Company Value Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
Metal Fabricate & Hardware — 2.4%
The Timken Co.		36,092	$	2,190,785
Miscellaneous - Manufacturing — 0.1%
DIRTT Environmental Solutions (a) (b)		64,423		87,615
Packaging & Containers — 3.0%
Graphic Packaging Holding Co.		90,890		1,821,436
Karat Packaging, Inc. (a)		6,249		124,043
Pactiv Evergreen, Inc.		79,608		800,856
				2,746,335
				19,489,793
Technology — 6.5%
Computers — 0.4%
KBR, Inc.		6,754		369,646
Semiconductors — 2.0%
Cohu, Inc. (a)		11,843		350,553
Kulicke & Soffa Industries, Inc. (b)		26,236		1,469,741
				1,820,294
Software — 4.1%
Avaya Holdings Corp. (a)		73,790		934,919
Donnelley Financial Solutions, Inc. (a)		27,120		902,011
IBEX Holdings Ltd. (a) (b)		13,734		218,920
Teradata Corp. (a)		33,240		1,638,400
				3,694,250
				5,884,190
Utilities — 1.1%
Electric — 0.3%
PNM Resources, Inc.		5,932		282,778
Gas — 0.8%
Northwest Natural Holding Co.		3,556		183,917
South Jersey Industries, Inc.		7,944		274,465
Southwest Gas Holdings, Inc.		3,556		278,399
				736,781
				1,019,559
TOTAL COMMON STOCK (Cost $94,256,113)				90,744,624
TOTAL EQUITIES (Cost $94,256,113)				90,744,624
MUTUAL FUNDS — 2.3%
Diversified Financial Services — 2.3%
State Street Navigator Securities Lending Government Money Market Portfolio (c)		2,055,914		2,055,914

MML Small Company Value Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
TOTAL MUTUAL FUNDS (Cost $2,055,914)			$	2,055,914
TOTAL LONG-TERM INVESTMENTS (Cost $96,312,027)				92,800,538
	Principal Amount
SHORT-TERM INVESTMENTS — 0.8%
Repurchase Agreement — 0.8%
Fixed Income Clearing Corp., Repurchase Agreement, dated 3/31/22, 0.000%, due 4/01/22 (d)	$	773,980		773,980
TOTAL SHORT-TERM INVESTMENTS (Cost $773,980)				773,980
TOTAL INVESTMENTS — 102.4% (Cost $97,086,007) (e)				93,574,518
Other Assets/(Liabilities) — (2.4)%				(2,151,175)
NET ASSETS — 100.0%			$	91,423,343

Notes to Portfolio of Investments

Percentages are stated as a percent of net assets.

(a) Non-income producing security.

(b) Denotes all or a portion of security on loan. The total value of securities on loan as of March 31, 2022, was $6,279,766 or 6.87% of net assets. Total securities on loan may be less than the amounts identified in the Portfolio of Investments. The Fund received $4,443,419 of non-cash collateral (U.S. Treasury and/or Agency obligations) related to securities lending activity. This amount is not reflected on the Portfolio of Investments. (Note 2).

(c) Represents investment of security lending cash collateral. (Note 2).

(d) Maturity value of $773,980. Collateralized by U.S. Government Agency obligations with a rate of 0.500%, maturity date of 4/15/24, and an aggregate market value, including accrued interest, of $789,560.

(e) See Note 3 for aggregate cost for federal tax purposes.

Forward contracts
Counterparty	Settlement Date	Currency Purchased		Currency Sold		Unrealized Appreciation (Depreciation)
UBS AG	6/30/22	USD	644,132	SEK	6,073,818	$	(3,333)
UBS AG	6/30/22	SEK	296,174	USD	31,525		47
						$	(3,286)

Currency Legend

SEK Swedish Krona

USD U.S. Dollar

MML Small/Mid Cap Value Fund —  Portfolio of Investments

March 31, 2022 (Unaudited)

	Number of Shares		Value
EQUITIES — 99.9%
COMMON STOCK — 99.9%
Basic Materials — 3.7%
Chemicals — 1.7%
Huntsman Corp.		38,350	$	1,438,508
Innospec, Inc.		11,993		1,109,952
Orion Engineered Carbons SA		9,536		152,290
Trinseo PLC		1,373		65,794
				2,766,544
Iron & Steel — 2.0%
Carpenter Technology Corp.		38,348		1,609,849
Reliance Steel & Aluminum Co.		9,025		1,654,734
				3,264,583
				6,031,127
Communications — 1.0%
Internet — 1.0%
Criteo SA Sponsored ADR (a)		61,771		1,682,642
Consumer, Cyclical — 17.2%
Airlines — 0.8%
SkyWest, Inc. (a)		44,837		1,293,547
Apparel — 4.1%
Carter's, Inc.		24,218		2,227,814
Kontoor Brands, Inc.		25,717		1,063,398
Ralph Lauren Corp.		16,110		1,827,518
Tapestry, Inc.		40,840		1,517,206
				6,635,936
Auto Parts & Equipment — 1.9%
Dana, Inc.		57,103		1,003,300
The Goodyear Tire & Rubber Co. (a)		145,570		2,080,195
				3,083,495
Entertainment — 0.8%
Scientific Games Corp. Class A (a)		21,270		1,249,613
Home Builders — 2.8%
KB Home		44,130		1,428,929
PulteGroup, Inc.		48,990		2,052,681
Taylor Morrison Home Corp. (a)		41,534		1,130,556
				4,612,166

MML Small/Mid Cap Value Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
Home Furnishing — 1.0%
MillerKnoll, Inc.		49,590	$	1,713,830
Lodging — 1.3%
Hilton Grand Vacations, Inc. (a)		39,910		2,075,719
Retail — 4.5%
Dine Brands Global, Inc.		26,441		2,061,076
Papa John's International, Inc.		17,911		1,885,670
Sally Beauty Holdings, Inc. (a)		104,090		1,626,927
Williams-Sonoma, Inc.		12,841		1,861,945
				7,435,618
				28,099,924
Consumer, Non-cyclical — 14.9%
Commercial Services — 5.8%
ADT, Inc.		220,730		1,675,341
Herc Holdings, Inc.		14,510		2,424,476
Korn Ferry		19,562		1,270,356
Robert Half International, Inc.		18,032		2,058,894
WillScot Mobile Mini Holdings Corp. (a)		52,540		2,055,890
				9,484,957
Food — 2.4%
The Hain Celestial Group, Inc. (a)		59,775		2,056,260
Nomad Foods Ltd. (a)		84,857		1,916,071
				3,972,331
Health Care – Products — 2.3%
Envista Holdings Corp. (a)		38,340		1,867,541
Integra LifeSciences Holdings Corp. (a)		29,160		1,873,822
				3,741,363
Health Care – Services — 4.4%
Acadia Healthcare Co., Inc. (a)		37,040		2,427,231
MEDNAX, Inc. (a)		95,610		2,244,923
Syneos Health, Inc. (a)		30,360		2,457,642
				7,129,796
				24,328,447
Energy — 3.0%
Oil & Gas — 3.0%
Coterra Energy, Inc.		89,661		2,418,157
HF Sinclair Corp.		60,285		2,402,357
				4,820,514
				4,820,514
Financial — 27.1%
Banks — 12.8%
BankUnited, Inc.		43,639		1,918,370

MML Small/Mid Cap Value Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
Comerica, Inc.		25,120	$	2,271,602
First BanCorp		52,719		691,673
First Citizens BancShares, Inc. Class A		3,539		2,355,558
First Hawaiian, Inc.		72,120		2,011,427
Synovus Financial Corp.		44,414		2,176,286
Texas Capital Bancshares, Inc. (a)		33,268		1,906,589
Umpqua Holdings Corp.		73,238		1,381,269
Webster Financial Corp.		37,379		2,097,709
Wintrust Financial Corp.		20,090		1,866,964
Zions Bancorp NA		33,578		2,201,374
				20,878,821
Diversified Financial Services — 2.1%
Moelis & Co. Class A		36,806		1,728,042
Stifel Financial Corp.		24,850		1,687,315
				3,415,357
Insurance — 4.2%
American Financial Group, Inc.		12,668		1,844,714
The Hanover Insurance Group, Inc.		13,598		2,033,173
Kemper Corp.		25,500		1,441,770
Selective Insurance Group, Inc.		17,507		1,564,426
				6,884,083
Real Estate Investment Trusts (REITS) — 8.0%
American Campus Communities, Inc.		31,732		1,776,040
Broadstone Net Lease, Inc.		71,390		1,554,874
Camden Property Trust		15,758		2,618,980
Cousins Properties, Inc.		46,070		1,856,160
CubeSmart		28,314		1,473,177
Physicians Realty Trust		124,753		2,188,168
STAG Industrial, Inc.		40,992		1,695,019
				13,162,418
				44,340,679
Industrial — 22.8%
Aerospace & Defense — 1.3%
Spirit AeroSystems Holdings, Inc. Class A		44,670		2,183,916
Building Materials — 0.8%
Masonite International Corp. (a)		14,691		1,306,618
Electrical Components & Equipment — 0.8%
Belden, Inc.		24,734		1,370,264
Electronics — 2.2%
Avnet, Inc.		47,576		1,931,110
Sensata Technologies Holding PLC (a)		34,290		1,743,646
				3,674,756
Engineering & Construction — 4.0%
AECOM		37,900		2,911,099
Arcosa, Inc.		22,255		1,274,099

MML Small/Mid Cap Value Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
Dycom Industries, Inc. (a)		23,870	$	2,273,856
				6,459,054
Hand & Machine Tools — 1.6%
Regal Rexnord Corp.		17,230		2,563,479
Machinery – Construction & Mining — 2.1%
Oshkosh Corp.		16,791		1,690,014
Vertiv Holdings Co.		126,890		1,776,460
				3,466,474
Machinery – Diversified — 2.2%
Applied Industrial Technologies, Inc.		14,500		1,488,570
Cactus, Inc. Class A		37,770		2,143,070
				3,631,640
Metal Fabricate & Hardware — 0.9%
The Timken Co.		23,740		1,441,018
Packaging & Containers — 2.3%
Berry Global Group, Inc. (a)		33,670		1,951,514
Sealed Air Corp.		26,020		1,742,299
				3,693,813
Transportation — 3.8%
Kirby Corp. (a)		31,060		2,242,221
Knight-Swift Transportation Holdings, Inc.		34,863		1,759,187
XPO Logistics, Inc. (a)		29,240		2,128,672
				6,130,080
Trucking & Leasing — 0.8%
GATX Corp.		10,460		1,290,032
				37,211,144
Technology — 7.7%
Computers — 1.9%
Genpact Ltd.		23,000		1,000,730
Lumentum Holdings, Inc. (a)		21,590		2,107,184
				3,107,914
Semiconductors — 2.1%
Kulicke & Soffa Industries, Inc. (b)		18,722		1,048,807
ON Semiconductor Corp. (a)		38,940		2,438,033
				3,486,840
Software — 3.7%
ACI Worldwide, Inc. (a)		58,370		1,838,071
Change Healthcare, Inc. (a)		105,500		2,299,900

MML Small/Mid Cap Value Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
CommVault Systems, Inc. (a)		27,696	$	1,837,630
				5,975,601
				12,570,355
Utilities — 2.5%
Electric — 1.6%
IDACORP, Inc.		23,001		2,653,396
Gas — 0.9%
Southwest Gas Holdings, Inc.		18,780		1,470,286
				4,123,682
TOTAL COMMON STOCK (Cost $138,457,254)				163,208,514
TOTAL EQUITIES (Cost $138,457,254)				163,208,514
TOTAL LONG-TERM INVESTMENTS (Cost $138,457,254)				163,208,514
	Principal Amount
SHORT-TERM INVESTMENTS — 0.5%
Repurchase Agreement — 0.5%
Fixed Income Clearing Corp., Repurchase Agreement, dated 3/31/22, 0.000%, due 4/01/22 (c)	$	849,870		849,870
TOTAL SHORT-TERM INVESTMENTS (Cost $849,870)				849,870
TOTAL INVESTMENTS — 100.4% (Cost $139,307,124) (d)				164,058,384
Other Assets/(Liabilities) — (0.4)%				(609,155)
NET ASSETS — 100.0%			$	163,449,229

Abbreviation Legend

ADR American Depositary Receipt

Notes to Portfolio of Investments

Percentages are stated as a percent of net assets.

(a) Non-income producing security.

(b) Denotes all or a portion of security on loan. The total value of securities on loan as of March 31, 2022, was $1,047,356 or 0.64% of net assets. Total securities on loan may be less than the amounts identified in the Portfolio of Investments. The Fund received $1,096,351 of non-cash collateral (U.S. Treasury and/or Agency obligations) related to securities lending activity. This amount is not reflected on the Portfolio of Investments. (Note 2).

(c) Maturity value of $849,870. Collateralized by U.S. Government Agency obligations with a rate of 0.500%, maturity date of 4/15/24, and an aggregate market value, including accrued interest, of $866,952.

(d) See Note 3 for aggregate cost for federal tax purposes.

MML Total Return Bond Fund —  Portfolio of Investments

March 31, 2022 (Unaudited)

	Number of Shares		Value
EQUITIES — 0.1%
COMMON STOCK — 0.1%
Communications — 0.1%
Telecommunications — 0.1%
Intelsat Emergence SA		7,210	$	209,991
TOTAL COMMON STOCK (Cost $676,994)				209,991
TOTAL EQUITIES (Cost $676,994)				209,991
	Principal Amount
BONDS & NOTES — 115.9%
BANK LOANS — 2.0%
Aerospace & Defense — 0.0%
TransDigm, Inc., 2020 Term Loan E, 1 mo. USD LIBOR + 2.250%
2.707% VRN 5/30/25	$	48,875		47,943
Airlines — 0.0%
American Airlines, Inc., 2017 1st Lien Term Loan, 1 mo. USD LIBOR + 1.750%
2.197% VRN 1/29/27		34,152		31,861
Auto Parts & Equipment — 0.1%
Clarios Global LP, 2021 USD Term Loan B, 1 mo. USD LIBOR + 3.250%
3.707% VRN 4/30/26		194,908		192,228
Beverages — 0.0%
Naked Juice LLC
Term Loan, SOFR + 3.250%
3.315% VRN 1/24/29		134,337		131,938
Delayed Draw Term Loan, SOFR + 3.250%
3.315% VRN 1/24/29		7,750		7,612
				139,550
Chemicals — 0.0%
Zep, Inc., 2017 1st Lien Term Loan, 3 mo. USD LIBOR + 4.000%
5.000% VRN 8/12/24		91,842		87,250
Commercial Services — 0.0%
Spin Holdco, Inc., 2021 Term Loan, 3 mo. USD LIBOR + 4.000%
4.750% VRN 3/04/28		99,250		98,368

MML Total Return Bond Fund —  Portfolio of Investments (Continued)

	Principal Amount		Value
Diversified Financial Services — 0.1%
Setanta Aircraft Leasing Designated Activity Company, Term Loan B, 3 mo. USD LIBOR + 2.000%
3.006% VRN 11/05/28	$	375,000	$	369,064
Environmental Controls — 0.0%
GFL Environmental, Inc., 2020 Term Loan, 3 mo. USD LIBOR + 3.000%
3.500% VRN 5/30/25		67,528		67,190
Food — 0.0%
Hostess Brands LLC, 2019 Term Loan, 3 mo. USD LIBOR + 2.250%
3.000% VRN 8/03/25		127,394		125,596
Health Care – Products — 0.2%
Avantor Funding, Inc., 2021 Term Loan B5, 1 mo. USD LIBOR + 2.250%
2.750% VRN 11/08/27		247,506		245,251
Medline Borrower LP, USD Term Loan B, 1 mo. USD LIBOR + 3.250%
3.750% - —% VRN 10/23/28 (a)		250,000		247,375
				492,626
Insurance — 0.2%
Acrisure LLC, 2020 Term Loan B, 1 mo. USD LIBOR + 3.500%
3.957% VRN 2/15/27		99,746		98,166
AmWINS Group, Inc., 2021 Term Loan B, 1 mo. USD LIBOR + 2.250%
3.000% VRN 2/19/28		494,199		485,551
				583,717
Internet — 0.1%
NortonLifeLock, Inc., 2022 Term Loan B,
0.000% - —% 1/28/29 (a)		250,000		246,668
Lodging — 0.0%
Caesars Resort Collection LLC, 2017 1st Lien Term Loan B, 1 mo. USD LIBOR + 2.750%
3.207% VRN 12/23/24		456		454
Media — 0.2%
Diamond Sports Group LLC
Term Loan, SOFR + 3.250%
3.545% VRN 8/24/26		64,505		21,811
Term Loan, SOFR + 8.000%
9.000% VRN 5/26/26		105,063		106,391
DirecTV Financing LLC, Term Loan, 3 mo. USD LIBOR + 5.000%
5.750% VRN 8/02/27		162,350		161,991
Sinclair Television Group, Inc., 2021 Term Loan B3, 1 mo. USD LIBOR + 3.000%
3.460% VRN 4/01/28		148,875		145,061
Virgin Media Bristol LLC, USD Term Loan N, 3 mo. USD LIBOR + 2.500%
2.897% VRN 1/31/28		275,000		271,263
				706,517
Packaging & Containers — 0.1%
Berry Global, Inc., 2021 Term Loan Z, 1 mo. USD LIBOR + 1.750%
2.071% VRN 7/01/26		151,015		148,678

MML Total Return Bond Fund —  Portfolio of Investments (Continued)

	Principal Amount		Value
Pharmaceuticals — 0.6%
Change Healthcare Holdings LLC, 2017 Term Loan B, 1 mo. USD LIBOR + 2.500%
3.500% VRN 3/01/24	$	43,501	$	43,207
Elanco Animal Health, Inc., Term Loan B, 1 mo. USD LIBOR + 1.750%
1.981% VRN 8/01/27		402,486		395,567
Endo Luxembourg Finance Company I SARL, 2021 Term Loan, 1 mo. USD LIBOR + 5.000%
5.750% VRN 3/27/28		480,159		448,699
Gainwell Acquisition Corp., Term Loan B, 3 mo. USD LIBOR + 4.000%
5.006% VRN 10/01/27		99,747		99,498
Grifols Worldwide Operations USA, Inc., USD 2019 Term Loan B, 1 mo. USD LIBOR + 2.000%
2.457% VRN 11/15/27		280,004		274,964
Horizon Therapeutics USA, Inc., 2021 Term Loan B2, 1 mo. USD LIBOR + 1.750%
2.250% VRN 3/15/28		372,188		368,001
ICON Luxembourg S.A.R.L.
US Term Loan, 3 mo. USD LIBOR + 2.250%
3.313% VRN 7/03/28		30,749		30,576
LUX Term Loan, 3 mo. USD LIBOR + 2.250%
3.313% VRN 7/03/28		123,414		122,720
Jazz Financing Lux S.A.R.L., USD Term Loan, 1 mo. USD LIBOR + 3.500%
4.000% VRN 5/05/28		148,875		148,187
Organon & Co, USD Term Loan, 3 mo. USD LIBOR + 3.000%
3.563% VRN 6/02/28		265,146		263,489
				2,194,908
Retail — 0.1%
Michaels Cos., Inc., 2021 Term Loan B, 3 mo. USD LIBOR + 4.250%
5.256% VRN 4/15/28		238,200		222,948
Whatabrands LLC, 2021 Term Loan B, 1 mo. USD LIBOR + 3.250%
3.750% - —% VRN 8/03/28 (a)		225,000		222,048
				444,996
Software — 0.1%
Athenahealth, Inc.
2022 Term Loan B, 1 mo. SOFR + 3.500%
0.000%VRN2/15/29 (a)		213,768		211,149
2022 Delayed Draw Term Loan, 3 mo. SOFR + 0.000%
0.000%VRN2/15/29 (a) (b)		36,232		35,788
				246,937
Telecommunications — 0.2%
Level 3 Financing, Inc., 2019 Term Loan B, 1 mo. USD LIBOR + 1.750%
2.207% VRN 3/01/27		600,000		588,186
TOTAL BANK LOANS (Cost $6,890,723)				6,812,737
CORPORATE DEBT — 29.6%
Aerospace & Defense — 0.2%
The Boeing Co.
1.167% 2/04/23		660,000		654,789

MML Total Return Bond Fund —  Portfolio of Investments (Continued)

	Principal Amount		Value
Agriculture — 0.8%
BAT Capital Corp.
4.390% 8/15/37	$	400,000	$	372,041
4.540% 8/15/47		1,050,000		936,573
5.650% 3/16/52		215,000		218,011
Imperial Brands Finance PLC
3.125% 7/26/24 (c)		320,000		317,288
3.500% 7/26/26 (c)		225,000		220,363
Imperial Tobacco Finance PLC
4.250% 7/21/25 (c)		415,000		418,709
Reynolds American, Inc.
5.850% 8/15/45		315,000		322,419
				2,805,404
Airlines — 0.0%
Northwest Airlines Pass-Through Trust
7.041% 10/01/23		12,371		12,370
Auto Manufacturers — 0.2%
General Motors Financial Co., Inc.
3.150% 6/30/22		550,000		551,362
3.450% 4/10/22		105,000		105,036
				656,398
Banks — 9.7%
Bank of America Corp.
SOFR + .910% 1.658% VRN 3/11/27		1,305,000		1,217,981
SOFR + 1.060% 2.087% VRN 6/14/29		1,000,000		915,155
SOFR + 1.320% 2.687% VRN 4/22/32		120,000		110,372
SOFR + 1.330% 2.972% VRN 2/04/33		670,000		628,691
3 mo. USD LIBOR + .790% 3.004% VRN 12/20/23		1,180,000		1,183,295
3 mo. USD LIBOR + .970% 3.458% VRN 3/15/25		415,000		417,801
3 mo. USD LIBOR + 1.070% 3.970% VRN 3/05/29		485,000		493,226
Citigroup, Inc.
SOFR + .528% 1.281% VRN 11/03/25		420,000		399,835
SOFR + .770% 1.462% VRN 6/09/27		725,000		666,702
SOFR + 1.167% 2.561% VRN 5/01/32		200,000		180,380
SOFR + 1.351% 3.057% VRN 1/25/33		505,000		473,293
SOFR + 1.280% 3.070% VRN 2/24/28		190,000		185,474
SOFR + 2.842% 3.106% VRN 4/08/26		335,000		332,523
SOFR + 1.939% 3.785% VRN 3/17/33		490,000		487,410
Credit Suisse Group AG
SOFR + .980% 1.305% VRN 2/02/27 (c)		495,000		442,726
SOFR + 2.044% 2.193% VRN 6/05/26 (c)		590,000		553,955
SOFR + 1.560% 2.593% VRN 9/11/25 (c)		420,000		406,084
SOFR + 1.730% 3.091% VRN 5/14/32 (c)		405,000		365,320
4.282% 1/09/28 (c)		65,000		65,017
DNB Bank ASA 1 year CMT + .680%
1.605% VRN 3/30/28 (c)		635,000		574,353
The Goldman Sachs Group, Inc.
SOFR + .486% 0.925% VRN 10/21/24		415,000		402,493
1.217% 12/06/23		1,030,000		1,005,730
SOFR + .798% 1.431% VRN 3/09/27		500,000		461,850
SOFR + .818% 1.542% VRN 9/10/27		1,450,000		1,327,028
SOFR + .913% 1.948% VRN 10/21/27		125,000		116,332
SOFR + 1.410% 3.102% VRN 2/24/33		705,000		664,603
3.200% 2/23/23		145,000		146,319

MML Total Return Bond Fund —  Portfolio of Investments (Continued)

	Principal Amount		Value
3 mo. USD LIBOR + 1.201% 3.272% VRN 9/29/25	$	325,000	$	325,372
3.500% 4/01/25		115,000		115,982
HSBC Holdings PLC
SOFR + .708% 0.976% VRN 5/24/25		505,000		479,798
SOFR + 1.732% 2.013% VRN 9/22/28		1,080,000		980,912
SOFR + 1.285% 2.206% VRN 8/17/29		610,000		549,723
SOFR + 1.187% 2.804% VRN 5/24/32		365,000		332,565
3 mo. USD LIBOR + 1.211% 3.803% VRN 3/11/25		110,000		110,596
JP Morgan Chase & Co.
SOFR + .580% 0.697% VRN 3/16/24		165,000		161,908
SOFR + .490% 0.768% VRN 8/09/25		255,000		241,918
3 mo. TSFR + .580% 0.969% VRN 6/23/25		1,060,000		1,012,884
SOFR + .885% 1.578% VRN 4/22/27		500,000		466,279
SOFR + 1.015% 2.069% VRN 6/01/29		250,000		230,819
SOFR + 1.180% 2.545% VRN 11/08/32		200,000		182,777
SOFR + 1.250% 2.580% VRN 4/22/32		180,000		164,931
SOFR + 1.170% 2.947% VRN 2/24/28 (d)		250,000		244,329
2.950% 10/01/26		895,000		889,284
SOFR + 1.260% 2.963% VRN 1/25/33		390,000		367,730
Lloyds Banking Group PLC
1 year CMT + .850% 1.627% VRN 5/11/27 (d)		515,000		472,834
3 mo. USD LIBOR + 1.205% 3.574% VRN 11/07/28		520,000		508,771
1 year CMT + 1.800% 3.750% VRN 3/18/28		200,000		199,707
1 year CMT + 3.500% 3.870% VRN 7/09/25		220,000		222,499
Macquarie Group Ltd.
SOFR + 1.532% 2.871% VRN 1/14/33 (c)		1,045,000		927,648
4.442% 6/21/33 (c)		240,000		242,664
Morgan Stanley
SOFR + .509% 0.791% VRN 1/22/25		285,000		273,775
SOFR + .560% 1.164% VRN 10/21/25		1,300,000		1,236,087
SOFR + .879% 1.593% VRN 5/04/27		505,000		469,785
SOFR + 1.178% 2.239% VRN 7/21/32		160,000		142,231
SOFR + 1.200% 2.511% VRN 10/20/32		285,000		256,903
SOFR + 1.290% 2.943% VRN 1/21/33		450,000		422,047
Natwest Group PLC 3 mo. USD LIBOR + 1.762%
4.269% VRN 3/22/25		775,000		784,496
Santander UK Group Holdings PLC
SOFR + .787% 1.089% VRN 3/15/25		540,000		514,000
SOFR + .989% 1.673% VRN 6/14/27		120,000		109,321
SOFR + 1.220% 2.469% VRN 1/11/28		200,000		186,281
3 mo. USD LIBOR + 1.080% 3.373% VRN 1/05/24		445,000		446,268
3 mo. USD LIBOR + 1.400% 3.823% VRN 11/03/28		485,000		477,683
Santander UK PLC
5.000% 11/07/23 (c)		435,000		444,940
UBS AG
0.700% 8/09/24 (c)		350,000		332,141
Wells Fargo & Co.
3 mo. USD LIBOR + .750% 2.164% VRN 2/11/26		335,000		324,382
SOFR + 2.100% 2.393% VRN 6/02/28		1,400,000		1,325,918
SOFR + 1.432% 2.879% VRN 10/30/30		490,000		465,772
SOFR + 1.500% 3.350% VRN 3/02/33		1,285,000		1,249,351
SOFR + 1.510% 3.526% VRN 3/24/28		320,000		319,125
SOFR + 4.502% 5.013% VRN 4/04/51		150,000		177,923
				33,614,307

MML Total Return Bond Fund —  Portfolio of Investments (Continued)

	Principal Amount		Value
Beverages — 0.5%
Anheuser-Busch Cos. LLC/Anheuser-Busch InBev Worldwide, Inc.
4.900% 2/01/46	$	880,000	$	978,233
Bacardi Ltd.
4.450% 5/15/25 (c)		700,000		717,209
				1,695,442
Biotechnology — 0.2%
Amgen, Inc.
3.350% 2/22/32		230,000		227,882
4.200% 2/22/52		135,000		139,540
4.400% 2/22/62		190,000		197,131
Grifols Escrow Issuer SA
4.750% 10/15/28 (c)		125,000		117,656
				682,209
Chemicals — 0.1%
International Flavors & Fragrances, Inc.
5.000% 9/26/48		380,000		416,583
Commercial Services — 0.1%
S&P Global, Inc.
4.750% 8/01/28 (c) (d)		385,000		414,340
Diversified Financial Services — 1.2%
AerCap Ireland Capital DAC / AerCap Global Aviation Trust
3.500% 1/15/25 (d)		1,000,000		981,016
AerCap Ireland Capital DAC/AerCap Global Aviation Trust
3.300% 1/30/32		615,000		554,480
American Express Co.
2.550% 3/04/27		385,000		375,259
Avolon Holdings Funding Ltd.
2.528% 11/18/27 (c)		389,000		344,201
2.875% 2/15/25 (c)		335,000		322,001
3.950% 7/01/24 (c)		130,000		129,686
Capital One Financial Corp. SOFR + 1.790%
3.273% VRN 3/01/30		535,000		516,969
Charles Schwab Corp.
2.900% 3/03/32		380,000		365,923
Park Aerospace Holdings Ltd.
4.500% 3/15/23 (c)		320,000		322,292
5.500% 2/15/24 (c) (d)		82,000		83,980
				3,995,807
Electric — 1.3%
Duke Energy Carolinas LLC
4.000% 9/30/42		1,000,000		1,023,481
Florida Power & Light Co.
3.990% 3/01/49		495,000		529,092
Jersey Central Power & Light Co.
4.700% 4/01/24 (c)		800,000		818,502
6.400% 5/15/36		425,000		515,823
MidAmerican Energy Co.
4.800% 9/15/43		1,400,000		1,579,982
				4,466,880

MML Total Return Bond Fund —  Portfolio of Investments (Continued)

	Principal Amount		Value
Engineering & Construction — 0.0%
Artera Services LLC
9.033% 12/04/25 (c)	$	95,000	$	93,812
Entertainment — 0.4%
Cdi Escrow Issuer, Inc.
5.750% 4/01/30 (c) (e)		115,000		116,150
Magallanes, Inc.
4.279% 3/15/32 (c)		190,000		190,817
5.050% 3/15/42 (c)		760,000		776,568
5.141% 3/15/52 (c)		380,000		388,871
				1,472,406
Environmental Controls — 0.1%
GFL Environmental, Inc.
5.125% 12/15/26 (c)		164,000		165,394
Food — 1.0%
JBS USA LUX SA / JBS USA Food Co. / JBS USA Finance, Inc.
4.375% 2/02/52 (c)		135,000		117,405
JBS USA LUX SA/JBS USA Finance, Inc.
6.750% 2/15/28 (c)		425,000		447,317
Kraft Heinz Foods Co.
4.875% 10/01/49		480,000		505,953
5.000% 6/04/42		360,000		384,750
Pilgrim's Pride Corp.
3.500% 3/01/32 (c)		360,000		315,000
5.875% 9/30/27 (c)		30,000		30,319
Post Holdings, Inc.
4.500% 9/15/31 (c)		400,000		353,800
4.625% 4/15/30 (c)		800,000		720,360
Smithfield Foods, Inc.
2.625% 9/13/31 (c)		635,000		556,936
				3,431,840
Health Care – Products — 0.2%
PerkinElmer, Inc.
2.250% 9/15/31		800,000		707,196
Health Care – Services — 1.8%
Aetna, Inc.
2.800% 6/15/23		485,000		486,985
Centene Corp.
2.450% 7/15/28		488,000		445,886
3.000% 10/15/30		453,000		417,778
4.250% 12/15/27		165,000		165,619
CommonSpirit Health
2.782% 10/01/30		495,000		461,375
3.910% 10/01/50		75,000		71,415
Fresenius Medical Care US Finance III, Inc.
1.875% 12/01/26 (c)		340,000		314,996
HCA, Inc.
2.375% 7/15/31		90,000		80,274
3.625% 3/15/32 (c)		350,000		342,926
4.125% 6/15/29		345,000		351,698
4.625% 3/15/52 (c)		195,000		196,577
5.000% 3/15/24		287,000		297,743

MML Total Return Bond Fund —  Portfolio of Investments (Continued)

	Principal Amount		Value
5.250% 6/15/49	$	775,000	$	848,342
5.500% 6/15/47		360,000		406,038
Humana, Inc.
2.150% 2/03/32		435,000		381,195
Molina Healthcare, Inc.
3.875% 11/15/30 (c)		90,000		86,625
3.875% 5/15/32 (c)		190,000		180,680
4.375% 6/15/28 (c)		300,000		295,273
Universal Health Services, Inc.
1.650% 9/01/26 (c)		385,000		353,647
				6,185,072
Household Products & Wares — 0.0%
Central Garden & Pet Co.
5.125% 2/01/28		123,000		122,101
Insurance — 1.0%
Aon Corp. / Aon Global Holdings PLC
3.900% 2/28/52		190,000		187,362
Athene Global Funding
SOFR + .700% 0.914% FRN 5/24/24 (c)		505,000		498,654
1.985% 8/19/28 (c)		555,000		488,510
3.205% 3/08/27 (c)		135,000		130,150
Berkshire Hathaway Finance Corp.
3.850% 3/15/52		405,000		413,903
Farmers Exchange Capital 3 mo. USD LIBOR + 3.454%
5.454% VRN 10/15/54 (c)		970,000		1,102,878
Teachers Insurance & Annuity Association of America
3.300% 5/15/50 (c)		225,000		199,087
4.270% 5/15/47 (c)		130,000		137,275
Willis North America, Inc.
4.500% 9/15/28		310,000		319,098
				3,476,917
Internet — 0.1%
Tencent Holdings Ltd.
3.680% 4/22/41 (c)		335,000		295,828
Media — 1.1%
CCO Holdings LLC/CCO Holdings Capital Corp.
4.500% 5/01/32		250,000		229,050
4.750% 2/01/32 (c)		200,000		186,408
Charter Communications Operating LLC/Charter Communications Operating Capital
3.750% 2/15/28		92,000		90,924
4.800% 3/01/50		330,000		313,319
4.908% 7/23/25		205,000		212,822
5.250% 4/01/53		265,000		268,387
5.375% 5/01/47		95,000		97,493
5.750% 4/01/48		280,000		297,546
Cox Communications, Inc.
2.600% 6/15/31 (c)		385,000		347,903
CSC Holdings LLC
4.125% 12/01/30 (c)		200,000		175,373
4.500% 11/15/31 (c)		375,000		335,625
5.375% 2/01/28 (c)		30,000		29,117
Diamond Sports Group LLC / Diamond Sports Finance Co.
5.375% 8/15/26 (c)		665,000		257,688

MML Total Return Bond Fund —  Portfolio of Investments (Continued)

	Principal Amount		Value
Sinclair Television Group, Inc.
4.125% 12/01/30 (c)	$	215,000	$	191,709
Time Warner Cable, Inc.
5.500% 9/01/41		170,000		176,625
5.875% 11/15/40		170,000		182,851
Virgin Media Secured Finance PLC
5.500% 5/15/29 (c)		255,000		252,555
The Walt Disney Co.
3.600% 1/13/51		320,000		316,225
				3,961,620
Mining — 0.1%
Indonesia Asahan Aluminium Persero PT
6.530% 11/15/28 (c)		200,000		222,222
Miscellaneous - Manufacturing — 0.2%
GE Capital International Funding Co. Unlimited Co.
4.418% 11/15/35		540,000		579,861
General Electric Co.
6.750% 3/15/32		162,000		204,107
				783,968
Oil & Gas — 0.7%
Antero Resources Corp.
8.375% 7/15/26 (c)		32,000		35,250
Exxon Mobil Corp.
3.452% 4/15/51		393,000		380,750
KazMunay Gas National Co.
4.750% 4/19/27		200,000		191,512
Occidental Petroleum Corp.
0.000% 10/10/36		230,000		123,338
Pertamina Persero PT
3.100% 8/27/30 (c)		200,000		188,500
Petroleos Mexicanos
5.950% 1/28/31 (d)		50,000		46,151
6.625% 6/15/35		900,000		806,634
7.690% 1/23/50		115,000		100,481
Saudi Arabian Oil Co.
1.625% 11/24/25		200,000		189,656
Shell International Finance BV
3.750% 9/12/46		25,000		25,271
Sunoco LP / Sunoco Finance Corp.
4.500% 4/30/30 (c)		126,000		116,277
Transocean Guardian Ltd.
5.875% 1/15/24 (c)		92,172		88,255
Transocean Pontus Ltd.
6.125% 8/01/25 (c)		106,395		106,129
Transocean Poseidon Ltd.
6.875% 2/01/27 (c)		91,875		90,956
				2,489,160
Oil & Gas Services — 0.1%
Archrock Partners LP / Archrock Partners Finance Corp.
6.250% 4/01/28 (c)		180,000		177,406
Transocean Proteus Ltd.
6.250% 12/01/24 (c)		41,500		41,189

MML Total Return Bond Fund —  Portfolio of Investments (Continued)

	Principal Amount		Value
USA Compression Partners LP/USA Compression Finance Corp.
6.875% 4/01/26	$	30,000	$	30,352
6.875% 9/01/27		96,000		96,369
				345,316
Packaging & Containers — 0.5%
Amcor Finance USA, Inc.
3.625% 4/28/26		650,000		660,242
Ardagh Packaging Finance PLC / Ardagh Holdings USA, Inc.
5.250% 8/15/27 (c)		190,000		176,225
5.250% 8/15/27 (c) (d)		200,000		185,500
Berry Global, Inc.
1.570% 1/15/26		410,000		381,179
4.875% 7/15/26 (c)		330,000		334,198
Mauser Packaging Solutions Holding Co.
5.500% 4/15/24 (c)		30,000		29,881
Sealed Air Corp.
4.000% 12/01/27 (c)		30,000		29,250
				1,796,475
Pharmaceuticals — 1.4%
AbbVie, Inc.
4.450% 5/14/46		400,000		428,123
4.500% 5/14/35		390,000		418,638
Bayer US Finance II LLC
4.250% 12/15/25 (c)		75,000		76,356
4.375% 12/15/28 (c)		850,000		870,634
4.625% 6/25/38 (c)		795,000		825,423
4.875% 6/25/48 (c)		210,000		225,642
Cigna Corp.
3.400% 3/15/51		250,000		225,126
4.125% 11/15/25		60,000		61,919
CVS Health Corp.
5.050% 3/25/48		1,090,000		1,236,943
Embecta Corp.
5.000% 2/15/30 (c)		200,000		188,641
Endo DAC / Endo Finance LLC / Endo Finco, Inc.
6.000% 6/30/28 (c) (d)		656,000		370,640
				4,928,085
Pipelines — 1.1%
Energy Transfer LP
4.950% 6/15/28		300,000		314,600
5.000% 5/15/50		380,000		384,313
5.150% 3/15/45		400,000		404,059
5.350% 5/15/45		50,000		51,497
5.400% 10/01/47		394,000		411,038
5.500% 6/01/27		254,000		272,873
Galaxy Pipeline Assets Bidco Ltd.
2.160% 3/31/34 (c)		384,360		353,553
Plains All American Pipeline LP/PAA Finance Corp.
3.800% 9/15/30		300,000		294,303
Rockies Express Pipeline LLC
4.800% 5/15/30 (c)		250,000		236,875
4.950% 7/15/29 (c)		200,000		193,748

MML Total Return Bond Fund —  Portfolio of Investments (Continued)

	Principal Amount		Value
Ruby Pipeline LLC
8.000% STEP 4/01/22 (c)	$	431,818	$	398,352
TransMontaigne Partners LP/TLP Finance Corp.
6.125% 2/15/26		81,000		80,043
Venture Global Calcasieu Pass LLC
3.875% 8/15/29 (c)		365,000		354,351
				3,749,605
Real Estate Investment Trusts (REITS) — 1.3%
American Campus Communities Operating Partnership LP
3.750% 4/15/23		650,000		656,564
CyrusOne LP/CyrusOne Finance Corp.
2.150% 11/01/30		385,000		384,407
3.450% 11/15/29		275,000		283,250
GLP Capital LP/GLP Financing II, Inc.
4.000% 1/15/30		35,000		33,971
5.250% 6/01/25		205,000		211,009
5.300% 1/15/29		310,000		325,677
5.375% 4/15/26		715,000		747,128
SL Green Operating Partnership LP
3.250% 10/15/22		500,000		501,482
Ventas Realty LP
3.500% 2/01/25		1,500,000		1,505,717
				4,649,205
Retail — 0.2%
Michaels Cos., Inc.
5.250% 5/01/28 (c)		170,000		156,077
Starbucks Corp.
3.000% 2/14/32		385,000		368,470
				524,547
Savings & Loans — 0.4%
Nationwide Building Society
SOFR + 1.290% 2.972% VRN 2/16/28 (c)		340,000		324,000
3 mo. USD LIBOR + 1.181% 3.622% VRN 4/26/23 (c)		270,000		270,054
3 mo. USD LIBOR + 1.064% 3.766% VRN 3/08/24 (c)		525,000		528,014
3 mo. USD LIBOR + 1.392% 4.363% VRN 8/01/24 (c)		190,000		193,017
				1,315,085
Semiconductors — 0.1%
Broadcom, Inc.
4.300% 11/15/32		360,000		365,811
Intel Corp.
3.050% 8/12/51 (d)		75,000		66,779
				432,590
Software — 0.3%
Change Healthcare Holdings LLC/Change Healthcare Finance, Inc.
5.750% 3/01/25 (c)		144,000		144,000
Oracle Corp.
2.800% 4/01/27		140,000		133,797
2.875% 3/25/31		195,000		177,939

MML Total Return Bond Fund —  Portfolio of Investments (Continued)

	Principal Amount		Value
3.950% 3/25/51	$	753,000	$	657,938
				1,113,674
Telecommunications — 3.1%
AT&T, Inc.
2.550% 12/01/33		634,000		562,749
3.800% 12/01/57		1,692,000		1,542,748
4.750% 5/15/46		710,000		770,756
5.250% 3/01/37		270,000		306,967
CommScope, Inc.
4.750% 9/01/29 (c)		274,000		252,765
Intelsat Jackson Holdings S.A.
6.500% 3/15/30		1,052,000		1,049,370
Intelsat Jackson Holdings S.A., Escrow
8.500% 10/15/24 (c) (f) (g)		171,000		—
Intelsat Jackson Holdings S.A., Escrow
9.750% 7/15/25 (c) (f) (g)		585,000		—
Level 3 Financing, Inc.
3.400% 3/01/27 (c)		60,000		56,550
3.875% 11/15/29 (c)		460,000		423,200
Lumen Technologies, Inc.
5.375% 6/15/29 (c)		385,000		342,136
Sprint Corp.
7.875% 9/15/23		33,000		35,063
Sprint Spectrum Co. LLC/Sprint Spectrum Co. II LLC/Sprint Spectrum Co. III LLC
4.738% 3/20/25 (c)		1,057,500		1,080,345
T-Mobile USA, Inc.
2.550% 2/15/31		900,000		816,237
3.750% 4/15/27		375,000		377,556
3.875% 4/15/30		345,000		346,367
4.375% 4/15/40		290,000		291,487
4.750% 2/01/28		119,000		121,261
Verizon Communications, Inc.
2.100% 3/22/28		170,000		158,740
2.355% 3/15/32 (c)		520,000		468,903
2.550% 3/21/31		510,000		473,256
Vodafone Group PLC
4.875% 6/19/49		1,052,000		1,144,199
				10,620,655
Transportation — 0.1%
Union Pacific Corp.
3.500% 2/14/53		195,000		190,685
TOTAL CORPORATE DEBT (Cost $107,148,549)				102,487,987
MUNICIPAL OBLIGATIONS — 1.1%
California State University
Revenue, 2.274% 11/01/34		180,000		155,179
Revenue, 2.374% 11/01/35		20,000		17,191
City of New York NY, General Obligation
5.206% 10/01/31		750,000		825,565
Jersey City Municipal Utilities Authority, Revenue Bond
5.470% 5/15/27		745,000		794,101

MML Total Return Bond Fund —  Portfolio of Investments (Continued)

	Principal Amount		Value
New York City Transitional Finance Authority Future Tax Secured Revenue, Revenue
2.450% 11/01/34	$	560,000	$	473,812
New York State Dormitory Authority, Revenue Bond
5.289% 3/15/33		750,000		836,650
University of California, Revenue
2.247% 5/15/34		290,000		255,624
University of Michigan, Revenue
3.504% 4/01/52		380,000		386,399
				3,744,521
TOTAL MUNICIPAL OBLIGATIONS (Cost $4,075,096)				3,744,521
NON-U.S. GOVERNMENT AGENCY OBLIGATIONS — 14.9%
Commercial Mortgage-Backed Securities — 2.6%
BAMLL Commercial Mortgage Securities Trust, Series 2018-PARK, Class A,
4.091% VRN 8/10/38 (c) (h)		620,000		637,695
BX Commercial Mortgage Trust, Series 2019-XL, Class A, 1 mo. USD LIBOR + .920%
1.317% FRN 10/15/36 (c)		1,313,924		1,305,794
BX Trust, Series 2019-OC11, Class A
3.202% 12/09/41 (c)		180,000		174,177
CALI Mortgage Trust, Series 2019-101C, Class A
3.957% 3/10/39 (c)		415,000		420,915
CPT Mortgage Trust, Series 2019-CPT, Class A
2.865% 11/13/39 (c)		355,000		335,796
DC Office Trust, Series 2019-MTC, Class A
2.965% 9/15/45 (c)		375,000		354,902
GS Mortgage Securities Corp. II, Series 2021-ARDN, Class A, 1 mo. USD LIBOR + 1.250%
1.647% FRN 11/15/36 (c)		1,570,000		1,554,796
Hudson Yards Mortgage Trust
Series 2019-55HY, Class A, 2.943% VRN 12/10/41 (c) (h)		375,000		357,703
Series 2019-30HY, Class A, 3.228% 7/10/39 (c)		385,000		373,820
JP Morgan Chase Commercial Mortgage Securities Trust, Series 2019-OSB, Class A
3.397% 6/05/39 (c)		385,000		379,817
MKT Mortgage Trust, Series 2020-525M, Class A
2.694% 2/12/40 (c)		465,000		432,989
Natixis Commercial Mortgage Securities Trust, 1 mo. USD LIBOR + .950%
1.347% FRN 8/15/38 (c)		1,100,000		1,091,363
One Bryant Park Trust, Series 2019-OBP, Class A
2.516% 9/15/54 (c)		490,000		453,397
RBS Commercial Funding, Inc. Trust, Series 2013-GSP, Class A,
3.834% VRN 1/15/32 (c) (h)		670,000		669,016
SFAVE Commercial Mortgage Securities Trust, Series 2015-5AVE, Class A1,
3.872% VRN 1/05/43 (c) (h)		655,000		621,642
				9,163,822
Home Equity Asset-Backed Securities — 1.2%
Aegis Asset Backed Securities Trust Mortgage Pass-Through Ctfs, Series 2005-4, Class M2, 1 mo. USD LIBOR + .705%
1.162% FRN 10/25/35		780,000		765,834
Citigroup Mortgage Loan Trust, Series 2006-HE2, Class M1, 1 mo. USD LIBOR + .435%
0.892% FRN 8/25/36		977,738		971,584

MML Total Return Bond Fund —  Portfolio of Investments (Continued)

	Principal Amount		Value
Morgan Stanley Capital, Inc. Trust, Series 2006-HE1, Class A4, 1 mo. USD LIBOR + .580%
1.037% FRN 1/25/36	$	1,085,162	$	1,074,949
Opteum Mortgage Acceptance Corp.
1.207% 11/25/35		1,450,000		1,390,986
				4,203,353
Other Asset-Backed Securities — 4.7%
Aimco CLO 11 Ltd., Series 2020-11A, Class AR, 3 mo. USD LIBOR + 1.130%
1.371% FRN 10/17/34 (c)		750,000		741,899
AIMCO CLO Series 2015-A, Series 2015-AA, Class BR2, 3 mo. USD LIBOR + 1.600%
1.841% FRN 10/17/34 (c)		1,100,000		1,087,758
Allegro CLO XII Ltd., Series 2020-1A, Class A1, 3 mo. USD LIBOR + 1.250%
1.505% FRN 1/21/32 (c)		850,000		844,732
Cedar Funding XIV CLO Ltd., Series 2021-14A, Class A, 3 mo. USD LIBOR + 1.100%
1.341% FRN 7/15/33 (c)		800,000		793,398
Dryden Senior Loan Fund, Series 2013-26A, Class AR, 3 mo. USD LIBOR + .900%
1.141% FRN 4/15/29 (c)		934,966		931,045
Eaton Vance CLO 2020-1 Ltd., Series 2020-1A, Class AR, 3 mo. USD LIBOR + 1.170%
1.411% FRN 10/15/34 (c)		990,000		984,654
Flatiron CLO Ltd., Series 2021-1A, Class B, 3 mo. USD LIBOR + 1.600%
1.848% FRN 7/19/34 (c)		850,000		835,745
GSAMP Trust 2005-HE5, Series 2005-HE5, Class M3, 1 mo. USD LIBOR + .690%
1.147% FRN 11/25/35		646,473		643,024
Highbridge Loan Management ., Series 2021-16A, Class A1, 3 mo. USD LIBOR + 1.140%
1.395% FRN 1/23/35 (c)		1,000,000		991,625
LCM LP, Series 21A, Class AR, 3 mo. USD LIBOR + .880%
1.134% FRN 4/20/28 (c)		281,336		280,250
Magnetite Ltd., Series 2012-7A, Class A1R2, 3 mo. USD LIBOR + .800%
1.041% FRN 1/15/28 (c)		950,088		946,074
Progress Residential Trust
Series 2019-SFR2, Class A, 3.147% 5/17/36 (c)		2,665,407		2,639,048
Series 2019-SFR2, Class F, 4.837% 5/17/36 (c)		2,250,000		2,196,254
Rockford Tower CLO Ltd., Series 2021-1A, Class A1, 3 mo. USD LIBOR + 1.170%
1.424% FRN 7/20/34 (c)		850,000		844,692
Saxon Asset Securities Trust, Series 2007-2, Class A2C, 1 mo. USD LIBOR + .240%
0.697% FRN 5/25/47		1,664,279		1,335,633
				16,095,831
Student Loans Asset-Backed Securities — 2.6%
Access Group, Inc., Series 2015-1, Class A, 1 mo. USD LIBOR + .700%
1.157% FRN 7/25/56 (c)		395,804		391,005
Collegiate Funding Services Education Loan Trust, Series 2005-A, Class A4, 3 mo. USD LIBOR + .200%
1.166% FRN 3/28/35		1,287,178		1,276,596

MML Total Return Bond Fund —  Portfolio of Investments (Continued)

	Principal Amount		Value
Nelnet Student Loan Trust, Series 2015-1A, Class A, 1 mo. USD LIBOR + .590%
1.047% FRN 4/25/46 (c)	$	770,762	$	766,671
SLM Student Loan Trust
Series 2007-7, Class A4, 3 mo. USD LIBOR + .330% 0.588% FRN 1/25/22		650,742		637,789
Series 2012-7, Class A3, 1 mo. USD LIBOR + .650% 1.107% FRN 5/26/26		693,292		684,368
Series 2012-2, Class A, 1 mo. USD LIBOR + .700% 1.157% FRN 1/25/29		806,931		780,873
Series 2008-6, Class A4, 3 mo. USD LIBOR + 1.100% 1.358% FRN 7/25/23		971,684		962,015
Series 2012-1, Class A3, 1 mo. USD LIBOR + .950% 1.407% FRN 9/25/28		683,092		669,976
Series 2008-9, Class A, 3 mo. USD LIBOR + 1.500% 1.758% FRN 4/25/23		711,204		710,986
Series 2008-5, Class B, 3 mo. USD LIBOR + 1.850% 2.108% FRN 7/25/73		700,000		689,657
Series 2008-9, Class B, 3 mo. USD LIBOR + 2.250% 2.508% FRN 10/25/83		1,430,000		1,434,904
				9,004,840
Whole Loan Collateral Collateralized Mortgage Obligations — 3.8%
American Home Mortgage Investment Trust, Series 2005-1, Class 6A, 6 mo. USD LIBOR + 2.000%
2.826% FRN 6/25/45		440,757		434,808
Chase Mortgage Finance Trust, Series 2007-A1, Class 11A4,
3.147% VRN 3/25/37 (h)		247,398		237,907
Citigroup Mortgage Loan Trust, Series 2005-11, Class A2A, 1 year CMT + 2.400%
2.470% FRN 10/25/35		192,803		194,925
Countrywide Alternative Loan Trust
Series 2006-OA6, Class 1A2, 1 mo. USD LIBOR + .420% 0.877% FRN 7/25/46		918,855		872,684
Series 2005-31, Class 2A1, 1 mo. USD LIBOR + .600% 1.057% FRN 8/25/35		934,632		872,791
Credit Suisse Mortgage Trust, Series 2018-RPL9, Class A,
3.850% VRN 9/25/57 (c) (h)		921,605		927,587
First Horizon Mortgage Pass-Through Trust, Series 2005-AR5, Class 2A1,
2.710% VRN 11/25/35 (h)		248,233		212,881
GS Mortgage-Backed Securities Trust, Series 2018-RPL1, Class A1A
3.750% 10/25/57 (c)		817,269		811,633
HarborView Mortgage Loan Trust, Series 2006-10, Class 1A1A, 1 mo. USD LIBOR + .200%
0.362% FRN 11/19/36		1,705,078		1,499,627
Impac CMB Trust, Series 2005-1, Class 1A1, 1 mo. USD LIBOR + .520%
0.977% FRN 4/25/35		524,671		502,066
IndyMac Index Mortgage Loan Trust
Series 2006-AR27, Class 1A3, 1 mo. USD LIBOR + .540% 0.997% FRN 10/25/36		1,322,493		683,749
Series 2005-AR19, Class A1, 2.770% VRN 10/25/35 (h)		707,074		626,156
JP Morgan Mortgage Trust, Series 2021-13, Class A3,
2.500% VRN 4/25/52 (c) (h)		1,835,316		1,696,018
Nomura Asset Acceptance Corp. Alternative Loan Trust, Series 2005-AR1, Class M1, 1 mo. USD LIBOR + 1.070%
1.527% FRN 2/25/35		713,917		701,403
Washington Mutual Mortgage Pass-Through Certificates
Series 2005-AR15, Class A1A1, 1 mo. USD LIBOR + .520% 0.977% FRN 11/25/45		1,785,688		1,686,098
Series 2005-AR13, Class A1A1, 1 mo. USD LIBOR + .580% 1.037% FRN 10/25/45		391,768		383,724
Series 2005-AR2, Class 2A21, 1 mo. USD LIBOR + .660% 1.117% FRN 1/25/45		338,596		334,875

MML Total Return Bond Fund —  Portfolio of Investments (Continued)

	Principal Amount		Value
Series 2006-1 Class 3A2, 5.750% 2/25/36	$	555,034	$	520,623
				13,199,555
TOTAL NON-U.S. GOVERNMENT AGENCY OBLIGATIONS (Cost $52,570,641)				51,667,401
SOVEREIGN DEBT OBLIGATIONS — 1.3%
Abu Dhabi Government International Bond
2.500% 9/30/29		200,000		195,845
Brazilian Government International Bond
2.875% 6/06/25		200,000		196,044
3.875% 6/12/30		200,000		184,290
CBB International Sukuk Co.
6.875% 10/05/25		200,000		219,410
Chile Government International Bond
3.240% 2/06/28		200,000		199,870
Colombia Government International Bond
3.000% 1/30/30 (d)		200,000		172,244
4.500% 1/28/26		200,000		199,774
Dominican Republic International Bond
4.500% 1/30/30 (c)		200,000		183,502
Hong Kong Airport Authority
3.250% 1/12/52 (c)		405,000		361,267
Hungary Government International Bond
2.125% 9/22/31 (c)		200,000		180,757
Mexico Government International Bond
2.659% 5/24/31 (d)		288,000		261,991
3.750% 1/11/28		300,000		305,484
Oman Government International Bond
5.625% 1/17/28		200,000		206,197
Panama Government International Bond
2.252% 9/29/32		200,000		176,894
3.160% 1/23/30 (d)		200,000		195,716
3.875% 3/17/28		200,000		204,306
Peruvian Government International Bond
2.844% 6/20/30		80,000		76,618
4.125% 8/25/27		200,000		207,826
Qatar Government International Bond
4.500% 4/23/28		200,000		217,750
Republic of South Africa Government International Bond
4.850% 9/30/29 (d)		250,000		244,239
Republic of Turkey International Bond
3.250% 3/23/23		200,000		196,296
Romanian Government International Bond
3.000% 2/14/31		110,000		101,555
				4,487,875
TOTAL SOVEREIGN DEBT OBLIGATIONS (Cost $4,851,469)				4,487,875
U.S. GOVERNMENT AGENCY OBLIGATIONS AND INSTRUMENTALITIES (i) — 35.9%
Collateralized Mortgage Obligations — 1.9%
Federal Home Loan Mortgage Corp.
Series 4818, Class CA, 3.000% 4/15/48		175,354		173,238
Series 4639, Class HZ, 3.250% STEP 4/15/53		1,574,768		1,600,732
Series 4846, Class PA, 4.000% 6/15/47		114,102		115,880

MML Total Return Bond Fund —  Portfolio of Investments (Continued)

	Principal Amount		Value
Federal National Mortgage Association REMICS
Series 2018-43, Class CT, 3.000% 6/25/48	$	313,164	$	309,110
Series 2018-57, Class QA, 3.500% 5/25/46		165,649		167,635
Series 2018-55, Class PA, 3.500% 1/25/47		122,864		124,525
Government National Mortgage Association
Series 2018-124, Class NW, 3.500% 9/20/48		310,829		312,796
Series 2019-15, Class GT, 3.500% 2/20/49		204,977		206,360
				3,010,276
Pass-Through Securities — 32.0%
Federal Home Loan Mortgage Corp.
Pool #G18592 3.000% 3/01/31		202,434		204,934
Pool #G18627 3.000% 1/01/32		636,948		644,815
Pool #G08710 3.000% 6/01/46		112,650		111,946
Pool #ZM1779 3.000% 9/01/46		563,442		559,334
Pool #G08726 3.000% 10/01/46		836,413		830,921
Pool #G08732 3.000% 11/01/46		304,287		302,289
Pool #G08741 3.000% 1/01/47		835,308		829,300
Pool #G07848 3.500% 4/01/44		1,370,257		1,404,188
Pool #G60023 3.500% 4/01/45		1,267,299		1,294,720
Pool #G08711 3.500% 6/01/46		376,769		382,802
Pool #G08716 3.500% 8/01/46		541,973		550,652
Pool #G67700 3.500% 8/01/46		624,141		634,916
Pool #G08722 3.500% 9/01/46		94,211		95,720
Pool #G08742 3.500% 1/01/47		428,398		434,856
Pool #G08757 3.500% 4/01/47		110,257		111,919
Pool #G67703 3.500% 4/01/47		911,840		927,581
Pool #G67706 3.500% 12/01/47		822,318		836,256
Pool #ZA5103 3.500% 12/01/47		4,445		4,504
Pool #G67707 3.500% 1/01/48		2,286,347		2,333,674
Pool #G08800 3.500% 2/01/48		39,131		39,562
Pool #G67708 3.500% 3/01/48		200,397		203,919
Pool #G67710 3.500% 3/01/48		1,054,924		1,069,838
Pool #G08816 3.500% 6/01/48		307,368		309,985
Pool #G61556 3.500% 8/01/48		540,634		547,939
Pool #G60344 4.000% 12/01/45		681,498		708,810
Pool #G67711 4.000% 3/01/48		750,104		778,994
Pool #G67713 4.000% 6/01/48		3,104		3,212
Pool #G08843 4.500% 10/01/48		161,450		168,255
Pool #G08833 5.000% 7/01/48		116,028		122,885
Pool #G08844 5.000% 10/01/48		103,234		108,835
Federal National Mortgage Association
Pool #MA4152 2.000% 10/01/40		873,665		819,682
Pool #MA4176 2.000% 11/01/40		1,707,116		1,601,635
Pool #MA4333 2.000% 5/01/41		205,910		193,187
Pool #BQ7006 2.000% 1/01/52 (e)		3,724,575		3,464,175
Pool #BL6060 2.455% 4/01/40		570,000		507,166
Pool #MA1607 3.000% 10/01/33		333,607		333,781
Pool #MA3811 3.000% 10/01/49		225,415		218,910
Pool #AB4262 3.500% 1/01/32		288,098		293,390
Pool #MA1512 3.500% 7/01/33		145,499		148,717
Pool #MA1148 3.500% 8/01/42		832,239		853,245
Pool #CA0996 3.500% 1/01/48		143,697		145,483
Pool #MA3305 3.500% 3/01/48		29,419		29,693
Pool #BL1132 3.730% 1/01/29		920,000		965,705
Pool #MA1146 4.000% 8/01/42		338,301		354,538
Pool #AS9830 4.000% 6/01/47		162,101		167,298
Pool #MA3027 4.000% 6/01/47		224,012		231,193

MML Total Return Bond Fund —  Portfolio of Investments (Continued)

	Principal Amount		Value
Pool #AS9972 4.000% 7/01/47	$	144,565	$	149,064
Pool #931504 4.500% 7/01/39		45,263		47,755
Pool #CA1710 4.500% 5/01/48		543,606		564,722
Pool #CA1711 4.500% 5/01/48		172,585		180,011
Pool #CA2208 4.500% 8/01/48		321,973		335,676
Pool #AD6374 5.000% 5/01/40		9,988		10,722
Pool #AI2733 5.000% 5/01/41		86,587		94,045
Pool #977014 5.500% 5/01/38		12,359		13,529
Pool #985524 5.500% 6/01/38		13,477		14,760
Pool #988578 5.500% 8/01/38		109,201		119,315
Pool #995482 5.500% 1/01/39		68,958		75,403
Government National Mortgage Association II
Pool #MA4126 3.000% 12/20/46		687,050		686,632
Pool #MA4718 3.000% 9/20/47		75,746		75,677
Pool #MA4836 3.000% 11/20/47		265,114		264,870
Pool #MA6209 3.000% 10/20/49		219,915		215,108
Pool #MA3521 3.500% 3/20/46		272,330		278,491
Pool #MA3597 3.500% 4/20/46		84,396		86,305
Pool #MA3663 3.500% 5/20/46		123,924		126,728
Pool #MA3937 3.500% 9/20/46		102,125		104,340
Pool #MA4127 3.500% 12/20/46		368,744		376,165
Pool #MA4262 3.500% 2/20/47		375,499		383,055
Pool #MA4382 3.500% 4/20/47		96,893		98,843
Pool #MA4719 3.500% 9/20/47		120,879		123,085
Pool #MA4837 3.500% 11/20/47		248,384		252,374
Pool #MA4900 3.500% 12/20/47		895,939		910,329
Pool #MA4653 4.000% 8/20/47		48,010		49,519
Pool #MA4838 4.000% 11/20/47		202,707		208,823
Pool #MA4901 4.000% 12/20/47		87,270		89,903
Pool #MA5078 4.000% 3/20/48		302,520		311,175
Pool #MA5466 4.000% 9/20/48		333,474		341,998
Pool #MA5399 4.500% 8/20/48		630,254		655,887
Pool #MA3666 5.000% 5/20/46		96,450		103,632
Pool #MA4199 5.000% 1/20/47		95,812		101,783
Pool #MA4454 5.000% 5/20/47		162,183		173,500
Pool #MA4722 5.000% 9/20/47		193,649		206,011
Government National Mortgage Association II TBA
2.000% 4/21/52 (e)		1,925,000		1,831,307
2.500% 4/21/52 (e)		5,225,000		5,070,291
Uniform Mortgage Backed Securities TBA
2.000% 4/18/37 (e)		5,050,000		4,904,812
2.000% 4/13/52 (e)		1,900,000		1,764,477
2.000% 5/12/52 (e)		24,400,000		22,618,610
2.500% 4/18/37 (e)		3,950,000		3,903,402
2.500% 5/12/52 (e)		29,325,000		27,943,517
3.000% 4/13/52 (e)		3,650,000		3,573,293
3.000% 5/12/52 (e)		8,050,000		7,862,586
				114,186,889
U.S. Agencies Notes — 1.1%
Federal Home Loan Banks 1.040% 6/14/24		4,110,000		3,980,267
Whole Loans — 0.9%
Deutsche Alt-A Securities, Inc. Mortgage Loan Trust Series 2005-1, Series 2005-1, Class 1A3, 1 mo. USD LIBOR + .500%
0.957% FRN 2/25/35		1,620,084		1,565,492

MML Total Return Bond Fund —  Portfolio of Investments (Continued)

	Principal Amount		Value
MASTR Adjustable Rate Mortgages Trust 2007-1, Series 2007-1, Class I2A3, 12 mo. MTA + .740%
0.881% FRN 1/25/47	$	1,627,908	$	1,606,061
				3,171,553
TOTAL U.S. GOVERNMENT AGENCY OBLIGATIONS AND INSTRUMENTALITIES (Cost $125,544,956)				124,348,985
U.S. TREASURY OBLIGATIONS — 31.1%
U.S. Treasury Bonds & Notes — 31.1%
U.S. Treasury Bond
2.000% 11/15/41		18,132,000		16,431,920
2.250% 5/15/41		5,014,000		4,741,282
2.250% 2/15/52		6,054,000		5,802,384
2.375% 2/15/42		585,000		564,616
U.S. Treasury Inflation Index
0.125% 2/15/52		631,056		665,715
U.S. Treasury Note
0.750% 12/31/23		3,437,000		3,349,847
0.875% 1/31/24		9,608,000		9,370,161
1.250% 12/31/26		30,000		28,381
1.500% 2/29/24 (d)		12,762,000		12,585,396
1.500% 1/31/27		7,553,000		7,228,635
1.750% 3/15/25		11,850,000		11,611,151
1.875% 2/28/27 (d)		18,465,000		17,997,662
1.875% 2/15/32		1,015,000		974,558
2.250% 3/31/24		3,415,000		3,412,466
2.375% 3/31/27 (e)		12,825,000		12,870,093
				107,634,267
TOTAL U.S. TREASURY OBLIGATIONS (Cost $110,010,009)				107,634,267
TOTAL BONDS & NOTES (Cost $411,091,443)				401,183,773
TOTAL PURCHASED OPTIONS (#) — 0.0% (Cost $164,390)				45,644
RIGHTS — 0.0%
Communications — 0.0%
Intelsat Jackson Holdings S.A. (f) (g)		755		—
Intelsat Jackson Holdings S.A. (f) (g)		755		—
				—
TOTAL RIGHTS (Cost $0)				—

MML Total Return Bond Fund —  Portfolio of Investments (Continued)

	Number of Shares		Value
MUTUAL FUNDS — 9.5%
Diversified Financial Services — 9.5%
State Street Navigator Securities Lending Government Money Market Portfolio (j)		32,765,090	$	32,765,090
TOTAL MUTUAL FUNDS (Cost $32,765,090)				32,765,090
TOTAL LONG-TERM INVESTMENTS (Cost $444,697,917)				434,204,498
		Principal Amount
SHORT-TERM INVESTMENTS — 7.9%
Repurchase Agreement — 2.1%
Fixed Income Clearing Corp., Repurchase Agreement, dated 3/31/22, 0.000%, due 4/01/22 (k)	$	7,125,364		7,125,364
U.S. Treasury Bill — 5.8%
U.S. Treasury Bill
0.610% 8/25/22 (l)		4,275,000		4,259,441
0.651% 9/01/22 (l)		3,440,000		3,426,424
0.674% 8/25/22 (l)		1,125,000		1,120,906
0.983% 9/22/22 (l)		4,600,000		4,578,326
0.984% 9/22/22 (l)		6,800,000		6,767,960
				20,153,057
TOTAL SHORT-TERM INVESTMENTS (Cost $27,289,392)				27,278,421
TOTAL INVESTMENTS — 133.4% (Cost $471,987,309) (m)				461,482,919
Less Unfunded Loan Commitments — —%				(35,326)
NET INVESTMENTS — 133.4% (Cost $471,951,983)				461,447,593
Other Assets/(Liabilities) — (33.4)%				(115,430,322)
NET ASSETS — 100.0%			$	346,017,271

Abbreviation Legend

CLO Collateralized Loan Obligation

FRN Floating Rate Note

REMICS Real Estate Mortgage Investment Conduits

STEP Step Up Bond

TBA To Be Announced

VRN Variable Rate Note

Notes to Portfolio of Investments

Percentages are stated as a percent of net assets.

(a) All or a portion of the security represents unsettled loan commitments at March 31, 2022 where the rate will be determined at time of settlement

(b) Unfunded or partially unfunded loan commitments.

(c) Security is exempt from registration under Regulation S or Rule 144A of the Securities Act of 1933. These securities are considered restricted and may be resold in transactions exempt from registration. At March 31, 2022, the aggregate market value of these securities amounted to $58,007,258 or 16.76% of net assets.

(d) Denotes all or a portion of security on loan. The total value of securities on loan as of March 31, 2022, was $33,358,022 or 9.64% of net assets. Total securities on loan may be less than the amounts identified in the Portfolio of Investments. The Fund received $1,300,341 of non-cash collateral (U.S. Treasury and/or Agency obligations) related to securities lending activity. This amount is not reflected on the Portfolio of Investments. (Note 2).

(e) A portion of this security is purchased on a when-issued, delayed-delivery or forward commitment basis.

(f) Investment was valued using significant unobservable inputs.

(g) This security is fair valued in good faith in accordance with procedures approved by the Board of Trustees. At March 31, 2022, these securities amounted to a value of $0 or 0.00% of net assets.

(h) Certain variable rate securities are not based on a published reference rate and spread but are determined by the issuer or agent and are based on current market conditions. These securities do not indicate a reference rate and spread in their description above. The rates shown are the current interest rates at March 31, 2022.

(i) May contain securities that are issued by a U.S. Government Agency, but are unsecured and are not guaranteed by a U.S. Government Agency.

(j) Represents investment of security lending cash collateral. (Note 2).

(k) Maturity value of $7,125,364. Collateralized by U.S. Government Agency obligations with a rate of 0.500%, maturity date of 4/15/24, and an aggregate market value, including accrued interest, of $7,267,885.

(l) The rate shown represents yield-to-maturity.

(m) See Note 3 for aggregate cost for federal tax purposes.

(#) Exchange-Traded Options Purchased
Description	Expiration Date	Exercise Price		No. of Contracts		Notional Amount		Value		Premiums Paid (Received)		Unrealized Appreciation/ (Depreciation)
Call
Eurodollar 1 Year Mid Curve Future	9/16/22		98.38		51	USD	12,352,200	$	4,144	$	21,819	$	(17,675)
Eurodollar 1 Year Mid Curve Future	9/16/22		98.75		110	USD	26,642,000		5,500		65,435		(59,935)
Eurodollar Future	12/19/22		98.00		80	USD	19,468,000		36,000		77,136		(41,136)
								$	45,644	$	164,390	$	(118,746)

MML Total Return Bond Fund —  Portfolio of Investments (Continued)

Exchange-Traded Options Written
Description	Expiration Date	Exercise Price		No. of Contracts		Notional Amount		Value		Premiums Paid (Received)		Unrealized Appreciation/ (Depreciation)
Call
Eurodollar 1 Year Mid Curve Future	9/16/22		99.00		250	USD	60,550,000	$	(9,375)	$	(82,545)	$	73,170
Eurodollar 2 Year Mid Curve Future	12/16/22		97.63		61	USD	14,837,488		(41,937)		(42,616)		679
Eurodollar 2 Year Mid Curve Future	12/16/22		97.88		115	USD	27,972,313		(53,906)		(63,803)		9,897
Eurodollar Future	12/19/22		98.25		80	USD	19,468,000		(25,000)		(51,819)		26,819
								$	(130,218)	$	(240,783)	$	110,565
Put
Eurodollar 1 Year Mid Curve Future	9/16/22		97.63		314	USD	76,050,800	$	(675,100)	$	(214,990)	$	(460,110)
Eurodollar 2 Year Mid Curve Future	12/16/22		97.38		176	USD	42,809,800		(205,700)		(179,595)		(26,105)
Eurodollar Future	12/19/22		97.25		80	USD	19,468,000		(79,000)		(36,819)		(42,181)
								$	(959,800)	$	(431,404)	$	(528,396)
								$	(1,090,018)	$	(672,187)	$	(417,831)

Interest Rate Swaptions Written

Paid by Fund

Received by Fund

Description

Counterparty

Expiration Date

Rate

Frequency

Rate

Frequency

Notional Amount

Value

Premium Paid (Received)

Unrealized Appreciation/ (Depreciation)

Put

2-Year Interest Rate Swap, 6/10/24

JP Morgan Chase Bank N.A.

6/08/22

3-Month SOFR

Quarterly

1.95%##

Semi-Annually

USD

19,270,000

$

(298,300)

$

(55,498)

$

(242,802)

Futures contracts
	Expiration Date	Number of Contracts		Notional Amount		Value/ Net Unrealized Appreciation/ (Depreciation)
Short
U.S. Treasury Ultra 10 Year	6/21/22		32	$	(4,356,187)	$	21,187
U.S. Treasury Note 2 Year	6/30/22		4		(858,868)		11,180
						$	32,367

Centrally Cleared Interest Rate Swaps
Paid by the Fund		Received by the Fund
Rate/ Reference	Frequency	Rate/ Reference		Frequency		Termination Date		Notional Amount		Value		Upfront Premium Paid/ (Received)		Unrealized Appreciation (Depreciation)
3-Month USD LIBOR BBA	Quarterly		Fixed 1.072%		Semi-Annually		7/24/25	USD	3,875,000	$	(134,778)	$	—	$	(134,778)
3-Month USD LIBOR BBA	Quarterly		Fixed 1.034%		Semi-Annually		7/24/25	USD	5,725,000		(203,331)		—		(203,331)
3-Month USD LIBOR BBA	Quarterly		Fixed 1.025%		Semi-Annually		7/24/25	USD	7,750,000		(276,420)		—		(276,420)
3-Month USD LIBOR BBA	Quarterly		Fixed 1.390%		Semi-Annually		9/28/25	USD	15,085,000		(413,098)		—		(413,098)
3-Month USD LIBOR BBA	Quarterly		Fixed 1.688%		Semi-Annually		12/07/25	USD	10,555,000		(214,390)		—		(214,390)
Fixed 1.808%	Semi-Annually		3-Month USD LIBOR BBA		Quarterly		7/24/53	USD	325,000		30,031		—		30,031
Fixed 1.785%	Semi-Annually		3-Month USD LIBOR BBA		Quarterly		7/24/53	USD	475,000		46,203		—		46,203
Fixed 1.773%	Semi-Annually		3-Month USD LIBOR BBA		Quarterly		7/24/53	USD	650,000		64,982		—		64,982
Fixed 1.870%	Semi-Annually		3-Month USD LIBOR BBA		Quarterly		9/28/53	USD	1,285,000		97,891		—		97,891
Fixed 1.743%	Semi-Annually		3-Month USD LIBOR BBA		Quarterly		12/07/53	USD	880,000		87,690		—		87,690
										$	(915,220)	$	—	$	(915,220)

## Exercise Rate.

Currency Legend

USD U.S. Dollar

Notes to Portfolio of Investments (Unaudited)

1. The Funds

MML Series Investment Fund (the ’‘Trust’’) is registered under the Investment Company Act of 1940, as amended (the “1940 Act”), as an open-end management investment company. The Trust is organized under the laws of the Commonwealth of Massachusetts as a Massachusetts business trust pursuant to an Agreement and Declaration of Trust dated December 19, 1984, as restated May 14, 1993, and further amended and restated as of December 15, 2011, as it may be further amended from time to time. The Trust consists of the following series (each individually referred to as a ‘‘Fund’’ or collectively as the ‘‘Funds’’):

MML Conservative Allocation Fund (“Conservative Allocation Fund”)

MML Balanced Allocation Fund (“Balanced Allocation Fund”)

MML Moderate Allocation Fund (“Moderate Allocation Fund”)

MML Growth Allocation Fund (“Growth Allocation Fund”)

MML Aggressive Allocation Fund (“Aggressive Allocation Fund”)

MML American Funds Growth Fund (“MML American Funds Growth Fund”)

MML American Funds International Fund (“MML American Funds International Fund”)

MML American Funds Core Allocation Fund (“MML American Funds Core Allocation Fund”)

MML Blue Chip Growth Fund (“Blue Chip Growth Fund”)

MML Equity Income Fund (“Equity Income Fund”)

MML Equity Index Fund (“Equity Index Fund”)

MML Focused Equity Fund (“Focused Equity Fund”)

MML Foreign Fund (“Foreign Fund”)

MML Fundamental Equity Fund (“Fundamental Equity Fund”)

MML Fundamental Value Fund (“Fundamental Value Fund”)

MML Global Fund (“Global Fund”)

MML Growth & Income Fund (“Growth & Income Fund”)

MML Income & Growth Fund (“Income & Growth Fund”)

MML International Equity Fund (“International Equity Fund”)

MML Large Cap Growth Fund (“Large Cap Growth Fund”)

MML Managed Volatility Fund (“Managed Volatility Fund”)

MML Mid Cap Growth Fund (“Mid Cap Growth Fund”)

MML Mid Cap Value Fund (“Mid Cap Value Fund”)

MML Small Cap Growth Equity Fund (“Small Cap Growth Equity Fund”)

MML Small Company Value Fund (“Small Company Value Fund”)

MML Small/Mid Cap Value Fund (“Small/Mid Cap Value Fund”)

MML Total Return Bond Fund (“Total Return Bond Fund”)

The Trust makes shares of the Funds available for the investment of assets of various separate investment accounts established by Massachusetts Mutual Life Insurance Company (“MassMutual”) and by life insurance companies which are subsidiaries of MassMutual. Shares of the Trust are not offered to the general public. MassMutual, MML Bay State Life Insurance Company, C.M. Life Insurance Company, and the Allocation Funds (as defined below), which are “funds of funds” series of the Trust, are the record owners of all of the outstanding shares of the Funds.

The Conservative Allocation Fund, Balanced Allocation Fund, Moderate Allocation Fund, Growth Allocation Fund, and Aggressive Allocation Fund (the “Allocation Funds”) invest their investable assets in shares of series of the Trust and MML Series Investment Fund II, advised by MML Investment Advisers, LLC and non-affiliated mutual funds (together, the “MML Underlying Funds”).

MML American Funds Growth Fund and MML American Funds International Fund (each a “Feeder Fund,” collectively the “Feeder Funds”) invest all of their assets in Class 1 shares of the Growth and International Funds, respectively, each a series of the American Funds Insurance Series (each a “Master Fund,” collectively the “Master Funds”). Each Master Fund is an open-end investment company and organized as a Massachusetts business trust. Each Feeder Fund has an investment objective that is consistent with its corresponding Master Fund. Each Master Fund intends to comply with the requirements under Subchapter M of the Internal Revenue Code of 1986, as amended (the “Code”), applicable to mutual funds and intends to distribute substantially all of its taxable income and capital gains to shareholders, which includes the applicable Feeder Fund, to qualify as a regulated investment company. The performance of the Feeder Funds is directly affected by the performance of the Master Funds. The MML American Funds Core Allocation Fund invests all of its investable assets in shares of various series of American Funds Insurance Series, which are managed by Capital Research and Management Company (together, the “American Underlying Funds” and collectively with the MML Underlying Funds, the “Underlying Funds”).

Notes to Portfolio of Investments (Unaudited) (Continued)

2. Significant Accounting Policies

Each Fund is an investment company and accordingly follows the investment company accounting and reporting guidance of the Financial Accounting Standards Board (“FASB”) Accounting Standards Codification Topic 946, Financial Services – Investment Companies.

Investment Valuation

The net asset value of each Fund’s shares is determined once daily as of the close of regular trading on the New York Stock Exchange (“NYSE”), on each day the NYSE is open for trading (a “business day”). The NYSE normally closes at 4:00 p.m. Eastern Time, but may close earlier on some days. If the NYSE is scheduled to close early, the business day will be considered to end as of the time of the NYSE’s scheduled close. A Fund will not treat an intraday disruption in NYSE trading or other event that causes an unscheduled closing of the NYSE as a close of business of the NYSE for these purposes and will instead fair value securities in accordance with procedures approved annually by the Board of Trustees (“Trustees”), and under the general oversight of the Trustees. On holidays and other days when the NYSE is closed, each Fund’s net asset value generally is not calculated and the Funds do not anticipate accepting buy or sell orders. However, the value of each Fund’s assets may still be affected on such days to the extent that a Fund or an Underlying Fund or Master Fund holds foreign securities that trade on days that foreign securities markets are open.

The net asset value of each of the Allocation Funds, Feeder Funds, and MML American Funds Core Allocation Fund is based upon the net asset value(s) of its corresponding Master Fund or Underlying Funds. Shares of the Master Fund and Underlying Funds are valued at their closing net asset values as reported on each business day.

The Prospectuses and Statements of Additional Information (“SAIs”) for the Underlying Funds and Master Funds, as applicable, explain the valuation methods for the Underlying Funds and Master Funds, including the circumstances under which the Underlying Funds or Master Funds, may use fair value pricing and the effects of doing so. Such Prospectuses and SAIs are available on the Securities and Exchange Commission’s (“SEC”) EDGAR database on its website at http://www.sec.gov.

Equity securities and derivative contracts that are actively traded on a national securities exchange or contract market are valued on the basis of information furnished by a pricing service, which provides the last reported sale price, or, in the case of futures contracts, the settlement price, for securities or derivatives listed on the exchange or contract market or the official closing price on the NASDAQ National Market System (“NASDAQ System”), or in the case of over-the-counter (“OTC”) securities for which an official closing price is unavailable or not reported on the NASDAQ System, the last reported bid price. Portfolio securities traded on more than one national securities exchange are valued at the last price at the close of the exchange representing the principal market for such securities. Debt securities are valued on the basis of valuations furnished by a pricing service, which generally determines valuations taking into account factors such as institutional-size trading in similar securities, yield, quality, coupon rate, maturity, type of issue, trading characteristics, and other market data. Shares of other open-end mutual funds are valued at their closing net asset values as reported on each business day.

Notes to Portfolio of Investments (Unaudited) (Continued)

Investments for which market quotations are readily available are marked to market daily based on those quotations. Market quotations may be provided by third-party vendors or market makers, and may be determined on the basis of a variety of factors, such as broker quotations, financial modeling, and other market data, such as market indexes and yield curves, counterparty information, and foreign exchange rates. U.S. Government and agency securities may be valued on the basis of market quotations or using a model that may incorporate market observable data such as reported sales of similar securities, broker quotes, yields, bids, offers, quoted market prices, and reference data. The fair values of OTC derivative contracts, including forward, swap, and option contracts related to interest rates, foreign currencies, credit standing of reference entities, equity prices, or commodity prices, may be based on market quotations or may be modeled using a series of techniques, including simulation models, depending on the contract and the terms of the transaction. The fair values of asset-backed securities and mortgage-backed securities are estimated based on models that consider the estimated cash flows of each debt tranche of the issuer, established benchmark yield, and estimated tranche-specific spread to the benchmark yield based on the unique attributes of the tranche, including, but not limited to, prepayment speed assumptions and attributes of the collateral.

Investments for which market quotations are not available or for which a pricing service or vendor does not provide a value, or for which such market quotations or values are considered by the investment adviser or subadviser to be unreliable (including, for example, certain foreign securities, thinly-traded securities, certain restricted securities, certain initial public offerings, or securities whose values may have been affected by a significant event) are stated at fair valuations determined in good faith by the Funds’ Valuation Committee¹ in accordance with procedures approved annually by the Trustees, and under the general oversight of the Trustees. The Funds’ Valuation Committee employs various methods to determine fair valuations including a regular review of significant inputs and assumptions and review of any related market activity. The Funds’ Valuation Committee reports to the Trustees at its regularly scheduled meetings. It is possible that fair value prices will be used by the Funds to a significant extent. The value determined for an investment using the Funds’ fair value procedures may differ from recent market prices for the investment and may be significantly different from the value realized upon the sale of such investment.

The Funds and certain Underlying Funds or Master Funds may invest in securities that are traded principally in foreign markets and that trade on weekends and other days when the Funds do not price their shares. As a result, the values of the Funds’ portfolio securities may change on days when the prices of the Funds’ shares are not calculated. The prices of the Funds’ shares will reflect any such changes when the prices of the Funds’ shares are next calculated, which is the next business day. The Funds may use fair value pricing more frequently for securities primarily traded in foreign markets because, among other things, most foreign markets close well before the Funds value their securities. The earlier close of these foreign markets gives rise to the possibility that significant events, including broad market moves, may have occurred in the interim. The Funds’ investments may be priced based on fair values provided by a third-party vendor, based on certain factors and methodologies applied by such vendor, in the event that there is movement in the U.S. market, between the close of the foreign market and the time the Funds calculate their net asset values. All assets and liabilities expressed in foreign currencies are converted into U.S. dollars at the mean between the buying and selling rates of such currencies against the U.S. dollar at the end of each business day.

For investments in equity or debt issued by privately held companies or funds (“Private Company” or collectively, the “Private Companies”) and any other investments fair valued using significant unobservable inputs, as described below, the fair valuation approaches used by third party service providers and/or the Funds’ subadvisers utilize one or a combination of, but not limited to, the following inputs:

¹ The voting members of the Valuation Committee consist of the President, Treasurer, Assistant Treasurers, Vice Presidents (except for the CCO, Secretary, and Assistant Secretaries) of the Trust, as well as such other members as the Trustees may from time to time designate. The non-voting members of the Valuation Committee consist of the CCO, Secretary, and Assistant Secretaries. The Valuation Committee reviews and determines the fair valuation of portfolio securities and the Funds’ pricing procedures in general.

Notes to Portfolio of Investments (Unaudited) (Continued)

Market approach: (i) recent market transactions, including subsequent rounds of financing, in the underlying investment or comparable issuers; (ii) recapitalizations and other transactions across the capital structure; and (iii) market multiples of comparable issuers.

Income approach: (i) future cash flows discounted to present value and adjusted as appropriate for liquidity, credit, and/or market risks; (ii) quoted prices for similar investments or assets in active markets; and (iii) other risk factors, such as interest rates, yield curves, volatilities, prepayment speeds, loss severities, credit risks, recovery rates, liquidation amounts, and/or default rates.

Cost approach: (i) audited or unaudited financial statements, investor communications, and financial or operational metrics issued by the Private Company; (ii) changes in the valuation of relevant indexes or publicly traded companies comparable to the Private Company; (iii) relevant news and other public sources; and (iv) known secondary market transactions in the Private Company’s interests and merger or acquisition activity in companies comparable to the Private Company.

Investments in series of preferred stock issued by Private Companies are typically valued utilizing Market approach in determining the enterprise value of the company. Such investments often contain rights and preferences that differ from other series of preferred and common stock of the same issuer. Valuation techniques such as the current value method (“CVM”), an option pricing model (“OPM”), a probability weighted expected return model (“PWERM”), or a hybrid of those techniques are used in allocating enterprise value of the company, as deemed appropriate under the circumstances. The CVM allocates value among the various parts of a company’s capital structure assuming that the value of convertible preferred stock is represented by the most favorable claim the preferred stockholders have on the enterprise value as of the valuation date. The use of OPM and PWERM techniques involve a determination of the exit scenarios of the investment in order to appropriately allocate the enterprise value of the company among the various parts of its capital structure.

The Private Companies are not subject to the public company disclosure, timing, and reporting standards as other investments held by a Fund. Typically, the most recently available information for a Private Company is as of a date that is earlier than the date a Fund is calculating its net asset value. This factor may result in a difference between the value of the investment and the price a Fund could receive upon the sale of the investment.

Fair value is defined as the price that would be received to sell an asset or paid to transfer a liability in an orderly transaction between market participants at the measurement date. A three-tier hierarchy is utilized to maximize the use of observable market data inputs and minimize the use of unobservable inputs and to establish classification of fair value measurements for disclosure purposes. Inputs refer broadly to the assumptions that market participants would use in pricing the asset or liability, including assumptions about risk, for example, the risk inherent in a particular valuation technique used to measure fair value (such as a pricing model) and/or the risk inherent in the inputs to the valuation technique. Inputs may be observable or unobservable. Observable inputs are inputs that reflect the assumptions market participants would use in pricing the asset or liability and are developed based on market data obtained from sources independent of the reporting entity. Unobservable inputs are inputs that reflect the reporting entity’s own assumptions about the assumptions market participants would use in pricing the asset or liability and are developed based on the best information available in the circumstances. The three-tier hierarchy of inputs is summarized in the three broad Levels listed below. The inputs or methodology used for valuing investments are not necessarily an indication of the risk associated with investing in those investments and the determination of the significance of a particular input to the fair value measurement in its entirety requires judgment and consideration of factors specific to each security.

Notes to Portfolio of Investments (Unaudited) (Continued)

Level 1 – quoted prices (unadjusted) in active markets for identical investments that the Funds can access at the measurement date

Level 2 – other significant observable inputs (including quoted prices for similar investments, interest rates, prepayment speeds, credit risk, etc.)

Level 3 – significant unobservable inputs, to the extent observable inputs are not available (including the Funds’ own assumptions in determining the fair value of investments)

The availability of observable inputs can vary from security to security and is affected by a wide variety of factors, including, for example, the type of security, whether the security is new and not yet established in the marketplace, the liquidity of markets, and other characteristics particular to the security. To the extent that valuation is based on models or inputs that are less observable or unobservable in the market, the determination of fair value requires more judgment. Accordingly, the degree of judgment exercised in determining fair value is greatest for instruments categorized in Level 3.

Changes in valuation techniques may result in transfers in or out of an investment’s assigned Level within the hierarchy. In addition, in periods of market dislocation, the observability of prices and inputs may be reduced for many instruments. This condition, as well as changes related to liquidity of investments, could cause a security to be reclassified between Levels.

In certain cases, the inputs used to measure fair value may fall into different Levels of the fair value hierarchy. In such cases, for disclosure purposes, the Level in the fair value hierarchy within which the fair value measurement falls is determined based on the lowest Level input that is significant to the overall fair value measurement.

The Allocation Funds, Feeder Funds, and MML American Funds Core Allocation Fund characterized all investments at Level 1, as of March 31, 2022. The Focused Equity Fund, Fundamental Value Fund, Income & Growth Fund, Large Cap Growth Fund, Small Cap Growth Equity Fund, and Small/Mid Cap Value Fund characterized all long-term investments at Level 1, and all short-term investments at Level 2, as of March 31, 2022. For each Fund noted in the preceding sentences, the level classification by major category of investments is the same as the category presentation in the Portfolio of Investments.

The following is the aggregate value by input level, as of March 31, 2022, for the remaining Funds’ investments:

Blue Chip Growth Fund

Asset Investments	Level 1		Level 2			Level 3		Total
Common Stock	$	544,413,508	$	6,404,998	*	$	1,239,185	$	552,057,691
Preferred Stock		—		—			857,256		857,256
Mutual Funds		1,288,549		—			—		1,288,549
Short-Term Investments		100		1,773,199			—		1,773,299
Total Investments	$	545,702,157	$	8,178,197		$	2,096,441	$	555,976,795

Equity Income Fund

Asset Investments	Level 1		Level 2			Level 3		Total
Common Stock	$	429,596,878	$	12,931,170	*	$	—	$	442,528,048
Preferred Stock		6,588,636		—			—		6,588,636
Mutual Funds		9,238,457		—			—		9,238,457
Short-Term Investments		1,075		5,600,854			—		5,601,929
Total Investments	$	445,425,046	$	18,532,024		$	—	$	463,957,070

Notes to Portfolio of Investments (Unaudited) (Continued)

Equity Index Fund

Asset Investments	Level 1		Level 2		Level 3		Total
Common Stock	$	728,865,263	$	—	$	—	$	728,865,263
Mutual Funds		—		385,515		—		385,515
Short-Term Investments		—		2,945,502		—		2,945,502
Total Investments	$	728,865,263	$	3,331,017	$	—	$	732,196,280
Asset Derivatives
Futures Contracts	$	201,882	$	—	$	—	$	201,882

Foreign Fund

Asset Investments	Level 1		Level 2		Level 3		Total
Common Stock*
Australia	$	826,575	$	8,217,837	$	—	$	9,044,412
Austria		—		681,824		—		681,824
Belgium		—		5,515,969		—		5,515,969
Canada		660,345		—		—		660,345
Cayman Islands		—		5,895,741		—		5,895,741
Finland		—		1,189,606		—		1,189,606
France		—		20,819,639		—		20,819,639
Germany		—		26,160,926		—		26,160,926
Ireland		2,140,181		6,538,097		—		8,678,278
Israel		2,461,028		—		—		2,461,028
Italy		—		1,062,782		—		1,062,782
Japan		—		50,335,285		—		50,335,285
Luxembourg		—		2,150,381		—		2,150,381
Netherlands		2,776,168		10,004,241		—		12,780,409
Norway		—		2,078,480		—		2,078,480
Republic of Korea		—		4,038,924		—		4,038,924
Singapore		—		3,536,051		—		3,536,051
Sweden		—		5,683,020		—		5,683,020
Switzerland		—		21,413,970		—		21,413,970
United Kingdom		919,805		34,127,331		—		35,047,136
Mutual Funds		4,049,203		—		—		4,049,203
Total Investments	$	13,833,305	$	209,450,104	$	—	$	223,283,409

Fundamental Equity Fund

Asset Investments	Level 1		Level 2			Level 3		Total
Common Stock	$	223,051,193	$	974,672	*	$	—	$	224,025,865
Short-Term Investments		—		648,435			—		648,435
Total Investments	$	223,051,193	$	1,623,107		$	—	$	224,674,300

Global Fund

Asset Investments	Level 1		Level 2		Level 3		Total
Common Stock*
Austria	$	—	$	557,589	$	—	$	557,589
Canada		6,657,791		—		—		6,657,791
Denmark		—		797,154		—		797,154
France		—		17,933,283		—		17,933,283
Germany		—		6,811,987		—		6,811,987
Ireland		14,480,568		3,286,535		—		17,767,103
Israel		1,594,000		—		—		1,594,000
Japan		—		3,092,511		—		3,092,511
Mexico		435,148		—		—		435,148
Netherlands		—		4,041,186		—		4,041,186
Republic of Korea		—		1,350,440		—		1,350,440
Spain		—		1,442,535		—		1,442,535
Sweden		—		2,044,240		—		2,044,240
Switzerland		—		14,471,384		—		14,471,384
United Kingdom		694,198		11,776,716		—		12,470,914
United States		72,093,851		—		—		72,093,851
Warrants		83,333		—		—		83,333
Short-Term Investments		—		515,689		—		515,689
Total Investments	$	96,038,889	$	68,121,249	$	—	$	164,160,138

Notes to Portfolio of Investments (Unaudited) (Continued)

Growth & Income Fund

Asset Investments	Level 1		Level 2			Level 3		Total
Common Stock	$	151,974,933	$	6,140,299	*	$	—	$	158,115,232
Short-Term Investments		—		835,080			—		835,080
Total Investments	$	151,974,933	$	6,975,379		$	—	$	158,950,312

International Equity Fund

Asset Investments	Level 1		Level 2			Level 3		Total
Common Stock*
Australia	$	—	$	1,702,343		$	—	$	1,702,343
Belgium		—		2,665,241			—		2,665,241
Canada		9,478,961		—			—		9,478,961
Cayman Islands		792,576		4,658,122			—		5,450,698
Denmark		—		4,972,236			—		4,972,236
Finland		—		489,459			—		489,459
France		—		40,468,350			—		40,468,350
Germany		—		37,004,201			—		37,004,201
Hong Kong		—		3,609,861			—		3,609,861
India		704,927		786,437			—		1,491,364
Indonesia		—		97,505			—		97,505
Ireland		2,462,185		2,045,908			—		4,508,093
Israel		2,067,955		—			—		2,067,955
Italy		—		5,386,278			—		5,386,278
Japan		—		24,390,662			—		24,390,662
Mexico		957,060		—			—		957,060
Netherlands		—		11,451,060			—		11,451,060
Portugal		—		1,147,825			—		1,147,825
Republic of Korea		—		1,273,684			—		1,273,684
Singapore		—		2,134,776			—		2,134,776
Spain		—		2,423,124			—		2,423,124
Sweden		—		3,362,530			—		3,362,530
Switzerland		—		32,363,128			—		32,363,128
Taiwan		2,522,049		—			—		2,522,049
United Kingdom		1,316,316		28,707,330			—		30,023,646
United States		991,518		—			—		991,518
Preferred Stock*
Republic of Korea		—		1,049,994			—		1,049,994
Mutual Funds		626,054		—			—		626,054
Short-Term Investments		—		3,242,074			—		3,242,074
Total Investments		21,919,601		215,432,128			—		237,351,729
Asset Derivatives
Forward Contracts	$	—	$	3,748		$	—	$	3,748
Liability Derivatives
Forward Contracts	$	—	$	(234	)	$	—	$	(234	)

Managed Volatility Fund

Asset Investments	Level 1			Level 2		Level 3			Total
Common Stock	$	130,517,868		$	—	$	—		$	130,517,868
Corporate Debt		—			—		—	+		—
Purchased Options		1,180,794			—		—			1,180,794
Short-Term Investments		—			1,802,855		—			1,802,855
Total Investments	$	131,698,662		$	1,802,855	$	—		$	133,501,517
Liability Derivatives
Written Options	$	(2,302,051	)	$	—	$	—		$	(2,302,051	)

Notes to Portfolio of Investments (Unaudited) (Continued)

Mid Cap Growth Fund

Asset Investments	Level 1		Level 2		Level 3		Total
Common Stock	$	373,284,774	$	5,171,949	$	981,778	$	379,438,501
Preferred Stock		—		—		1,719,646		1,719,646
Mutual Funds		—		4,007,609		—		4,007,609
Short-Term Investments		102		11,692,289		—		11,692,391
Total Investments	$	373,284,876	$	20,871,847	$	2,701,424	$	396,858,147

Mid Cap Value Fund

Asset Investments	Level 1		Level 2			Level 3		Total
Common Stock	$	312,401,290	$	21,970,200	*	$	—	$	334,371,490
Mutual Funds		6,996,188		2,365			—		6,998,553
Short-Term Investments		—		6,728,021			—		6,728,021
Total Investments	$	319,397,478	$	28,700,586		$	—	$	348,098,064
Asset Derivatives
Forward Contracts	$	—	$	68,129		$	—	$	68,129
Liability Derivatives
Forward Contracts	$	—	$	(49,624	)	$	—	$	(49,624	)

Small Company Value Fund

Asset Investments	Level 1		Level 2			Level 3		Total
Common Stock	$	90,022,878	$	721,746	*	$	—	$	90,744,624
Mutual Funds		2,055,914		—			—		2,055,914
Short-Term Investments		—		773,980			—		773,980
Total Investments	$	92,078,792	$	1,495,726		$	—	$	93,574,518
Asset Derivatives
Forward Contracts	$	—	$	47		$	—	$	47
Liability Derivatives
Forward Contracts	$	—	$	(3,333	)	$	—	$	(3,333	)

Total Return Bond Fund

Asset Investments	Level 1			Level 2			Level 3			Total
Common Stock	$	209,991		$	—		$	—		$	209,991
Bank Loans (Less Unfunded Loan Commitments)		—			6,776,949			—			6,776,949
Corporate Debt		—			102,487,987			—	+		102,487,987
Municipal Obligations		—			3,744,521			—			3,744,521
Non-U.S. Government Agency Obligations		—			51,667,401			—			51,667,401
Sovereign Debt Obligations		—			4,487,875			—			4,487,875
U.S. Government Agency Obligations and Instrumentalities		—			124,348,985			—			124,348,985
U.S. Treasury Obligations		—			107,634,267			—			107,634,267
Purchased Options		45,644			—			—			45,644
Rights		—			—			—	+		—
Mutual Funds		—			32,765,090			—			32,765,090
Short-Term Investments		—			27,278,421			—			27,278,421
Unfunded Loan Commitments**		—			462			—			462
Total Investments	$	255,635			461,191,958		$	—		$	461,447,593
Asset Derivatives
Futures Contracts		32,367			—			—			32,367
Swap Agreements		—			326,797			—			326,797
Total	$	32,367		$	326,797		$	—		$	359,164
Liability Derivatives
Swap Agreements		—			(1,242,017	)		—			(1,242,017	)
Written Options		(1,090,018	)		(298,300	)		—			(1,388,318	)
Total	$	(1,090,018	)	$	(1,540,317	)	$	—		$	(2,630,335	)

* Includes foreign equity securities whose values were adjusted to reflect market trading of comparable securities or other correlated instruments after the close of trading in their respective foreign markets, as applicable.

** Unfunded loan commitments are valued at the unrealized appreciation (depreciation) on the commitment.

+ Represents a security at $0 value as of March 31, 2022.

Notes to Portfolio of Investments (Unaudited) (Continued)

Derivative Instruments

Derivatives are financial instruments whose values are based on the values of one or more underlying indicators, such as a security, asset, currency, interest rate, or index. Derivative transactions can create investment leverage and may be highly volatile. Losses from derivatives can be substantially greater than the derivatives’ original cost and can sometimes be unlimited. A Fund may not be able to close out a derivative transaction at a favorable time or price.

Please refer to the Funds’ most recent Annual/Semiannual reports and Prospectus(es) and Statement(s) of Additional Information which can be found on the SEC’s EDGAR database on its website at http://www.sec.gov for more information regarding the derivatives and other investments held by the Fund(s).

Securities Lending

Each Fund, other than the Allocation Funds, Feeder Funds, and MML American Funds Core Allocation Fund, may lend its securities; however, lending cannot exceed 33% of the total assets of the Fund taken at current value. The Funds’ securities lending activities are governed by a Securities Lending Agency Agreement (“Lending Agreement”) between each Fund and the lending agent (“Agent”). The Lending Agreement authorizes the Agent to lend portfolio securities held by a Fund to approved borrowers (each, a “Borrower”).

Each Fund expects that in connection with any securities on loan, the loan will be secured continuously by collateral consisting of cash (U.S. currency) and/or securities (U.S. Treasury and Agency obligations) adjusted daily to have value at least equal to the current market value of the securities loaned. The market value of the loaned securities is determined at the close of business of a Fund and any additional collateral is delivered to the Fund the next business day. The Funds bear the risk of loss with respect to the investment of cash collateral. As with other extensions of credit, the Funds may bear the risk of delay in recovery of the loaned securities or even loss of rights in the collateral should the Borrower of the securities fail financially. Pursuant to the Lending Agreement, the Agent has provided indemnification to the Funds in the event of default by a Borrower with respect to a loan. The Funds receive compensation for lending their securities in the form of a securities loan fee paid by the Borrower, as well as a share of the income earned on investment of the cash collateral received for the loaned securities. At March 31, 2022, the Funds’ collateral was equal to or greater than 100% of the market value of securities on loan and all of the Funds' cash collateral was invested in the State Street Navigator Securities Lending Government Money Market Portfolio.

Subject to the terms of the Lending Agreement and the agreement between the Agent and the applicable Borrower (“Borrowing Agreement”), security loans can be terminated by the Agent, the Fund, or the Borrower and the related securities must be returned within the earlier of the customary settlement period for such securities or the period of time specified in the Borrowing Agreement.

Accounting for Investment Transactions

Investment transactions are accounted for on the trade date. Realized gains and losses on sales of investments and unrealized appreciation and depreciation of investments are computed by the specific identification cost method. Proceeds received from litigation, if any, are included in realized gains on investment transactions for any investments that are no longer held in the portfolio and as a reduction in cost for investments that continue to be held in the portfolio. Interest income, adjusted for amortization of discounts and premiums on debt securities, is earned from the settlement date and is recorded on the accrual basis. Dividend income is recorded on the ex-dividend date. Withholding taxes on foreign interest, dividends, and capital gains have been provided for in accordance with the applicable country’s tax rules and rates. Foreign dividend income is recorded on the ex-dividend date or as soon as practicable after the Fund determines the existence of a dividend declaration after exercising reasonable due diligence. Distributions received on securities that represent a return of capital or capital gains are recorded as a reduction of cost of investments and/or as a realized gain.

Notes to Portfolio of Investments (Unaudited) (Continued)

Foreign Currency Translation

The books and records of the Funds are maintained in U.S. dollars. The market values of foreign currencies, foreign securities, and other assets and liabilities denominated in foreign currencies are translated into U.S. dollars at the mean of the buying and selling rates of such currencies against the U.S. dollar at the end of each business day. Purchases and sales of foreign securities and income and expense items are translated at the rates of exchange prevailing on the respective dates of such transactions. The Funds do not isolate that portion of the results of operations arising from changes in the exchange rates from that portion arising from changes in the market prices of securities. Net realized foreign currency gains and losses resulting from changes in exchange rates include foreign currency gains and losses between trade date and settlement date on investment securities transactions, foreign currency transactions, and the difference between the amounts of dividends or interest recorded on the books of the Funds and the amount actually received.

Foreign Securities

The Global Fund invests a significant amount of its assets in foreign securities and each of the Foreign Fund and the International Equity Fund invests substantially all of its assets in foreign securities. The other Funds, except for the Allocation Funds, Feeder Funds, and MML American Funds Core Allocation Fund may also invest in foreign securities. In addition, certain Underlying Funds or Master Funds may invest in foreign securities. Foreign securities, including American Depositary Receipts, are subject to additional risks compared to securities of U.S. issuers, including international trade, currency, political, regulatory, and diplomatic risks. In addition, fluctuations in currency exchange rates may adversely affect the values of foreign securities and the price of a Fund’s shares. Emerging markets securities are subject to greater risks than securities issued in developed foreign markets, including less liquidity, greater price volatility, higher relative rates of inflation, greater political, economic, and social instability, greater custody and operational risks, and greater volatility in currency exchange rates.

Federal Income Tax

It is each Fund’s intent to continue to comply with the provisions of Subchapter M of the Code, applicable to a regulated investment company. Under such provisions, the Funds would not be subject to federal income taxes on their ordinary income and net realized capital gains to the extent they are distributed or deemed to have been distributed to their shareholders. Therefore, the Funds have not made any provision for federal income tax.

Dividends and Distributions to Shareholders

Dividends from net investment income and distributions of any net realized capital gains of each Fund are declared and paid annually and at other times as may be required to satisfy tax or regulatory requirements.

Distributions to shareholders are recorded on the ex-dividend date. Income and capital gain distributions are determined in accordance with income tax regulations, which may differ from generally accepted accounting principles. As a result, net investment income and net realized capital gains on investment transactions for a reporting period may differ significantly from distributions during such period.

3. Federal Income Tax Information

At March 31, 2022, the aggregate cost of investments and the unrealized appreciation (depreciation) in the value of all investments owned by the Fund(s), as computed on a federal income tax basis, were as follows:

Notes to Portfolio of Investments (Unaudited) (Continued)

	Federal Income Tax Cost		Tax Basis Unrealized Appreciation		Tax Basis Unrealized (Depreciation)		Net Unrealized Appreciation (Depreciation)
Conservative Allocation Fund	$	370,825,406	$	20,785,001	$	(17,669,278)	$	3,115,723
Balanced Allocation Fund		418,025,330		29,507,393		(14,233,210)		15,274,183
Moderate Allocation Fund		1,512,919,215		108,947,277		(40,272,051)		68,675,226
Growth Allocation Fund		1,037,330,968		96,050,362		(24,438,942)		71,611,420
Aggressive Allocation Fund		145,398,303		12,164,316		(3,510,254)		8,654,062
MML American Funds Growth Fund		187,177,515		76,337,389		–		76,337,389
MML American Funds International Fund		51,143,718		2,845,579		–		2,845,579
MML American Funds Core Allocation Fund		743,177,057		125,659,112		(14,342,071)		111,317,041
Blue Chip Growth Fund		365,509,300		222,146,412		(31,678,917)		190,467,495
Equity Income Fund		340,133,920		128,132,533		(4,309,383)		123,823,150
Equity Index Fund		356,776,463		387,607,693		(12,187,876)		375,419,817
Focused Equity Fund		285,333,800		62,448,451		(2,667,591)		59,780,860
Foreign Fund		211,664,190		31,397,757		(19,778,538)		11,619,219
Fundamental Equity Fund		192,671,543		39,748,312		(7,745,555)		32,002,757
Fundamental Value Fund		144,872,956		49,711,034		(2,335,156)		47,375,878
Global Fund		126,506,471		41,031,001		(3,377,334)		37,653,667
Growth & Income Fund		87,480,451		72,518,495		(1,048,634)		71,469,861
Income & Growth Fund		188,292,078		63,579,477		(3,700,552)		59,878,925
International Equity Fund		246,294,729		13,390,171		(22,333,171)		(8,943,000)
Large Cap Growth Fund		162,790,318		49,090,824		(3,486,730)		45,604,094
Managed Volatility Fund		75,009,763		61,575,770		(3,084,016)		58,491,754
Mid Cap Growth Fund		315,054,179		105,591,036		(23,787,068)		81,803,968
Mid Cap Value Fund		304,776,150		49,146,101		(5,824,187)		43,321,914
Small Cap Growth Equity Fund		162,142,719		28,885,636		(11,572,494)		17,313,142
Small Company Value Fund		97,086,007		4,123,682		(7,635,171)		(3,511,489)
Small/Mid Cap Value Fund		139,307,124		30,241,690		(5,490,430)		24,751,260
Total Return Bond Fund		471,951,983		2,062,202		(12,566,592)		(10,504,390)

4. New Accounting Pronouncements

In January 2021, FASB issued Accounting Standards Update 2021-01 — Reference Rate Reform (Topic 848) — Scope (“ASU 2021-01”) as an update to Accounting Standards Update 2020-04 — Reference Rate Reform (Topic 848) — Facilitation of the Effects of Reference Rate Reform on Financial Reporting (“ASU 2020-04”) in order to make certain clarifications. ASU 2020-04 and ASU 2021-01 provide optional temporary financial reporting relief from the effect of certain types of contract modifications due to the planned discontinuation of the London Interbank Offered Rate (“LIBOR”) and other interbank-offered based reference rates. The temporary relief provided by ASU 2020-04 and ASU 2021-01 is effective for certain reference rate-related contract modifications that occur during the period March 12, 2020 through December 31, 2022. Management does not believe the impact of adopting ASU 2020-04 and ASU 2021-01 will have a material impact on the financial statements.

Notes to Portfolio of Investments (Unaudited) (Continued)

5. Coronavirus (COVID-19) Pandemic

The global pandemic outbreak of the novel coronavirus known as COVID-19 and efforts to contain its spread have produced, and will likely continue to produce, substantial market volatility, severe market dislocations and liquidity constraints in many markets, exchange trading suspensions and closures, higher default rates, and global business disruption, and they may result in future significant adverse effects, such as declines in global financial markets and a substantial economic downturn or recession throughout the world. Such factors may have a significant adverse effect on a Fund’s performance and have the potential to impair the ability of a Fund’s investment adviser, subadviser, or other service providers to serve the Fund and could lead to disruptions that negatively impact the Fund.

6. Russian-Ukraine War

In February 2022, Russia commenced a large-scale military attack on Ukraine. The outbreak of hostilities between the two countries could result in more widespread conflict and could have a severe adverse effect on the regional and the global financial markets and economies. One or more of the Underlying Funds hold positions in securities or other instruments that are economically tied to Russia. Investments in Russia are subject to political, economic, legal, market, and currency risks, as well as the risks related to the economic sanctions on Russia, Russian individuals, and Russian corporate and banking entities imposed by the United States and/or other countries. Such sanctions — which affect companies in many sectors, including energy, financial services, and defense, among others — could adversely affect the global energy and financial markets and, thus, could affect the value of a Fund’s investments, even beyond any direct exposure the Fund may have to Russian issuers or the adjoining geographic regions. Such sanctions may also result in a decline in the value and liquidity of Russian securities and a weakening of the ruble, and will impair a Fund’s ability to buy, sell, receive, or deliver Russian securities. In addition, certain transactions have been or may be prohibited and/or existing investments have or may become illiquid (e.g., because transacting in certain existing investments is prohibited), which could cause a Fund to sell other portfolio holdings at a disadvantageous time or price in order to meet shareholder redemptions.

7. Subsequent Events

Effective April 29, 2022, the MML Growth & Income Fund changed its name to the MML Sustainable Equity Fund.