Fidelity (logo) Investments®||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||| Fidelity Management & Research Company
82 Devonshire Street
Boston MA 02109-3614
617 563 7000

|||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||| April 5, 2010

U.S. Securities & Exchange Commission

100 F Street, N.E.

Washington, D.C. 20549

Attention: File Room

RE:	Fidelity Devonshire Trust (the trust):
	Fidelity Equity-Income Fund Fidelity Large Cap Growth Fund Fidelity Large Cap Value Fund Fidelity Mid Cap Growth Fund Fidelity Mid Cap Value Fund Fidelity Series All-Sector Equity Fund Fidelity Series Large Cap Value Fund Fidelity Telecom and Utilities Fund (the funds)
	File Nos. (002-24389) and (811-01352)

__________________________________________________________________________

Ladies and Gentlemen:

Transmitted herewith pursuant to Rule 497(j) under the Securities Act of 1933 is certification that the Prospectuses and Statements of Additional Information with respect to the above referenced funds do not differ from those filed in the most recent post-effective amendment, which was filed electronically.

||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||	Very truly yours,
	/s/ Scott C. Goebel
	Scott C. Goebel
	Secretary of the Trust