Fidelity (logo) Investments®||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||| Fidelity Management & Research Company
245 Summer Street
Boston MA 02210
617 563 7000

|||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||| April 2, 2013

U.S. Securities & Exchange Commission

100 F Street, N.E.

Washington, D.C. 20549

Attention: File Room

RE:	Fidelity Devonshire Trust (the trust):
	Fidelity Advisor Series Equity-Income Fund
	Fidelity Advisor Series Stock Selector Large Cap Value Fund
	Fidelity Equity-Income Fund
	Fidelity Large Cap Growth Fund
	Fidelity Mid Cap Value Fund
	Fidelity Series All-Sector Equity Fund
	Fidelity Series Equity-Income Fund
	Fidelity Series Large Cap Value Fund
	Fidelity Series Stock Selector Large Cap Value Fund
	Fidelity Stock Selector Large Cap Value Fund (the funds)
	File Nos. (002-24389) and (811-01352)

__________________________________________________________________________

Ladies and Gentlemen:

Transmitted herewith pursuant to Rule 497(j) under the Securities Act of 1933 is certification that the Prospectuses and Statements of Additional Information with respect to the above referenced funds do not differ from those filed in the most recent post-effective amendment, which was filed electronically.

||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||	Very truly yours,
	/s/ Scottt C. Goebel
	Scott C. Goebel
	Secretary of the Trust