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Represents the number of directors of reporting entity.

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Number of Collaboration Agreements

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Related Party Transaction, Potential Required Amounts of Transaction

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Royalty Expense, Percent of Cost of Goods Sold

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Royalty Expense, Percent of Sales

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Royalty Expense, Sales Revenue Benchmark

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Shareholder Ownership Percentage

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Shareholder Ownership Percentage of Related Party in Affiliate

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Carrying value as of the balance sheet date of obligations incurred (and for which invoices have typically been received) and payable to vendors for goods and services received that are used in an entity's business. Used to reflect the current portion of the liabilities (due within one year or within the normal operating cycle if longer).

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Number of securities into which the class of warrant or right may be converted. For example, but not limited to, 500,000 warrants may be converted into 1,000,000 shares.

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Face (par) amount of debt instrument at time of issuance.

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Carrying amount as of the balance sheet date of obligations due all related parties. For classified balance sheets, represents the current portion of such liabilities (due within one year or within the normal operating cycle if longer).

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Total number of nonredeemable preferred shares (or preferred stock redeemable solely at the option of the issuer) issued to shareholders (includes related preferred shares that were issued, repurchased, and remain in the treasury). May be all or portion of the number of preferred shares authorized. Excludes preferred shares that are classified as debt.

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Amount of transactions with related party during the financial reporting period.

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Expenses recognized resulting from transactions (excluding transactions that are eliminated in consolidated or combined financial statements) with related party.

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Line items represent financial concepts included in a table. These concepts are used to disclose reportable information associated with domain members defined in one or many axes to the table.

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The aggregate costs incurred (1) in a planned search or critical investigation aimed at discovery of new knowledge with the hope that such knowledge will be useful in developing a new product or service, a new process or technique, or in bringing about a significant improvement to an existing product or process; or (2) to translate research findings or other knowledge into a plan or design for a new product or process or for a significant improvement to an existing product or process whether intended for sale or the entity's use, during the reporting period charged to research and development projects, including the costs of developing computer software up to the point in time of achieving technological feasibility, and costs allocated in accounting for a business combination to in-process projects deemed to have no alternative future use.

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The number of shares issued or sold by the subsidiary or equity method investee per stock transaction.

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Related Party Transactions - (Details)		1 Months Ended		3 Months Ended		12 Months Ended
Related Party Transactions - (Details)	Jan. 06, 2018 shares	Dec. 31, 2017 USD ($) shares	Jan. 31, 2016 USD ($)	Mar. 31, 2018 USD ($)	Jun. 30, 2017 USD ($)	Dec. 31, 2018 USD ($) director agreement shares	Dec. 31, 2017 USD ($) shares	Aug. 31, 2018 USD ($)	Mar. 07, 2017 shares	Sep. 30, 2016 USD ($) shares
Related Party Transaction [Line Items]
Number of collaboration agreements \| agreement						2
Research and development expenses						$ 6,018,000	$ 6,512,000
Related party payable		$ 2,303,000				$ 100,000	$ 2,303,000
Preferred shares issued (in shares) \| shares		8,000				8,000	8,000
Common stock, shares issued (shares) \| shares	409,091
2017 Private Placement
Related Party Transaction [Line Items]
Preferred shares issued (in shares) \| shares									1
Issued with September 2016 Convertible Notes
Related Party Transaction [Line Items]
Warrants issued to purchase common stock (in shares) \| shares										1,205,840
Convertible Debt
Related Party Transaction [Line Items]
Convertible debt principal										$ 18,087,500
Technology Access Fee
Related Party Transaction [Line Items]
Payments to Intrexon			$ 10,000,000
Development Milestone
Related Party Transaction [Line Items]
Possible required payments to Intrexon						$ 30,000,000
Commercialization Milestone
Related Party Transaction [Line Items]
Possible required payments to Intrexon						22,500,000
Milestones
Related Party Transaction [Line Items]
Payments to Intrexon						0
Affiliated Entity
Related Party Transaction [Line Items]
Research and development expenses - related party						500,000	$ 5,200,000
Trade payables		$ 2,300,000				100,000	2,300,000
Research and development expenses						$ (34,000)	$ 5,720,000
Ownership percent of related party						50.00%
Ownership percent of company common stock						17.00%
Cash royalty, net sales benchmark						$ 25,000,000
Royalty expense, percent of cost of goods sold (percentage)						33.33%
Payments to Intrexon						$ 0
Affiliated Entity \| Minimum
Related Party Transaction [Line Items]
Cash royalty, percent of sales						7.00%
Affiliated Entity \| Maximum
Related Party Transaction [Line Items]
Cash royalty, percent of sales						14.00%
Affiliated Entity \| 2017 Private Placement
Related Party Transaction [Line Items]
Warrants issued to purchase common stock (in shares) \| shares		545,456					545,456		259,176
Preferred shares issued (in shares) \| shares									3,016
Common stock, shares issued (shares) \| shares		545,456
Affiliated Entity \| Issued with September 2016 Convertible Notes
Related Party Transaction [Line Items]
Warrants issued to purchase common stock (in shares) \| shares										450,835
Affiliated Entity \| Convertible Debt
Related Party Transaction [Line Items]
Convertible debt principal										$ 6,762,500
Affiliated Entity \| Direct Expenses for Work Performed
Related Party Transaction [Line Items]
Research and development expenses - related party						$ 100,000	$ 1,300,000
Affiliated Entity \| Pass-through Costs
Related Party Transaction [Line Items]
Research and development expenses - related party						400,000	3,900,000
Affiliated Entity \| Reduction For Settlement Of Disputed Charges
Related Party Transaction [Line Items]
Research and development expenses - related party						$ 500,000	500,000
Affiliated Entity \| 2012 Exclusive Channel Collaboration Agreement Dispute
Related Party Transaction [Line Items]
Research and development expenses				$ 1,400,000	$ 1,100,000
Related party payable		$ 700,000			$ 200,000		$ 700,000	$ 200,000
Reduction in due to related parties				$ 500,000
Director
Related Party Transaction [Line Items]
Number of directors \| director						2