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Related Party Transactions (Details)	3 Months Ended
Related Party Transactions (Details)	Mar. 31, 2017 USD ($) director agreement	Mar. 31, 2016 USD ($)	Mar. 07, 2017 shares	Dec. 31, 2016 USD ($)	Sep. 30, 2016 USD ($) shares
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]
Number of exclusive channel collaboration agreements (agreement) \| agreement	2
Royalty and milestone payments incurred	$ 0
Warrants Issued With September 2016 Convertible Notes
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]
Warrants issued to purchase common stock (in shares) \| shares					6,029,174
Convertible Debt
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]
Convertible debt principal					$ 18,087,500
Affiliated Entity
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]
Outstanding payables to Intrexon	$ 1,500,000			$ 900,000
Shareholder ownership percentage of related party in affiliate, more than	50.00%
Shareholder ownership percentage	38.00%
Affiliated Entity \| Private Placement
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]
Warrants issued to purchase common stock (in shares) \| shares			1,295,875
Preferred stock, shares issued (shares) \| shares			3,016
Affiliated Entity \| Warrants Issued With September 2016 Convertible Notes
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]
Warrants issued to purchase common stock (in shares) \| shares					2,254,168
Affiliated Entity \| Convertible Debt
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]
Convertible debt principal					$ 6,762,500
Director
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]
Number of directors (director) \| director	2
2012 ECC \| Affiliated Entity
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]
Expenses for work performed	$ 1,500,000	$ 1,300,000
2015 ECC \| Affiliated Entity
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]
Expenses for work performed	0	0
Direct Expenses for Work Performed \| Affiliated Entity
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]
Expenses for work performed	300,000	500,000
Pass-through Costs \| Affiliated Entity
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]
Expenses for work performed	$ 1,200,000	$ 800,000