Related Party Transactions (Details)	1 Months Ended	3 Months Ended		6 Months Ended
	Jan. 31, 2016 USD ($)	Jun. 30, 2016 USD ($) agreement	Jun. 30, 2015 USD ($)	Jun. 30, 2016 USD ($) director agreement	Jun. 30, 2015 USD ($)	Dec. 31, 2015 USD ($)
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]
Number of exclusive channel collaboration agreements (agreement) | agreement		2		2
Technology Access Fee
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]
Fees to Intrexon	$ 10,000,000
Affiliated Entity
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]
Expenses for work performed		$ 900,000	$ 1,100,000	$ 2,200,000	$ 2,900,000
Outstanding payables to Intrexon		$ 900,000		$ 900,000		$ 10,700,000
Shareholder ownership percentage of related party in affiliate, more than		50.00%		50.00%
Shareholder ownership percentage		38.00%		38.00%
Affiliated Entity | 2015 ECC
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]
Expenses for work performed				$ 0	0
Affiliated Entity | Direct Expenses for Work Performed
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]
Expenses for work performed		$ 300,000	800,000	800,000	1,700,000
Affiliated Entity | Pass-through Costs
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]
Expenses for work performed		$ 600,000	$ 300,000	$ 1,400,000	$ 1,200,000
Director
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]
Number of directors (director) | director				2