Represents the number of directors of reporting entity.

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Number of Collaboration Agreements

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Royalty Expense, Percent of Sales

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Royalty Expense, Sales Revenue Benchmark

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- Definition

Shareholder Ownership Percentage

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- Definition

Carrying value as of the balance sheet date of obligations incurred (and for which invoices have typically been received) and payable to vendors for goods and services received that are used in an entity's business. Used to reflect the current portion of the liabilities (due within one year or within the normal operating cycle if longer).

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-Name Accounting Standards Codification
-Glossary Current Liabilities
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- Definition

Number of securities into which the class of warrant or right may be converted. For example, but not limited to, 500,000 warrants may be converted into 1,000,000 shares.

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- Definition

Face (par) amount of debt instrument at time of issuance.

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- Definition

Total number of nonredeemable preferred shares (or preferred stock redeemable solely at the option of the issuer) issued to shareholders (includes related preferred shares that were issued, repurchased, and remain in the treasury). May be all or portion of the number of preferred shares authorized. Excludes preferred shares that are classified as debt.

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- Definition

Amount of transactions with related party during the financial reporting period.

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- Definition

Expenses recognized resulting from transactions (excluding transactions that are eliminated in consolidated or combined financial statements) with related party.

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Line items represent financial concepts included in a table. These concepts are used to disclose reportable information associated with domain members defined in one or many axes to the table.

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Related Party Transactions (Details)	1 Months Ended	12 Months Ended
Related Party Transactions (Details)	Jan. 31, 2016 USD ($)	Dec. 31, 2016 USD ($) director agreement shares	Dec. 31, 2015 USD ($) shares	Mar. 07, 2017 shares	Sep. 30, 2016 USD ($) shares
Related Party Transaction [Line Items]
Number of collaboration agreements \| agreement		2
Preferred shares issued (in shares) \| shares		0	0
2017 Private Placement \| Subsequent Event
Related Party Transaction [Line Items]
Preferred shares issued (in shares) \| shares				1
Issued with September 2016 Convertible Notes
Related Party Transaction [Line Items]
Warrants issued to purchase common stock \| shares					18,087,500
Convertible Debt
Related Party Transaction [Line Items]
Convertible debt principal					$ 18,087,500
Technology Access Fee
Related Party Transaction [Line Items]
Payments to Intrexon	$ 10,000,000
Development Milestone
Related Party Transaction [Line Items]
Possible required payments to Intrexon		$ 30,000,000
Commercialization Milestone
Related Party Transaction [Line Items]
Possible required payments to Intrexon		22,500,000
Milestones
Related Party Transaction [Line Items]
Payments to Intrexon		0
Affiliated Entity
Related Party Transaction [Line Items]
Research and development expenses - related party		3,700,000	$ 15,900,000
Trade payables		$ 900,000	10,700,000
Ownership percent of related party		50.00%
Ownership percent of company common stock		38.00%
Cash royalty, net sales benchmark		$ 100,000,000
Payments to Intrexon		$ 0
Affiliated Entity \| Minimum
Related Party Transaction [Line Items]
Cash royalty percent of sales		7.00%
Affiliated Entity \| Maximum
Related Party Transaction [Line Items]
Cash royalty percent of sales		14.00%
Affiliated Entity \| 2017 Private Placement \| Subsequent Event
Related Party Transaction [Line Items]
Warrants issued to purchase common stock \| shares				3,887,624
Preferred shares issued (in shares) \| shares				3,016
Affiliated Entity \| Issued with September 2016 Convertible Notes
Related Party Transaction [Line Items]
Warrants issued to purchase common stock \| shares					6,762,500
Affiliated Entity \| Convertible Debt
Related Party Transaction [Line Items]
Convertible debt principal					$ 6,762,500
Affiliated Entity \| Technology Access Fee
Related Party Transaction [Line Items]
Research and development expenses - related party			10,000,000
Affiliated Entity \| Direct Expenses for Work Performed
Related Party Transaction [Line Items]
Research and development expenses - related party		$ 1,200,000	3,100,000
Affiliated Entity \| Pass-through Costs
Related Party Transaction [Line Items]
Research and development expenses - related party		$ 2,500,000	$ 2,800,000
Director
Related Party Transaction [Line Items]
Number of directors \| director		2