The Securities and Exchange Commission has not necessarily reviewed the information in this filing and has not determined if it is accurate and complete.
The reader should not assume that the information is accurate and complete.

UNITED STATES SECURITIES AND EXCHANGE COMMISSION
Washington, D.C. 20549
FORM 13F

FORM 13F INFORMATION TABLE

OMB APPROVAL
OMB Number: 3235-0006
Estimated average burden
hours per response: 23.8

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COLUMN 1 COLUMN 2 COLUMN 3 COLUMN 4 COLUMN 5 COLUMN 6 COLUMN 7 COLUMN 8
VALUE SHRS OR SH/ PUT/ INVESTMENT OTHER VOTING AUTHORITY
NAME OF ISSUER TITLE OF CLASS CUSIP (x$1000) PRN AMT PRN CALL DISCRETION MANAGER SOLE SHARED NONE
180 DEGREE CAP CORP FUND 68235B109 1 500 SH   DFND 2 0 0 500
1ST CONSTITUTION BANCORP COMMON 31986N102 34 1,489 SH   DFND 2 0 0 1,489
1ST SOURCE CORP COMMON 336901103 12,177 227,901 SH   DFND 4 206,806 0 21,095
1ST SOURCE CORP COMMON 336901103 683 12,792 SH   DFND 2 9,650 0 3,142
1ST SOURCE CORP COMMON 336901103 38 710 SH   DFND 5 710 0 0
1ST SOURCE CORP COMMON 336901103 64 1,192 SH   DFND 16 0 0 1,192
21VIANET GROUP INC ADR 90138A103 2,326 239,842 SH   DFND 5 239,842 0 0
22ND CENTY GROUP INC COMMON 90137F103 58 23,596 SH   DFND 2 23,596 0 0
22ND CENTY GROUP INC COMMON 90137F103 0 24 SH   DFND 5 24 0 0
2U INC COMMON 90214J101 2,239 26,800 SH   DFND 2 22,100 0 4,700
2U INC COMMON 90214J101 4,312 51,609 SH   DFND 5 51,609 0 0
2U INC COMMON 90214J101 28,739 343,927 SH   DFND 4 319,879 0 24,048
2U INC COMMON 90214J101 92 1,100 SH   DFND 5 0 0 1,100
2U INC COMMON 90214J101 6,632 79,365 SH   DFND 8 79,365 0 0
3-D SYS CORP DEL COMMON 88554D205 4 311 SH   DFND 6 311 0 0
3-D SYS CORP DEL COMMON 88554D205 40 2,922 SH   DFND 5 0 0 2,922
3-D SYS CORP DEL COMMON 88554D205 306 22,174 SH   DFND 2 22,174 0 0
3-D SYS CORP DEL COMMON 88554D205 1 43 SH   DFND 2 43 0 0
3M CO COMMON 88579Y101 41,329 210,092 SH   OTR 2 167,617 9,369 33,106
3M CO COMMON 88579Y101 1,334 6,780 SH   OTR 21 0 6,780 0
3M CO COMMON 88579Y101 6 33 SH   OTR 4 0 33 0
3M CO COMMON 88579Y101 1,666 8,468 SH   OTR 1 7,824 0 644
3M CO COMMON 88579Y101 450 2,288 SH   OTR 13 0 2,288 0
3M CO COMMON 88579Y101 65,253 331,704 SH   DFND 2 327,194 0 4,510
3M CO COMMON 88579Y101 341 1,731 SH   DFND 2 1,731 0 0
3M CO COMMON 88579Y101 509 2,588 SH   DFND 1 2,557 0 31
3M CO COMMON 88579Y101 2,538 12,901 SH   DFND 19 12,901 0 0
3M CO COMMON 88579Y101 795 4,040 SH   DFND 16 0 0 4,040
3M CO COMMON 88579Y101 22,904 116,430 SH   DFND 5 44,395 0 72,035
3M CO COMMON 88579Y101 185,534 943,139 SH   DFND 4 856,184 0 86,955
3M CO COMMON 88579Y101 28,052 142,601 SH   DFND 8 134,391 5,400 2,810
3M CO COMMON 88579Y101 727 3,695 SH   DFND 10 3,695 0 0
3M CO COMMON 88579Y101 26,302 133,704 SH   DFND 5 133,704 0 0
3M CO OPTION 88579Y901 21,934 111,500 SH Call DFND 5 111,500 0 0
3M CO OPTION 88579Y951 13,377 68,000 SH Put DFND 5 68,000 0 0
51JOB INC ADR 316827104 3,327 34,073 SH   OTR 18 34,073 0 0
51JOB INC ADR 316827104 14,597 149,498 SH   DFND 4 108,588 0 40,910
51JOB INC ADR 316827104 56,309 576,696 SH   DFND 8 522,511 54,185 0
51JOB INC ADR 316827104 10,398 106,497 SH   DFND 15 106,497 0 0
51JOB INC ADR 316827104 4,817 49,339 SH   DFND 10 3,690 0 45,649
51JOB INC ADR 316827104 4,948 50,673 SH   DFND 2 50,673 0 0
51JOB INC ADR 316827104 31,880 326,503 SH   DFND 5 326,503 0 0
58 COM INC ADR 31680Q104 532 7,677 SH   DFND 4 7,677 0 0
58 COM INC ADR 31680Q104 241 3,474 SH   DFND 6 3,474 0 0
58 COM INC ADR 31680Q104 3,750 54,077 SH   DFND 5 54,077 0 0
8X8 INC NEW COMMON 282914100 841 41,950 SH   DFND 2 41,950 0 0
8X8 INC NEW COMMON 282914100 1,052 52,473 SH   DFND 4 52,473 0 0
A10 NETWORKS INC COMMON 002121101 2 313 SH   OTR 1 0 0 313
A10 NETWORKS INC COMMON 002121101 36 5,740 SH   DFND 5 2,684 0 3,056
A10 NETWORKS INC COMMON 002121101 63 10,149 SH   DFND 2 10,149 0 0
A10 NETWORKS INC COMMON 002121101 1,366 219,288 SH   DFND 2 219,288 0 0
A10 NETWORKS INC COMMON 002121101 2,422 388,785 SH   DFND 5 388,785 0 0
AAC HLDGS INC COMMON 000307108 322 34,370 SH   DFND 5 34,370 0 0
AAC HLDGS INC COMMON 000307108 25 2,673 SH   DFND 2 2,673 0 0
AAON INC COMMON 000360206 279 8,381 SH   DFND 2 8,381 0 0
AAON INC COMMON 000360206 1,255 37,746 SH   DFND 5 37,746 0 0
AAR CORP COMMON 000361105 2,025 43,565 SH   DFND 5 43,565 0 0
AAR CORP COMMON 000361105 2,959 63,646 SH   DFND 2 60,146 0 3,500
AAR CORP COMMON 000361105 3,152 67,790 SH   DFND 4 44,490 0 23,300
AARONS INC COMMON 002535300 3,421 78,727 SH   DFND 4 75,868 0 2,859
AARONS INC COMMON 002535300 3,409 78,458 SH   DFND 2 78,458 0 0
AARONS INC COMMON 002535300 2,240 51,549 SH   DFND 5 51,549 0 0
ABAXIS INC COMMON 002567105 8 91 SH   OTR 21 0 91 0
ABAXIS INC COMMON 002567105 370 4,453 SH   DFND 2 4,453 0 0
ABAXIS INC COMMON 002567105 133 1,608 SH   DFND 8 1,608 0 0
ABAXIS INC COMMON 002567105 244 2,938 SH   DFND 5 2,938 0 0
ABAXIS INC COMMON 002567105 1,190 14,340 SH   DFND 4 14,340 0 0
ABAXIS INC COMMON 002567105 22 260 SH   DFND 5 0 0 260
ABB LTD ADR 000375204 378 17,348 SH   OTR 1 0 0 17,348
ABB LTD ADR 000375204 80 3,690 SH   DFND 1 0 0 3,690
ABB LTD ADR 000375204 212 9,755 SH   DFND 5 6,811 0 2,944
ABB LTD ADR 000375204 4,482 205,900 SH   DFND 4 205,900 0 0
ABB LTD ADR 000375204 959 44,060 SH   DFND 5 44,060 0 0
ABB LTD ADR 000375204 646 29,680 SH   DFND 2 0 0 29,680
ABBOTT LABS COMMON 002824100 3,538 58,005 SH   OTR 21 0 58,005 0
ABBOTT LABS COMMON 002824100 38,202 626,371 SH   OTR 2 412,661 133,705 80,005
ABBOTT LABS COMMON 002824100 790 12,959 SH   OTR 1 10,453 0 2,506
ABBOTT LABS COMMON 002824100 2,688 44,067 SH   OTR 13 0 44,067 0
ABBOTT LABS COMMON 002824100 6 102 SH   OTR 4 0 102 0
ABBOTT LABS COMMON 002824100 86 1,403 SH   OTR 11 0 1,403 0
ABBOTT LABS COMMON 002824100 46,504 762,488 SH   DFND 8 693,916 17,400 51,172
ABBOTT LABS COMMON 002824100 187,477 3,073,893 SH   DFND 2 3,043,665 0 30,228
ABBOTT LABS COMMON 002824100 1,999 32,781 SH   DFND 10 32,781 0 0
ABBOTT LABS COMMON 002824100 18,079 296,419 SH   DFND 11 296,419 0 0
ABBOTT LABS COMMON 002824100 10,534 172,709 SH   DFND 5 14,204 0 158,505
ABBOTT LABS COMMON 002824100 2,545 41,730 SH   DFND 15 41,730 0 0
ABBOTT LABS COMMON 002824100 299,988 4,918,650 SH   DFND 4 4,268,527 0 650,123
ABBOTT LABS COMMON 002824100 1,499 24,570 SH   DFND 16 0 0 24,570
ABBOTT LABS COMMON 002824100 1,505 24,684 SH   DFND 1 22,295 0 2,389
ABBOTT LABS COMMON 002824100 2,535 41,561 SH   DFND 19 41,561 0 0
ABBOTT LABS COMMON 002824100 28,560 468,276 SH   DFND 5 468,276 0 0
ABBOTT LABS COMMON 002824100 3,724 61,067 SH   DFND 2 61,067 0 0
ABBOTT LABS OPTION 002824900 1,269 20,800 SH Call DFND 5 20,800 0 0
ABBOTT LABS OPTION 002824950 2,031 33,300 SH Put DFND 5 33,300 0 0
ABBVIE INC COMMON 00287Y109 3 29 SH   OTR 13 0 29 0
ABBVIE INC COMMON 00287Y109 43,385 468,264 SH   OTR 2 378,807 0 89,457
ABBVIE INC COMMON 00287Y109 973 10,506 SH   OTR 18 10,506 0 0
ABBVIE INC COMMON 00287Y109 77 834 SH   OTR 10 834 0 0
ABBVIE INC COMMON 00287Y109 1,048 11,311 SH   OTR 1 9,545 0 1,766
ABBVIE INC COMMON 00287Y109 83,309 899,178 SH   DFND 2 881,064 0 18,114
ABBVIE INC COMMON 00287Y109 4,683 50,546 SH   DFND 2 50,546 0 0
ABBVIE INC COMMON 00287Y109 236,476 2,552,359 SH   DFND 4 2,134,609 0 417,750
ABBVIE INC COMMON 00287Y109 849 9,164 SH   DFND 1 7,800 0 1,364
ABBVIE INC COMMON 00287Y109 1 14 SH   DFND 11 14 0 0
ABBVIE INC COMMON 00287Y109 151,826 1,638,700 SH   DFND 8 1,351,394 80,193 207,113
ABBVIE INC COMMON 00287Y109 25,167 271,637 SH   DFND 5 19,562 0 252,075
ABBVIE INC COMMON 00287Y109 146 1,572 SH   DFND 19 1,572 0 0
ABBVIE INC COMMON 00287Y109 801 8,647 SH   DFND 17 0 8,647 0
ABBVIE INC COMMON 00287Y109 57 618 SH   DFND 6 618 0 0
ABBVIE INC COMMON 00287Y109 981 10,589 SH   DFND 16 1,689 0 8,900
ABBVIE INC COMMON 00287Y109 135,801 1,465,737 SH   DFND 5 1,465,737 0 0
ABBVIE INC COMMON 00287Y109 4,807 51,886 SH   DFND 15 51,886 0 0
ABBVIE INC COMMON 00287Y109 9,103 98,256 SH   DFND 10 98,256 0 0
ABBVIE INC OPTION 00287Y909 9,191 99,200 SH Call DFND 5 99,200 0 0
ABBVIE INC OPTION 00287Y959 13,406 144,700 SH Put DFND 5 144,700 0 0
ABEONA THERAPEUTICS INC COMMON 00289Y107 1,148 71,726 SH   DFND 5 71,726 0 0
ABEONA THERAPEUTICS INC COMMON 00289Y107 1,600 100,000 SH   DFND 4 82,300 0 17,700
ABEONA THERAPEUTICS INC COMMON 00289Y107 100 6,263 SH   DFND 2 6,263 0 0
ABERCROMBIE & FITCH CO COMMON 002896207 3,774 154,173 SH   DFND 4 119,873 0 34,300
ABERCROMBIE & FITCH CO COMMON 002896207 2,377 97,120 SH   DFND 2 97,120 0 0
ABERCROMBIE & FITCH CO COMMON 002896207 39 1,598 SH   DFND 6 1,598 0 0
ABERCROMBIE & FITCH CO COMMON 002896207 766 31,281 SH   DFND 8 31,281 0 0
ABERCROMBIE & FITCH CO COMMON 002896207 0 17 SH   DFND 5 0 0 17
ABERCROMBIE & FITCH CO COMMON 002896207 3,837 156,747 SH   DFND 5 156,747 0 0
ABIOMED INC COMMON 003654100 124,960 305,489 SH   DFND 4 304,325 0 1,164
ABIOMED INC COMMON 003654100 1,950 4,768 SH   DFND 8 4,768 0 0
ABIOMED INC COMMON 003654100 11,203 27,388 SH   DFND 2 27,388 0 0
ABIOMED INC COMMON 003654100 103 252 SH   DFND 5 25 0 227
ABIOMED INC COMMON 003654100 450 1,100 SH   DFND 2 1,100 0 0
ABIOMED INC COMMON 003654100 510 1,247 SH   DFND 5 1,247 0 0
ABM INDS INC COMMON 000957100 6,096 208,898 SH   DFND 4 165,898 0 43,000
ABM INDS INC COMMON 000957100 4 140 SH   DFND 5 0 0 140
ABM INDS INC COMMON 000957100 2,096 71,843 SH   DFND 2 71,543 0 300
ABM INDS INC COMMON 000957100 3 95 SH   DFND 1 0 0 95
ABRAXAS PETE CORP COMMON 003830106 7,814 2,703,916 SH   DFND 4 2,060,816 0 643,100
ABRAXAS PETE CORP COMMON 003830106 2,061 713,108 SH   DFND 2 713,108 0 0
ACACIA COMMUNICATIONS INC COMMON 00401C108 2,238 64,280 SH   DFND 4 64,280 0 0
ACACIA COMMUNICATIONS INC COMMON 00401C108 8,058 231,476 SH   DFND 2 231,476 0 0
ACACIA RESH CORP COMMON 003881307 18 4,250 SH   DFND 16 0 0 4,250
ACACIA RESH CORP COMMON 003881307 2,887 695,564 SH   DFND 4 613,264 0 82,300
ACACIA RESH CORP COMMON 003881307 29 6,883 SH   DFND 5 6,883 0 0
ACACIA RESH CORP COMMON 003881307 98 23,561 SH   DFND 2 23,561 0 0
ACADIA HEALTHCARE COMPANY IN COMMON 00404A109 51 1,250 SH   OTR 2 350 0 900
ACADIA HEALTHCARE COMPANY IN COMMON 00404A109 29 708 SH   OTR 1 0 0 708
ACADIA HEALTHCARE COMPANY IN COMMON 00404A109 179,872 4,396,774 SH   DFND 4 4,271,155 0 125,619
ACADIA HEALTHCARE COMPANY IN COMMON 00404A109 17,694 432,516 SH   DFND 2 431,564 0 952
ACADIA HEALTHCARE COMPANY IN COMMON 00404A109 40,808 997,498 SH   DFND 8 704,097 0 293,401
ACADIA HEALTHCARE COMPANY IN COMMON 00404A109 9 219 SH   DFND 1 0 0 219
ACADIA PHARMACEUTICALS INC COMMON 004225108 27 1,775 SH   DFND 16 0 0 1,775
ACADIA PHARMACEUTICALS INC COMMON 004225108 1,079 70,630 SH   DFND 8 70,630 0 0
ACADIA PHARMACEUTICALS INC COMMON 004225108 600 39,290 SH   DFND 2 39,290 0 0
ACADIA PHARMACEUTICALS INC COMMON 004225108 5,904 386,644 SH   DFND 4 360,603 0 26,041
ACADIA RLTY TR COMMON 004239109 448 16,365 SH   DFND 2 16,365 0 0
ACADIA RLTY TR COMMON 004239109 8,381 306,198 SH   DFND 5 306,198 0 0
ACADIA RLTY TR COMMON 004239109 147 5,387 SH   DFND 4 5,387 0 0
ACCELERATE DIAGNOSTICS INC COMMON 00430H102 727 32,585 SH   DFND 5 32,585 0 0
ACCELERATE DIAGNOSTICS INC COMMON 00430H102 118 5,293 SH   DFND 2 5,293 0 0
ACCELERATE DIAGNOSTICS INC OPTION 00430H902 3,345 150,000 SH Call DFND 5 150,000 0 0
ACCELERON PHARMA INC COMMON 00434H108 28 567 SH   OTR 1 0 0 567
ACCELERON PHARMA INC COMMON 00434H108 476 9,808 SH   DFND 2 7,858 0 1,950
ACCELERON PHARMA INC COMMON 00434H108 9 176 SH   DFND 1 0 0 176
ACCELERON PHARMA INC COMMON 00434H108 1,969 40,574 SH   DFND 4 33,574 0 7,000
ACCELERON PHARMA INC OPTION 00434H908 1,213 25,000 SH Call DFND 5 25,000 0 0
ACCENTURE PLC IRELAND COMMON G1151C101 3,546 21,677 SH   OTR 13 0 21,677 0
ACCENTURE PLC IRELAND COMMON G1151C101 3,907 23,884 SH   OTR 21 0 23,884 0
ACCENTURE PLC IRELAND COMMON G1151C101 24,384 149,058 SH   OTR 2 77,936 63,499 7,623
ACCENTURE PLC IRELAND COMMON G1151C101 48 291 SH   OTR 4 0 291 0
ACCENTURE PLC IRELAND COMMON G1151C101 1,132 6,922 SH   OTR 1 3,052 0 3,870
ACCENTURE PLC IRELAND COMMON G1151C101 7 44 SH   OTR 5 0 44 0
ACCENTURE PLC IRELAND COMMON G1151C101 48 294 SH   OTR 11 0 294 0
ACCENTURE PLC IRELAND COMMON G1151C101 18,202 111,266 SH   DFND 10 73,211 0 38,055
ACCENTURE PLC IRELAND COMMON G1151C101 153,742 939,799 SH   DFND 5 939,799 0 0
ACCENTURE PLC IRELAND COMMON G1151C101 315,002 1,925,556 SH   DFND 2 1,914,319 0 11,237
ACCENTURE PLC IRELAND COMMON G1151C101 209,305 1,279,450 SH   DFND 8 1,114,183 12,370 152,897
ACCENTURE PLC IRELAND COMMON G1151C101 17,729 108,371 SH   DFND 11 108,371 0 0
ACCENTURE PLC IRELAND COMMON G1151C101 2,043 12,490 SH   DFND 16 0 0 12,490
ACCENTURE PLC IRELAND COMMON G1151C101 4,653 28,445 SH   DFND 1 24,237 0 4,208
ACCENTURE PLC IRELAND COMMON G1151C101 817,396 4,996,614 SH   DFND 4 4,239,097 0 757,517
ACCENTURE PLC IRELAND COMMON G1151C101 15,228 93,086 SH   DFND 5 66,382 0 26,704
ACCENTURE PLC IRELAND COMMON G1151C101 4,259 26,037 SH   DFND 15 26,037 0 0
ACCENTURE PLC IRELAND COMMON G1151C101 603 3,689 SH   DFND 2 3,689 0 0
ACCENTURE PLC IRELAND COMMON G1151C101 4,190 25,615 SH   DFND 19 25,615 0 0
ACCENTURE PLC IRELAND COMMON G1151C101 212 1,293 SH   DFND 17 0 1,293 0
ACCENTURE PLC IRELAND OPTION G1151C901 9,390 57,400 SH Call DFND 5 57,400 0 0
ACCENTURE PLC IRELAND OPTION G1151C951 14,707 89,900 SH Put DFND 5 89,900 0 0
ACCESS NATL CORP COMMON 004337101 88 3,090 SH   DFND 2 0 0 3,090
ACCO BRANDS CORP COMMON 00081T108 10 752 SH   OTR 2 0 0 752
ACCO BRANDS CORP COMMON 00081T108 8,672 626,164 SH   DFND 2 626,047 0 117
ACCO BRANDS CORP COMMON 00081T108 66,518 4,802,723 SH   DFND 4 4,137,523 0 665,200
ACCO BRANDS CORP COMMON 00081T108 342 24,725 SH   DFND 16 0 0 24,725
ACCURAY INC COMMON 004397105 23 5,704 SH   DFND 4 5,704 0 0
ACCURAY INC COMMON 004397105 70 16,982 SH   DFND 2 16,982 0 0
ACELRX PHARMACEUTICALS INC COMMON 00444T100 47 13,781 SH   DFND 5 13,781 0 0
ACER THERAPEUTICS INC COMMON 00444P108 1 45 SH   DFND 5 0 0 45
ACETO CORP COMMON 004446100 495 147,688 SH   DFND 5 147,688 0 0
ACHAOGEN INC COMMON 004449104 90 10,361 SH   DFND 2 6,661 0 3,700
ACHILLION PHARMACEUTICALS IN COMMON 00448Q201 47 16,638 SH   DFND 16 0 0 16,638
ACHILLION PHARMACEUTICALS IN COMMON 00448Q201 2,620 925,918 SH   DFND 4 868,825 0 57,093
ACHILLION PHARMACEUTICALS IN COMMON 00448Q201 78 27,612 SH   DFND 2 27,612 0 0
ACI WORLDWIDE INC COMMON 004498101 263 10,678 SH   OTR 21 0 10,678 0
ACI WORLDWIDE INC COMMON 004498101 285 11,549 SH   OTR 2 0 11,549 0
ACI WORLDWIDE INC COMMON 004498101 52 2,100 SH   OTR 1 0 0 2,100
ACI WORLDWIDE INC COMMON 004498101 335 13,593 SH   OTR 13 0 13,593 0
ACI WORLDWIDE INC COMMON 004498101 2,502 101,411 SH   DFND 2 94,298 0 7,113
ACI WORLDWIDE INC COMMON 004498101 3,777 153,113 SH   DFND 5 153,113 0 0
ACI WORLDWIDE INC COMMON 004498101 51 2,064 SH   DFND 1 0 0 2,064
ACI WORLDWIDE INC COMMON 004498101 3,632 147,235 SH   DFND 4 141,635 0 5,600
ACI WORLDWIDE INC COMMON 004498101 41 1,655 SH   DFND 16 0 0 1,655
ACLARIS THERAPEUTICS INC COMMON 00461U105 1,967 98,500 SH   DFND 4 63,600 0 34,900
ACLARIS THERAPEUTICS INC COMMON 00461U105 1,708 85,529 SH   DFND 2 85,529 0 0
ACM RESEARCH INC COMMON 00108J109 18 1,648 SH   DFND 2 1,648 0 0
ACM RESEARCH INC COMMON 00108J109 176 16,348 SH   DFND 5 16,348 0 0
ACNB CORP COMMON 000868109 47 1,387 SH   DFND 2 0 0 1,387
ACORDA THERAPEUTICS INC COMMON 00484M106 571 19,897 SH   DFND 5 19,897 0 0
ACORDA THERAPEUTICS INC COMMON 00484M106 6,319 220,173 SH   DFND 4 187,873 0 32,300
ACORDA THERAPEUTICS INC COMMON 00484M106 2,102 73,251 SH   DFND 2 73,251 0 0
ACORDA THERAPEUTICS INC BOND 00484MAA4 1,750 1,795,000 PRN   DFND 4 1,795,000 0 0
ACORDA THERAPEUTICS INC BOND 00484MAA4 2,109 2,164,000 PRN   DFND 8 1,954,000 0 210,000
ACORDA THERAPEUTICS INC BOND 00484MAA4 82 84,000 PRN   DFND 5 84,000 0 0
ACORDA THERAPEUTICS INC BOND 00484MAA4 2,518 2,583,000 PRN   DFND 2 2,583,000 0 0
ACTIVISION BLIZZARD INC COMMON 00507V109 13,520 177,151 SH   OTR 21 0 177,151 0
ACTIVISION BLIZZARD INC COMMON 00507V109 29 380 SH   OTR 1 0 0 380
ACTIVISION BLIZZARD INC COMMON 00507V109 16,849 220,767 SH   OTR 2 0 220,767 0
ACTIVISION BLIZZARD INC COMMON 00507V109 7,652 100,266 SH   OTR 13 0 100,266 0
ACTIVISION BLIZZARD INC COMMON 00507V109 82 1,073 SH   OTR 11 0 1,073 0
ACTIVISION BLIZZARD INC COMMON 00507V109 469,655 6,153,763 SH   DFND 4 6,082,590 0 71,173
ACTIVISION BLIZZARD INC COMMON 00507V109 51,823 679,026 SH   DFND 2 677,221 0 1,805
ACTIVISION BLIZZARD INC COMMON 00507V109 91 1,190 SH   DFND 10 1,190 0 0
ACTIVISION BLIZZARD INC COMMON 00507V109 919 12,042 SH   DFND 5 981 0 11,061
ACTIVISION BLIZZARD INC COMMON 00507V109 16,214 212,454 SH   DFND 8 212,454 0 0
ACTIVISION BLIZZARD INC COMMON 00507V109 419 5,496 SH   DFND 1 246 0 5,250
ACTIVISION BLIZZARD INC COMMON 00507V109 12,991 170,214 SH   DFND 11 170,214 0 0
ACTIVISION BLIZZARD INC COMMON 00507V109 59 775 SH   DFND 16 0 0 775
ACTIVISION BLIZZARD INC COMMON 00507V109 401 5,257 SH   DFND 19 5,257 0 0
ACTIVISION BLIZZARD INC COMMON 00507V109 80 1,053 SH   DFND 2 1,053 0 0
ACTIVISION BLIZZARD INC COMMON 00507V109 10,431 136,670 SH   DFND 5 136,670 0 0
ACTIVISION BLIZZARD INC COMMON 00507V109 2,058 26,960 SH   DFND 15 26,960 0 0
ACTIVISION BLIZZARD INC OPTION 00507V909 687 9,000 SH Call DFND 5 9,000 0 0
ACTIVISION BLIZZARD INC OPTION 00507V959 3,175 41,600 SH Put DFND 5 41,600 0 0
ACTUANT CORP COMMON 00508X203 4 122 SH   OTR 1 122 0 0
ACTUANT CORP COMMON 00508X203 6,521 222,167 SH   DFND 4 208,492 0 13,675
ACTUANT CORP COMMON 00508X203 5 167 SH   DFND 1 167 0 0
ACTUANT CORP COMMON 00508X203 146 4,986 SH   DFND 11 4,986 0 0
ACTUANT CORP COMMON 00508X203 120 4,100 SH   DFND 16 0 0 4,100
ACTUANT CORP COMMON 00508X203 2,304 78,500 SH   DFND 2 76,783 0 1,717
ACTUANT CORP COMMON 00508X203 8,432 287,307 SH   DFND 5 287,307 0 0
ACUITY BRANDS INC COMMON 00508Y102 7,689 66,363 SH   DFND 5 66,363 0 0
ACUITY BRANDS INC COMMON 00508Y102 140,613 1,213,545 SH   DFND 4 1,180,354 0 33,191
ACUITY BRANDS INC COMMON 00508Y102 3,424 29,551 SH   DFND 2 29,551 0 0
ACUITY BRANDS INC COMMON 00508Y102 16 140 SH   DFND 6 140 0 0
ACUITY BRANDS INC COMMON 00508Y102 5 43 SH   DFND 19 43 0 0
ACUSHNET HOLDINGS CORP COMMON 005098108 2,342 95,762 SH   DFND 2 95,762 0 0
ACUSHNET HOLDINGS CORP COMMON 005098108 96,390 3,940,739 SH   DFND 4 3,613,706 0 327,033
ACUSHNET HOLDINGS CORP COMMON 005098108 45 1,828 SH   DFND 5 0 0 1,828
ACUSHNET HOLDINGS CORP COMMON 005098108 3,816 156,021 SH   DFND 8 156,021 0 0
ACXIOM CORP COMMON 005125109 1,360 45,413 SH   DFND 2 45,413 0 0
ACXIOM CORP COMMON 005125109 127 4,255 SH   DFND 4 4,255 0 0
ACXIOM CORP COMMON 005125109 1,643 54,858 SH   DFND 5 54,858 0 0
ADAM NAT RES FD INC FUND 00548F105 15 744 SH   DFND 2 0 0 744
ADAMAS PHARMACEUTICALS INC COMMON 00548A106 3,734 144,575 SH   DFND 4 128,775 0 15,800
ADAMAS PHARMACEUTICALS INC COMMON 00548A106 361 13,962 SH   DFND 2 13,962 0 0
ADAMS DIVERSIFIED EQUITY FD FUND 006212104 63 4,096 SH   DFND 2 0 0 4,096
ADAMS RES & ENERGY INC COMMON 006351308 6 150 SH   DFND 16 0 0 150
ADAMS RES & ENERGY INC COMMON 006351308 647 15,050 SH   DFND 4 14,150 0 900
ADAMS RES & ENERGY INC COMMON 006351308 19 440 SH   DFND 2 440 0 0
ADAMS RES & ENERGY INC COMMON 006351308 61 1,418 SH   DFND 5 1,418 0 0
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ADDUS HOMECARE CORP COMMON 006739106 4,296 75,048 SH   DFND 2 75,048 0 0
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ADVANSIX INC COMMON 00773T101 5,231 142,797 SH   DFND 2 142,348 0 449
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ADVERUM BIOTECHNOLOGIES INC COMMON 00773U108 679 128,154 SH   DFND 2 128,154 0 0
ADVERUM BIOTECHNOLOGIES INC COMMON 00773U108 984 185,730 SH   DFND 5 185,730 0 0
ADVERUM BIOTECHNOLOGIES INC COMMON 00773U108 9,620 1,815,062 SH   DFND 4 1,688,165 0 126,897
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AERIE PHARMACEUTICALS INC COMMON 00771V108 488 7,224 SH   DFND 2 7,224 0 0
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AFFILIATED MANAGERS GROUP COMMON 008252108 40,788 274,350 SH   DFND 4 258,650 0 15,700
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AGNICO EAGLE MINES LTD COMMON 008474108 92 2,017 SH   DFND 10 2,017 0 0
AGNICO EAGLE MINES LTD COMMON 008474108 271 5,919 SH   DFND 5 840 0 5,079
AGNICO EAGLE MINES LTD COMMON 008474108 4,809 104,927 SH   DFND 4 104,927 0 0
AGNICO EAGLE MINES LTD COMMON 008474108 1,155 25,197 SH   DFND 5 25,197 0 0
AGNICO EAGLE MINES LTD COMMON 008474108 97 2,118 SH   DFND 6 2,118 0 0
AGNICO EAGLE MINES LTD COMMON 008474108 92 2,000 SH   DFND 16 0 0 2,000
AGNICO EAGLE MINES LTD COMMON 008474108 941 20,537 SH   DFND 19 20,537 0 0
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AGREE REALTY CORP COMMON 008492100 0 6 SH   DFND 5 0 0 6
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AIMMUNE THERAPEUTICS INC COMMON 00900T107 236 8,794 SH   DFND 2 8,794 0 0
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AIR PRODS & CHEMS INC COMMON 009158106 72 464 SH   OTR 4 0 464 0
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AIR PRODS & CHEMS INC COMMON 009158106 465 2,988 SH   OTR 1 1,418 0 1,570
AIR PRODS & CHEMS INC COMMON 009158106 104,716 672,419 SH   DFND 2 670,879 0 1,540
AIR PRODS & CHEMS INC COMMON 009158106 514 3,301 SH   DFND 2 3,301 0 0
AIR PRODS & CHEMS INC COMMON 009158106 286,347 1,838,738 SH   DFND 4 1,725,388 0 113,350
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AIR PRODS & CHEMS INC COMMON 009158106 5,844 37,528 SH   DFND 5 37,528 0 0
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AIR PRODS & CHEMS INC COMMON 009158106 609 3,908 SH   DFND 10 3,908 0 0
AIR PRODS & CHEMS INC COMMON 009158106 1,252 8,040 SH   DFND 16 0 0 8,040
AIR PRODS & CHEMS INC COMMON 009158106 60,324 387,360 SH   DFND 8 387,360 0 0
AIR PRODS & CHEMS INC COMMON 009158106 38 243 SH   DFND 6 243 0 0
AIR PRODS & CHEMS INC COMMON 009158106 4,171 26,784 SH   DFND 19 26,784 0 0
AIR PRODS & CHEMS INC OPTION 009158906 9,531 61,200 SH Call DFND 5 61,200 0 0
AIR PRODS & CHEMS INC OPTION 009158956 10,060 64,600 SH Put DFND 5 64,600 0 0
AIR TRANSPORT SERVICES GRP I COMMON 00922R105 268 11,855 SH   DFND 2 11,855 0 0
AIR TRANSPORT SERVICES GRP I COMMON 00922R105 4,463 197,548 SH   DFND 5 197,548 0 0
AIRCASTLE LTD COMMON G0129K104 531 25,898 SH   DFND 2 10,534 0 15,364
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AIRCASTLE LTD COMMON G0129K104 15 719 SH   DFND 5 0 0 719
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AK STL CORP BOND 001546AP5 251 219,000 PRN   DFND 5 219,000 0 0
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AK STL HLDG CORP COMMON 001547108 19,192 4,422,207 SH   DFND 4 4,203,707 0 218,500
AK STL HLDG CORP COMMON 001547108 2,033 468,388 SH   DFND 2 468,388 0 0
AK STL HLDG CORP COMMON 001547108 1,073 247,287 SH   DFND 5 58,495 0 188,792
AK STL HLDG CORP COMMON 001547108 4,043 931,602 SH   DFND 5 931,602 0 0
AKAMAI TECHNOLOGIES INC COMMON 00971T101 3 44 SH   OTR 1 0 0 44
AKAMAI TECHNOLOGIES INC COMMON 00971T101 7,053 96,312 SH   DFND 2 96,312 0 0
AKAMAI TECHNOLOGIES INC COMMON 00971T101 75 1,023 SH   DFND 5 149 0 874
AKAMAI TECHNOLOGIES INC COMMON 00971T101 3,047 41,605 SH   DFND 4 31,805 0 9,800
AKAMAI TECHNOLOGIES INC COMMON 00971T101 3,178 43,395 SH   DFND 2 43,395 0 0
AKAMAI TECHNOLOGIES INC COMMON 00971T101 168,639 2,302,868 SH   DFND 5 2,302,868 0 0
AKAMAI TECHNOLOGIES INC OPTION 00971T901 4,760 65,000 SH Call DFND 5 65,000 0 0
AKAMAI TECHNOLOGIES INC OPTION 00971T951 4,760 65,000 SH Put DFND 5 65,000 0 0
AKAMAI TECHNOLOGIES INC BOND 00971TAG6 3,616 3,594,000 PRN   DFND 4 1,493,000 2,101,000 0
AKAMAI TECHNOLOGIES INC BOND 00971TAG6 1,958 1,946,000 PRN   DFND 2 1,946,000 0 0
AKAMAI TECHNOLOGIES INC BOND 00971TAG6 3,975 3,950,000 PRN   DFND 8 3,950,000 0 0
AKCEA THERAPEUTICS INC COMMON 00972L107 69 2,923 SH   DFND 2 2,923 0 0
AKCEA THERAPEUTICS INC COMMON 00972L107 33 1,395 SH   DFND 5 0 0 1,395
AKCEA THERAPEUTICS INC OPTION 00972L907 1,280 54,000 SH Call DFND 5 54,000 0 0
AKCEA THERAPEUTICS INC OPTION 00972L957 1,280 54,000 SH Put DFND 5 54,000 0 0
AKEBIA THERAPEUTICS INC COMMON 00972D105 377 37,808 SH   DFND 5 37,808 0 0
AKEBIA THERAPEUTICS INC COMMON 00972D105 1,369 137,200 SH   DFND 4 88,700 0 48,500
AKEBIA THERAPEUTICS INC COMMON 00972D105 1,217 121,938 SH   DFND 2 121,938 0 0
AKEBIA THERAPEUTICS INC OPTION 00972D905 50 5,000 SH Call DFND 5 5,000 0 0
AKORN INC COMMON 009728106 4,009 241,639 SH   DFND 5 241,639 0 0
AKORN INC COMMON 009728106 2,539 153,025 SH   DFND 4 153,025 0 0
AKORN INC COMMON 009728106 2,305 138,964 SH   DFND 2 138,964 0 0
AKORN INC COMMON 009728106 2 100 SH   DFND 1 0 0 100
AKORN INC OPTION 009728906 3,524 212,400 SH Call DFND 5 212,400 0 0
AKORN INC OPTION 009728956 1,659 100,000 SH Put DFND 5 100,000 0 0
AKOUSTIS TECHNOLOGIES INC COMMON 00973N102 0 26 SH   DFND 5 26 0 0
ALAMO GROUP INC COMMON 011311107 908 10,045 SH   DFND 4 10,045 0 0
ALAMO GROUP INC COMMON 011311107 2,814 31,138 SH   DFND 5 31,138 0 0
ALAMO GROUP INC COMMON 011311107 339 3,756 SH   DFND 2 3,756 0 0
ALAMOS GOLD INC NEW COMMON 011532108 0 1 SH   DFND 19 1 0 0
ALARM COM HLDGS INC COMMON 011642105 12 298 SH   DFND 6 298 0 0
ALARM COM HLDGS INC COMMON 011642105 5,212 129,072 SH   DFND 5 129,072 0 0
ALARM COM HLDGS INC COMMON 011642105 701 17,357 SH   DFND 4 17,357 0 0
ALARM COM HLDGS INC COMMON 011642105 253 6,260 SH   DFND 2 6,260 0 0
ALARM COM HLDGS INC COMMON 011642105 905 22,422 SH   DFND 2 22,422 0 0
ALASKA AIR GROUP INC COMMON 011659109 825 13,660 SH   OTR 21 0 13,660 0
ALASKA AIR GROUP INC COMMON 011659109 289 4,791 SH   OTR 13 0 4,791 0
ALASKA AIR GROUP INC COMMON 011659109 17 278 SH   OTR 11 0 278 0
ALASKA AIR GROUP INC COMMON 011659109 2,963 49,057 SH   OTR 2 623 48,434 0
ALASKA AIR GROUP INC COMMON 011659109 5 79 SH   OTR 1 0 0 79
ALASKA AIR GROUP INC COMMON 011659109 376 6,228 SH   DFND 1 5,347 0 881
ALASKA AIR GROUP INC COMMON 011659109 27,138 449,386 SH   DFND 2 449,115 0 271
ALASKA AIR GROUP INC COMMON 011659109 72 1,187 SH   DFND 2 1,187 0 0
ALASKA AIR GROUP INC COMMON 011659109 4,162 68,918 SH   DFND 11 68,918 0 0
ALASKA AIR GROUP INC COMMON 011659109 14,088 233,291 SH   DFND 5 233,291 0 0
ALASKA AIR GROUP INC COMMON 011659109 371 6,141 SH   DFND 5 172 0 5,969
ALASKA AIR GROUP INC COMMON 011659109 1,590 26,329 SH   DFND 4 26,329 0 0
ALASKA COMMUNICATIONS SYS GR COMMON 01167P101 11 6,123 SH   DFND 4 6,123 0 0
ALBANY INTL CORP COMMON 012348108 5 78 SH   OTR 1 0 0 78
ALBANY INTL CORP COMMON 012348108 730 12,138 SH   DFND 2 5,838 0 6,300
ALBANY INTL CORP COMMON 012348108 114 1,894 SH   DFND 4 1,894 0 0
ALBEMARLE CORP COMMON 012653101 3 34 SH   OTR 1 0 0 34
ALBEMARLE CORP COMMON 012653101 5 55 SH   OTR 5 0 55 0
ALBEMARLE CORP COMMON 012653101 3 35 SH   OTR 21 0 35 0
ALBEMARLE CORP COMMON 012653101 85 904 SH   DFND 6 904 0 0
ALBEMARLE CORP COMMON 012653101 0 3 SH   DFND 8 3 0 0
ALBEMARLE CORP COMMON 012653101 13,704 145,276 SH   DFND 5 101,928 0 43,348
ALBEMARLE CORP COMMON 012653101 3,507 37,176 SH   DFND 2 37,164 0 12
ALBEMARLE CORP COMMON 012653101 2,610 27,674 SH   DFND 4 27,674 0 0
ALBEMARLE CORP COMMON 012653101 3,749 39,744 SH   DFND 2 39,744 0 0
ALBIREO PHARMA INC COMMON 01345P106 45 1,262 SH   DFND 5 1,262 0 0
ALBIREO PHARMA INC COMMON 01345P106 66 1,849 SH   DFND 2 1,849 0 0
ALCOA CORP COMMON 013872106 53 1,130 SH   OTR 10 1,130 0 0
ALCOA CORP COMMON 013872106 30 639 SH   OTR 2 454 0 185
ALCOA CORP COMMON 013872106 211 4,504 SH   OTR 18 4,504 0 0
ALCOA CORP COMMON 013872106 67 1,431 SH   DFND 16 1,081 0 350
ALCOA CORP COMMON 013872106 52,220 1,113,903 SH   DFND 4 1,027,635 0 86,268
ALCOA CORP COMMON 013872106 450 9,600 SH   DFND 19 9,600 0 0
ALCOA CORP COMMON 013872106 625 13,329 SH   DFND 2 13,290 0 39
ALCOA CORP COMMON 013872106 190 4,052 SH   DFND 1 4,052 0 0
ALCOA CORP COMMON 013872106 14,822 316,173 SH   DFND 8 290,339 25,834 0
ALCOA CORP COMMON 013872106 236 5,030 SH   DFND 15 5,030 0 0
ALCOA CORP COMMON 013872106 5,637 120,244 SH   DFND 5 42,523 0 77,721
ALCOA CORP COMMON 013872106 2,837 60,523 SH   DFND 10 60,523 0 0
ALCOA CORP COMMON 013872106 55 1,177 SH   DFND 6 1,177 0 0
ALDER BIOPHARMACEUTICALS INC COMMON 014339105 82 5,200 SH   DFND 16 0 0 5,200
ALDER BIOPHARMACEUTICALS INC COMMON 014339105 4,393 278,050 SH   DFND 4 260,925 0 17,125
ALDER BIOPHARMACEUTICALS INC COMMON 014339105 13 828 SH   DFND 5 0 0 828
ALDER BIOPHARMACEUTICALS INC COMMON 014339105 195 12,359 SH   DFND 2 12,359 0 0
ALDEYRA THERAPEUTICS INC COMMON 01438T106 26 3,234 SH   DFND 2 3,234 0 0
ALDEYRA THERAPEUTICS INC COMMON 01438T106 477 60,062 SH   DFND 5 60,062 0 0
ALEXANDER & BALDWIN INC NEW COMMON 014491104 1,845 78,499 SH   DFND 2 71,935 0 6,564
ALEXANDER & BALDWIN INC NEW COMMON 014491104 800 34,035 SH   DFND 5 34,035 0 0
ALEXANDER & BALDWIN INC NEW COMMON 014491104 8,188 348,434 SH   DFND 4 315,968 0 32,466
ALEXANDERS INC COMMON 014752109 166 433 SH   DFND 2 433 0 0
ALEXANDERS INC COMMON 014752109 2,119 5,537 SH   DFND 5 5,537 0 0
ALEXANDERS INC COMMON 014752109 52 135 SH   DFND 4 135 0 0
ALEXANDRIA REAL ESTATE EQ IN COMMON 015271109 56 444 SH   OTR 13 0 444 0
ALEXANDRIA REAL ESTATE EQ IN COMMON 015271109 65 514 SH   OTR 21 0 514 0
ALEXANDRIA REAL ESTATE EQ IN COMMON 015271109 209 1,656 SH   OTR 2 1,445 211 0
ALEXANDRIA REAL ESTATE EQ IN COMMON 015271109 9 72 SH   OTR 1 72 0 0
ALEXANDRIA REAL ESTATE EQ IN COMMON 015271109 2 19 SH   OTR 4 0 19 0
ALEXANDRIA REAL ESTATE EQ IN COMMON 015271109 14,494 114,880 SH   DFND 8 114,880 0 0
ALEXANDRIA REAL ESTATE EQ IN COMMON 015271109 290 2,301 SH   DFND 6 2,301 0 0
ALEXANDRIA REAL ESTATE EQ IN COMMON 015271109 247 1,955 SH   DFND 1 1,942 0 13
ALEXANDRIA REAL ESTATE EQ IN COMMON 015271109 59,297 469,974 SH   DFND 4 426,892 0 43,082
ALEXANDRIA REAL ESTATE EQ IN COMMON 015271109 0 3 SH   DFND 11 3 0 0
ALEXANDRIA REAL ESTATE EQ IN COMMON 015271109 124 982 SH   DFND 10 982 0 0
ALEXANDRIA REAL ESTATE EQ IN COMMON 015271109 18 145 SH   DFND 5 0 0 145
ALEXANDRIA REAL ESTATE EQ IN COMMON 015271109 286 2,270 SH   DFND 16 0 0 2,270
ALEXANDRIA REAL ESTATE EQ IN COMMON 015271109 5,008 39,692 SH   DFND 2 39,674 0 18
ALEXANDRIA REAL ESTATE EQ IN COMMON 015271109 1,062 8,418 SH   DFND 19 8,418 0 0
ALEXCO RESOURCE CORP COMMON 01535P106 0 130 SH   DFND 2 130 0 0
ALEXION PHARMACEUTICALS INC COMMON 015351109 198 1,593 SH   OTR 1 0 0 1,593
ALEXION PHARMACEUTICALS INC COMMON 015351109 5 39 SH   OTR 5 0 39 0
ALEXION PHARMACEUTICALS INC COMMON 015351109 256 2,061 SH   OTR 2 2,061 0 0
ALEXION PHARMACEUTICALS INC COMMON 015351109 3 25 SH   OTR 11 0 25 0
ALEXION PHARMACEUTICALS INC COMMON 015351109 41,321 332,834 SH   DFND 8 224,675 1,660 106,499
ALEXION PHARMACEUTICALS INC COMMON 015351109 781 6,293 SH   DFND 11 6,293 0 0
ALEXION PHARMACEUTICALS INC COMMON 015351109 2,032 16,365 SH   DFND 2 16,365 0 0
ALEXION PHARMACEUTICALS INC COMMON 015351109 87 700 SH   DFND 1 487 0 213
ALEXION PHARMACEUTICALS INC COMMON 015351109 135 1,090 SH   DFND 16 0 0 1,090
ALEXION PHARMACEUTICALS INC COMMON 015351109 34,479 277,718 SH   DFND 2 277,527 0 191
ALEXION PHARMACEUTICALS INC COMMON 015351109 83,752 674,607 SH   DFND 4 553,960 0 120,647
ALEXION PHARMACEUTICALS INC COMMON 015351109 12,744 102,654 SH   DFND 5 94,706 0 7,948
ALEXION PHARMACEUTICALS INC COMMON 015351109 2,968 23,903 SH   DFND 10 23,903 0 0
ALEXION PHARMACEUTICALS INC COMMON 015351109 586 4,717 SH   DFND 15 4,717 0 0
ALEXION PHARMACEUTICALS INC COMMON 015351109 65 527 SH   DFND 19 527 0 0
ALEXION PHARMACEUTICALS INC COMMON 015351109 1,870 15,066 SH   DFND 5 15,066 0 0
ALEXION PHARMACEUTICALS INC OPTION 015351909 10,925 88,000 SH Call DFND 5 88,000 0 0
ALEXION PHARMACEUTICALS INC OPTION 015351959 11,831 95,300 SH Put DFND 5 95,300 0 0
ALGONQUIN PWR UTILS CORP COMMON 015857105 34 3,512 SH   DFND 1 0 0 3,512
ALGONQUIN PWR UTILS CORP COMMON 015857105 915 94,648 SH   DFND 5 94,648 0 0
ALGONQUIN PWR UTILS CORP COMMON 015857105 142 14,681 SH   DFND 6 14,681 0 0
ALIBABA GROUP HLDG LTD ADR 01609W102 60 323 SH   OTR 11 0 323 0
ALIBABA GROUP HLDG LTD ADR 01609W102 12,682 68,354 SH   OTR 2 3,865 64,489 0
ALIBABA GROUP HLDG LTD ADR 01609W102 364 1,961 SH   OTR 1 250 0 1,711
ALIBABA GROUP HLDG LTD ADR 01609W102 25,718 138,620 SH   OTR 4 0 0 138,620
ALIBABA GROUP HLDG LTD ADR 01609W102 9,095 49,022 SH   OTR 21 0 49,022 0
ALIBABA GROUP HLDG LTD ADR 01609W102 5,384 29,022 SH   OTR 13 0 29,022 0
ALIBABA GROUP HLDG LTD ADR 01609W102 64,610 348,248 SH   OTR 18 348,248 0 0
ALIBABA GROUP HLDG LTD ADR 01609W102 1,268 6,835 SH   OTR 8 0 0 6,835
ALIBABA GROUP HLDG LTD ADR 01609W102 30,032 161,871 SH   DFND 17 159,860 2,011 0
ALIBABA GROUP HLDG LTD ADR 01609W102 346,542 1,867,847 SH   DFND 15 1,812,278 0 55,569
ALIBABA GROUP HLDG LTD ADR 01609W102 813,465 4,384,548 SH   DFND 8 2,786,428 179,633 1,418,487
ALIBABA GROUP HLDG LTD ADR 01609W102 369 1,991 SH   DFND 19 1,991 0 0
ALIBABA GROUP HLDG LTD ADR 01609W102 8,959 48,289 SH   DFND 2 48,289 0 0
ALIBABA GROUP HLDG LTD ADR 01609W102 1,765 9,513 SH   DFND 1 6,733 0 2,780
ALIBABA GROUP HLDG LTD ADR 01609W102 9,224 49,716 SH   DFND 11 49,716 0 0
ALIBABA GROUP HLDG LTD ADR 01609W102 51,274 276,367 SH   DFND 16 262,606 0 13,761
ALIBABA GROUP HLDG LTD ADR 01609W102 195,751 1,055,091 SH   DFND 10 380,024 0 675,067
ALIBABA GROUP HLDG LTD ADR 01609W102 1,114,354 6,006,330 SH   DFND 4 4,692,004 223,694 1,090,632
ALIBABA GROUP HLDG LTD ADR 01609W102 5,943 32,034 SH   DFND 6 32,034 0 0
ALIBABA GROUP HLDG LTD ADR 01609W102 296,750 1,599,471 SH   DFND 2 1,565,330 0 34,141
ALIBABA GROUP HLDG LTD ADR 01609W102 34,540 186,170 SH   DFND 5 12,582 0 173,588
ALIBABA GROUP HLDG LTD ADR 01609W102 118,957 641,176 SH   DFND 5 641,176 0 0
ALIBABA GROUP HLDG LTD OPTION 01609W902 137,645 741,900 SH Call DFND 5 741,900 0 0
ALIBABA GROUP HLDG LTD OPTION 01609W952 164,602 887,200 SH Put DFND 5 887,200 0 0
ALICO INC COMMON 016230104 21 667 SH   DFND 2 667 0 0
ALICO INC COMMON 016230104 5 162 SH   DFND 5 162 0 0
ALIGN TECHNOLOGY INC COMMON 016255101 6 18 SH   OTR 1 0 0 18
ALIGN TECHNOLOGY INC COMMON 016255101 109 319 SH   OTR 10 319 0 0
ALIGN TECHNOLOGY INC COMMON 016255101 6,033 17,632 SH   DFND 5 17,632 0 0
ALIGN TECHNOLOGY INC COMMON 016255101 17,177 50,204 SH   DFND 4 48,315 0 1,889
ALIGN TECHNOLOGY INC COMMON 016255101 14 40 SH   DFND 6 40 0 0
ALIGN TECHNOLOGY INC COMMON 016255101 11,092 32,420 SH   DFND 2 32,394 0 26
ALIGN TECHNOLOGY INC COMMON 016255101 15,068 44,041 SH   DFND 5 31,434 0 12,607
ALIGN TECHNOLOGY INC COMMON 016255101 739 2,161 SH   DFND 2 2,161 0 0
ALIO GOLD INC COMMON 01627X108 769 527,050 SH   DFND 5 527,050 0 0
ALJ REGIONAL HLDGS INC COMMON 001627108 1 509 SH   DFND 5 509 0 0
ALKERMES PLC COMMON G01767105 2 45 SH   OTR 1 0 0 45
ALKERMES PLC COMMON G01767105 2,897 70,386 SH   DFND 4 67,419 0 2,967
ALKERMES PLC COMMON G01767105 6 146 SH   DFND 6 146 0 0
ALKERMES PLC COMMON G01767105 47 1,150 SH   DFND 16 0 0 1,150
ALKERMES PLC COMMON G01767105 179 4,343 SH   DFND 2 4,343 0 0
ALKERMES PLC OPTION G01767905 8,236 200,100 SH Call DFND 5 200,100 0 0
ALKERMES PLC OPTION G01767955 148 3,600 SH Put DFND 5 3,600 0 0
ALLEGHANY CORP DEL COMMON 017175100 6 10 SH   OTR 5 0 10 0
ALLEGHANY CORP DEL COMMON 017175100 853 1,484 SH   DFND 10 1,484 0 0
ALLEGHANY CORP DEL COMMON 017175100 6,024 10,477 SH   DFND 5 10,476 0 1
ALLEGHANY CORP DEL COMMON 017175100 1,051 1,828 SH   DFND 8 1,482 100 246
ALLEGHANY CORP DEL COMMON 017175100 185,881 323,289 SH   DFND 4 318,456 0 4,833
ALLEGHANY CORP DEL COMMON 017175100 13,045 22,689 SH   DFND 2 22,608 0 81
ALLEGHANY CORP DEL COMMON 017175100 12,013 20,893 SH   DFND 5 20,893 0 0
ALLEGHANY CORP DEL COMMON 017175100 200 347 SH   DFND 2 347 0 0
ALLEGHANY CORP DEL COMMON 017175100 34 60 SH   DFND 16 0 0 60
ALLEGHENY TECHNOLOGIES INC COMMON 01741R102 1,913 76,153 SH   DFND 2 73,518 0 2,635
ALLEGHENY TECHNOLOGIES INC COMMON 01741R102 100 3,986 SH   DFND 4 3,986 0 0
ALLEGHENY TECHNOLOGIES INC BOND 01741RAG7 14,976 7,912,000 PRN   DFND 5 7,912,000 0 0
ALLEGIANCE BANCSHARES INC COMMON 01748H107 104 2,391 SH   DFND 2 0 0 2,391
ALLEGIANCE BANCSHARES INC COMMON 01748H107 1,157 26,700 SH   DFND 4 21,900 0 4,800
ALLEGIANCE BANCSHARES INC COMMON 01748H107 11 250 SH   DFND 5 0 0 250
ALLEGIANT TRAVEL CO COMMON 01748X102 565 4,069 SH   DFND 2 2,619 0 1,450
ALLEGIANT TRAVEL CO COMMON 01748X102 65 465 SH   DFND 4 465 0 0
ALLEGION PUB LTD CO COMMON G0176J109 1,753 40,546 SH   OTR 2 39,053 1,440 53
ALLEGION PUB LTD CO COMMON G0176J109 151 3,500 SH   OTR 1 2,320 0 1,180
ALLEGION PUB LTD CO COMMON G0176J109 159 2,058 SH   OTR 18 2,058 0 0
ALLEGION PUB LTD CO COMMON G0176J109 40 926 SH   OTR 13 0 926 0
ALLEGION PUB LTD CO COMMON G0176J109 5 115 SH   OTR 4 0 115 0
ALLEGION PUB LTD CO COMMON G0176J109 3,512 45,403 SH   DFND 15 45,403 0 0
ALLEGION PUB LTD CO COMMON G0176J109 224,344 3,010,679 SH   DFND 4 2,234,120 0 776,559
ALLEGION PUB LTD CO COMMON G0176J109 7,540 97,471 SH   DFND 10 92,750 0 4,721
ALLEGION PUB LTD CO COMMON G0176J109 57,221 739,674 SH   DFND 8 676,357 10,950 52,367
ALLEGION PUB LTD CO COMMON G0176J109 397 9,171 SH   DFND 1 8,690 0 481
ALLEGION PUB LTD CO COMMON G0176J109 52,494 757,080 SH   DFND 2 751,692 0 5,388
ALLEGION PUB LTD CO COMMON G0176J109 1,187 15,344 SH   DFND 16 9,074 0 6,270
ALLEGION PUB LTD CO COMMON G0176J109 1,879 24,289 SH   DFND 19 24,289 0 0
ALLEGION PUB LTD CO COMMON G0176J109 4 46 SH   DFND 6 46 0 0
ALLEGION PUB LTD CO COMMON G0176J109 4,211 54,437 SH   DFND 5 54,437 0 0
ALLENA PHARMACEUTICALS INC COMMON 018119107 631 48,455 SH   DFND 5 48,455 0 0
ALLENA PHARMACEUTICALS INC COMMON 018119107 1,130 86,755 SH   DFND 2 86,755 0 0
ALLENA PHARMACEUTICALS INC COMMON 018119107 1,355 104,000 SH   DFND 4 67,300 0 36,700
ALLERGAN PLC COMMON G0177J108 720 4,318 SH   OTR 1 2,489 0 1,829
ALLERGAN PLC COMMON G0177J108 7,340 44,026 SH   OTR 2 42,625 1,303 98
ALLERGAN PLC COMMON G0177J108 136 814 SH   OTR 18 814 0 0
ALLERGAN PLC COMMON G0177J108 140 842 SH   OTR 13 0 842 0
ALLERGAN PLC COMMON G0177J108 18 110 SH   OTR 4 0 110 0
ALLERGAN PLC COMMON G0177J108 40,497 242,903 SH   DFND 5 6,321 0 236,582
ALLERGAN PLC COMMON G0177J108 117,268 703,380 SH   DFND 8 648,671 27,397 27,312
ALLERGAN PLC COMMON G0177J108 1,657 9,939 SH   DFND 15 9,939 0 0
ALLERGAN PLC COMMON G0177J108 419,121 2,513,920 SH   DFND 4 2,271,947 0 241,973
ALLERGAN PLC COMMON G0177J108 8,557 51,323 SH   DFND 10 51,323 0 0
ALLERGAN PLC COMMON G0177J108 685 4,110 SH   DFND 16 0 0 4,110
ALLERGAN PLC COMMON G0177J108 2,232 13,385 SH   DFND 1 12,438 0 947
ALLERGAN PLC COMMON G0177J108 61,170 366,904 SH   DFND 5 366,904 0 0
ALLERGAN PLC COMMON G0177J108 117,041 702,023 SH   DFND 2 696,515 0 5,508
ALLERGAN PLC COMMON G0177J108 2,290 13,735 SH   DFND 19 13,735 0 0
ALLERGAN PLC COMMON G0177J108 376 2,254 SH   DFND 6 2,254 0 0
ALLERGAN PLC COMMON G0177J108 2,798 16,780 SH   DFND 2 16,780 0 0
ALLERGAN PLC OPTION G0177J908 11,854 71,100 SH Call DFND 5 71,100 0 0
ALLERGAN PLC OPTION G0177J958 31,710 190,200 SH Put DFND 5 190,200 0 0
ALLETE INC COMMON 018522300 7 88 SH   DFND 5 0 0 88
ALLETE INC COMMON 018522300 3,621 46,772 SH   DFND 2 46,572 0 200
ALLETE INC COMMON 018522300 2,865 37,009 SH   DFND 4 37,009 0 0
ALLETE INC COMMON 018522300 44 574 SH   DFND 6 574 0 0
ALLIANCE DATA SYSTEMS CORP COMMON 018581108 3 13 SH   OTR 1 0 0 13
ALLIANCE DATA SYSTEMS CORP COMMON 018581108 10 43 SH   OTR 2 43 0 0
ALLIANCE DATA SYSTEMS CORP COMMON 018581108 89 381 SH   OTR 18 381 0 0
ALLIANCE DATA SYSTEMS CORP COMMON 018581108 100,779 432,159 SH   DFND 2 430,867 0 1,292
ALLIANCE DATA SYSTEMS CORP COMMON 018581108 11,501 49,317 SH   DFND 10 39,163 0 10,154
ALLIANCE DATA SYSTEMS CORP COMMON 018581108 52,328 224,389 SH   DFND 8 183,747 1,660 38,982
ALLIANCE DATA SYSTEMS CORP COMMON 018581108 338,652 1,452,194 SH   DFND 4 1,181,205 0 270,989
ALLIANCE DATA SYSTEMS CORP COMMON 018581108 16,289 69,852 SH   DFND 5 69,852 0 0
ALLIANCE DATA SYSTEMS CORP COMMON 018581108 833 3,570 SH   DFND 2 3,570 0 0
ALLIANCE DATA SYSTEMS CORP COMMON 018581108 2,077 8,906 SH   DFND 19 8,906 0 0
ALLIANCE DATA SYSTEMS CORP COMMON 018581108 10 43 SH   DFND 5 0 0 43
ALLIANCE DATA SYSTEMS CORP COMMON 018581108 289 1,240 SH   DFND 16 0 0 1,240
ALLIANCE DATA SYSTEMS CORP COMMON 018581108 688 2,950 SH   DFND 15 2,950 0 0
ALLIANCE ONE INTL INC COMMON 018772301 2,463 155,376 SH   DFND 5 155,376 0 0
ALLIANCE ONE INTL INC COMMON 018772301 27 1,704 SH   DFND 2 1,704 0 0
ALLIANCE RES PARTNER L P COMMON 01877R108 389 21,181 SH   DFND 5 0 0 21,181
ALLIANCE RES PARTNER L P COMMON 01877R108 1,062 57,897 SH   DFND 5 57,897 0 0
ALLIANCE RES PARTNER L P COMMON 01877R108 29,147 1,588,373 SH   DFND 20 1,588,373 0 0
ALLIANCEBERNSTEIN HOLDING LP COMMON 01881G106 933 32,669 SH   OTR 2 32,669 0 0
ALLIANCEBERNSTEIN HOLDING LP COMMON 01881G106 21,316 746,631 SH   DFND 20 746,631 0 0
ALLIANCEBERNSTEIN HOLDING LP COMMON 01881G106 347 12,164 SH   DFND 5 0 0 12,164
ALLIANT ENERGY CORP COMMON 018802108 3 67 SH   OTR 1 0 0 67
ALLIANT ENERGY CORP COMMON 018802108 728 17,205 SH   DFND 2 17,205 0 0
ALLIANT ENERGY CORP COMMON 018802108 1 25 SH   DFND 1 0 0 25
ALLIANT ENERGY CORP COMMON 018802108 2,496 58,981 SH   DFND 2 58,895 0 86
ALLIANT ENERGY CORP COMMON 018802108 628 14,838 SH   DFND 5 14,838 0 0
ALLIANT ENERGY CORP COMMON 018802108 61 1,442 SH   DFND 5 0 0 1,442
ALLIANT ENERGY CORP COMMON 018802108 5,250 124,062 SH   DFND 4 124,062 0 0
ALLIED MOTION TECHNOLOGIES I COMMON 019330109 47 976 SH   DFND 5 976 0 0
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AMTRUST FINL SVCS INC COMMON 032359309 6 400 SH   DFND 5 0 0 400
AMTRUST FINL SVCS INC COMMON 032359309 13,688 939,484 SH   DFND 4 939,484 0 0
AMTRUST FINL SVCS INC COMMON 032359309 12,950 888,785 SH   DFND 8 888,785 0 0
AMTRUST FINL SVCS INC COMMON 032359309 8,552 586,980 SH   DFND 5 586,980 0 0
AMTRUST FINL SVCS INC COMMON 032359309 15,738 1,080,192 SH   DFND 2 1,080,192 0 0
AMYRIS INC COMMON 03236M200 1,424 222,906 SH   DFND 5 222,906 0 0
AMYRIS INC COMMON 03236M200 31 4,922 SH   DFND 2 4,922 0 0
ANADARKO PETE CORP COMMON 032511107 125 1,705 SH   OTR 1 122 0 1,583
ANADARKO PETE CORP COMMON 032511107 4,105 56,040 SH   OTR 2 54,120 0 1,920
ANADARKO PETE CORP COMMON 032511107 1,112 15,187 SH   DFND 19 15,187 0 0
ANADARKO PETE CORP COMMON 032511107 9,693 132,332 SH   DFND 4 118,672 0 13,660
ANADARKO PETE CORP COMMON 032511107 74,296 1,014,274 SH   DFND 8 1,007,807 6,457 10
ANADARKO PETE CORP COMMON 032511107 369 5,042 SH   DFND 11 5,042 0 0
ANADARKO PETE CORP COMMON 032511107 5,056 69,025 SH   DFND 5 1,381 0 67,644
ANADARKO PETE CORP COMMON 032511107 19,075 260,411 SH   DFND 2 226,682 0 33,729
ANADARKO PETE CORP COMMON 032511107 16 218 SH   DFND 1 200 0 18
ANADARKO PETE CORP COMMON 032511107 10,671 145,680 SH   DFND 5 145,680 0 0
ANADARKO PETE CORP COMMON 032511107 239 3,267 SH   DFND 2 3,267 0 0
ANADARKO PETE CORP OPTION 032511907 17,866 243,900 SH Call DFND 5 243,900 0 0
ANADARKO PETE CORP OPTION 032511957 15,339 209,400 SH Put DFND 5 209,400 0 0
ANALOG DEVICES INC COMMON 032654105 9,126 95,144 SH   OTR 21 0 95,144 0
ANALOG DEVICES INC COMMON 032654105 1,879 19,593 SH   OTR 13 0 19,593 0
ANALOG DEVICES INC COMMON 032654105 67 703 SH   OTR 4 0 703 0
ANALOG DEVICES INC COMMON 032654105 10,365 108,059 SH   OTR 2 52,188 50,109 5,762
ANALOG DEVICES INC COMMON 032654105 857 8,938 SH   OTR 1 4,388 0 4,550
ANALOG DEVICES INC COMMON 032654105 324 3,378 SH   OTR 18 3,378 0 0
ANALOG DEVICES INC COMMON 032654105 128,935 1,344,198 SH   DFND 8 1,122,369 16,840 204,989
ANALOG DEVICES INC COMMON 032654105 827 8,617 SH   DFND 2 8,617 0 0
ANALOG DEVICES INC COMMON 032654105 3,632 37,870 SH   DFND 1 36,641 0 1,229
ANALOG DEVICES INC COMMON 032654105 2,814 29,340 SH   DFND 16 0 0 29,340
ANALOG DEVICES INC COMMON 032654105 3,099 32,313 SH   DFND 5 13,244 0 19,069
ANALOG DEVICES INC COMMON 032654105 18,015 187,808 SH   DFND 15 187,808 0 0
ANALOG DEVICES INC COMMON 032654105 241,526 2,517,995 SH   DFND 2 2,510,186 0 7,809
ANALOG DEVICES INC COMMON 032654105 1,504,829 15,688,370 SH   DFND 4 13,967,665 0 1,720,705
ANALOG DEVICES INC COMMON 032654105 5,933 61,858 SH   DFND 19 61,858 0 0
ANALOG DEVICES INC COMMON 032654105 21,056 219,517 SH   DFND 10 159,154 4,082 56,281
ANALOG DEVICES INC OPTION 032654905 13,525 141,000 SH Call DFND 5 141,000 0 0
ANALOG DEVICES INC OPTION 032654955 7,194 75,000 SH Put DFND 5 75,000 0 0
ANAPTYSBIO INC COMMON 032724106 51 714 SH   OTR 1 0 0 714
ANAPTYSBIO INC COMMON 032724106 7,207 101,446 SH   DFND 4 92,546 0 8,900
ANAPTYSBIO INC COMMON 032724106 1,742 24,520 SH   DFND 2 24,268 0 252
ANAPTYSBIO INC COMMON 032724106 16 221 SH   DFND 1 0 0 221
ANDEAVOR COMMON 03349M105 1,991 15,176 SH   OTR 2 5 15,171 0
ANDEAVOR COMMON 03349M105 1,427 10,878 SH   OTR 21 0 10,878 0
ANDEAVOR COMMON 03349M105 568 4,328 SH   OTR 13 0 4,328 0
ANDEAVOR COMMON 03349M105 6 49 SH   OTR 1 0 0 49
ANDEAVOR COMMON 03349M105 477 3,636 SH   DFND 5 30 0 3,606
ANDEAVOR COMMON 03349M105 948 7,230 SH   DFND 15 7,230 0 0
ANDEAVOR COMMON 03349M105 95 724 SH   DFND 19 724 0 0
ANDEAVOR COMMON 03349M105 222,822 1,698,594 SH   DFND 4 1,375,137 0 323,457
ANDEAVOR COMMON 03349M105 790 6,020 SH   DFND 16 0 0 6,020
ANDEAVOR COMMON 03349M105 10,825 82,517 SH   DFND 8 41,032 7,410 34,075
ANDEAVOR COMMON 03349M105 3,283 25,029 SH   DFND 10 25,029 0 0
ANDEAVOR COMMON 03349M105 60,468 460,957 SH   DFND 2 460,898 0 59
ANDEAVOR COMMON 03349M105 18 141 SH   DFND 6 141 0 0
ANDEAVOR COMMON 03349M105 9,230 70,359 SH   DFND 5 70,359 0 0
ANDEAVOR COMMON 03349M105 26 197 SH   DFND 2 197 0 0
ANDEAVOR COMMON 03349M105 1 5 SH   DFND 1 0 0 5
ANDEAVOR OPTION 03349M905 4,854 37,000 SH Call DFND 5 37,000 0 0
ANDEAVOR OPTION 03349M955 7,871 60,000 SH Put DFND 5 60,000 0 0
ANDEAVOR LOGISTICS LP COMMON 03350F106 4,592 107,980 SH   DFND 5 700 0 107,280
ANDEAVOR LOGISTICS LP COMMON 03350F106 1 14 SH   DFND 6 14 0 0
ANDEAVOR LOGISTICS LP COMMON 03350F106 13,036 306,521 SH   DFND 5 306,521 0 0
ANDEAVOR LOGISTICS LP COMMON 03350F106 133,474 3,138,356 SH   DFND 20 3,138,356 0 0
ANDERSONS INC COMMON 034164103 1,535 44,879 SH   DFND 5 44,879 0 0
ANDERSONS INC COMMON 034164103 287 8,379 SH   DFND 2 5,379 0 3,000
ANDERSONS INC COMMON 034164103 174 5,086 SH   DFND 4 5,086 0 0
ANGIODYNAMICS INC COMMON 03475V101 1,137 51,126 SH   DFND 2 51,126 0 0
ANGIODYNAMICS INC COMMON 03475V101 14,809 665,852 SH   DFND 4 612,827 0 53,025
ANGIODYNAMICS INC COMMON 03475V101 1,040 46,745 SH   DFND 5 46,745 0 0
ANGIODYNAMICS INC COMMON 03475V101 122 5,500 SH   DFND 16 0 0 5,500
ANGLOGOLD ASHANTI LTD ADR 035128206 52 6,378 SH   DFND 4 6,378 0 0
ANGLOGOLD ASHANTI LTD ADR 035128206 0 2 SH   DFND 5 0 0 2
ANGLOGOLD ASHANTI LTD ADR 035128206 46 5,570 SH   DFND 10 5,570 0 0
ANGLOGOLD ASHANTI LTD ADR 035128206 2,519 306,865 SH   DFND 5 306,865 0 0
ANGLOGOLD ASHANTI LTD ADR 035128206 392 47,793 SH   DFND 8 39,667 8,126 0
ANHEUSER BUSCH INBEV SA/NV ADR 03524A108 1,617 16,046 SH   OTR 2 1,000 15,046 0
ANHEUSER BUSCH INBEV SA/NV ADR 03524A108 993 9,856 SH   OTR 21 0 9,856 0
ANHEUSER BUSCH INBEV SA/NV ADR 03524A108 26 263 SH   OTR 1 0 0 263
ANHEUSER BUSCH INBEV SA/NV ADR 03524A108 248 2,460 SH   OTR 13 0 2,460 0
ANHEUSER BUSCH INBEV SA/NV ADR 03524A108 4,339 43,058 SH   DFND 4 43,058 0 0
ANHEUSER BUSCH INBEV SA/NV ADR 03524A108 1,206 11,967 SH   DFND 5 5,518 0 6,449
ANHEUSER BUSCH INBEV SA/NV ADR 03524A108 6,111 60,645 SH   DFND 2 60,645 0 0
ANHEUSER BUSCH INBEV SA/NV ADR 03524A108 6,976 69,235 SH   DFND 5 69,235 0 0
ANHEUSER BUSCH INBEV SA/NV ADR 03524A108 66 651 SH   DFND 2 0 0 651
ANI PHARMACEUTICALS INC COMMON 00182C103 105 1,565 SH   DFND 2 1,565 0 0
ANI PHARMACEUTICALS INC COMMON 00182C103 2,198 32,901 SH   DFND 5 32,901 0 0
ANI PHARMACEUTICALS INC COMMON 00182C103 62 933 SH   DFND 4 933 0 0
ANI PHARMACEUTICALS INC BOND 00182CAA1 2,913 2,564,000 PRN   DFND 5 2,564,000 0 0
ANIKA THERAPEUTICS INC COMMON 035255108 94 2,926 SH   DFND 2 2,926 0 0
ANIKA THERAPEUTICS INC COMMON 035255108 146 4,569 SH   DFND 5 4,569 0 0
ANIKA THERAPEUTICS INC COMMON 035255108 45 1,408 SH   DFND 4 1,408 0 0
ANIXTER INTL INC COMMON 035290105 4,580 72,359 SH   DFND 5 72,359 0 0
ANIXTER INTL INC COMMON 035290105 797 12,587 SH   DFND 2 12,587 0 0
ANIXTER INTL INC COMMON 035290105 817 12,912 SH   DFND 4 12,912 0 0
ANNALY CAP MGMT INC COMMON 035710409 708 68,800 SH   OTR 2 68,800 0 0
ANNALY CAP MGMT INC COMMON 035710409 29 2,800 SH   DFND 2 2,800 0 0
ANNALY CAP MGMT INC COMMON 035710409 328 31,913 SH   DFND 6 31,913 0 0
ANNALY CAP MGMT INC COMMON 035710409 788 76,552 SH   DFND 5 19,996 0 56,556
ANNALY CAP MGMT INC COMMON 035710409 12,555 1,220,124 SH   DFND 5 1,220,124 0 0
ANNALY CAP MGMT INC COMMON 035710409 1,623 157,764 SH   DFND 4 150,045 0 7,719
ANNALY CAP MGMT INC COMMON 035710409 735 71,458 SH   DFND 8 71,458 0 0
ANSYS INC COMMON 03662Q105 295 1,692 SH   DFND 2 1,692 0 0
ANSYS INC COMMON 03662Q105 3,728 21,404 SH   DFND 2 21,367 0 37
ANSYS INC COMMON 03662Q105 3,529 20,262 SH   DFND 4 20,262 0 0
ANSYS INC COMMON 03662Q105 52 301 SH   DFND 5 160 0 141
ANSYS INC COMMON 03662Q105 14,868 85,358 SH   DFND 5 85,358 0 0
ANTARES PHARMA INC COMMON 036642106 76 29,317 SH   DFND 2 29,317 0 0
ANTARES PHARMA INC COMMON 036642106 1,069 414,277 SH   DFND 5 414,277 0 0
ANTERO MIDSTREAM GP LP COMMON 03675Y103 3 179 SH   OTR 1 179 0 0
ANTERO MIDSTREAM GP LP COMMON 03675Y103 571 30,271 SH   DFND 5 260 0 30,011
ANTERO MIDSTREAM GP LP COMMON 03675Y103 408 21,654 SH   DFND 5 21,654 0 0
ANTERO MIDSTREAM GP LP COMMON 03675Y103 46 2,452 SH   DFND 1 244 0 2,208
ANTERO MIDSTREAM GP LP COMMON 03675Y103 1,775 94,108 SH   DFND 2 94,108 0 0
ANTERO MIDSTREAM GP LP COMMON 03675Y103 30,970 1,642,095 SH   DFND 4 1,642,095 0 0
ANTERO MIDSTREAM GP LP COMMON 03675Y103 139 7,379 SH   DFND 11 7,379 0 0
ANTERO MIDSTREAM PARTNERS LP COMMON 03673L103 1,028 34,814 SH   DFND 5 208 0 34,606
ANTERO MIDSTREAM PARTNERS LP COMMON 03673L103 47,927 1,623,533 SH   DFND 20 1,623,533 0 0
ANTERO MIDSTREAM PARTNERS LP COMMON 03673L103 1 25 SH   DFND 6 25 0 0
ANTERO MIDSTREAM PARTNERS LP COMMON 03673L103 248 8,400 SH   DFND 2 0 0 8,400
ANTERO MIDSTREAM PARTNERS LP COMMON 03673L103 3,815 129,250 SH   DFND 5 129,250 0 0
ANTERO RES CORP COMMON 03674X106 10 460 SH   OTR 21 0 460 0
ANTERO RES CORP COMMON 03674X106 23,062 1,080,172 SH   DFND 4 1,078,870 0 1,302
ANTERO RES CORP COMMON 03674X106 296 13,886 SH   DFND 8 7,404 1,660 4,822
ANTERO RES CORP COMMON 03674X106 2 104 SH   DFND 2 104 0 0
ANTERO RES CORP COMMON 03674X106 22,633 1,060,098 SH   DFND 2 1,060,098 0 0
ANTERO RES CORP COMMON 03674X106 27 1,252 SH   DFND 10 1,252 0 0
ANTERO RES CORP COMMON 03674X106 225 10,539 SH   DFND 5 0 0 10,539
ANTERO RES CORP COMMON 03674X106 26 1,230 SH   DFND 16 0 0 1,230
ANTHEM INC COMMON 036752103 375 1,577 SH   OTR 1 0 0 1,577
ANTHEM INC COMMON 036752103 854 3,588 SH   OTR 18 3,588 0 0
ANTHEM INC COMMON 036752103 186 780 SH   OTR 10 780 0 0
ANTHEM INC COMMON 036752103 141,915 596,205 SH   DFND 8 442,813 23,682 129,710
ANTHEM INC COMMON 036752103 1,716 7,209 SH   DFND 5 198 0 7,011
ANTHEM INC COMMON 036752103 18,424 77,401 SH   DFND 2 76,310 0 1,091
ANTHEM INC COMMON 036752103 135 567 SH   DFND 16 567 0 0
ANTHEM INC COMMON 036752103 98,911 415,539 SH   DFND 5 415,539 0 0
ANTHEM INC COMMON 036752103 6,674 28,040 SH   DFND 10 28,040 0 0
ANTHEM INC COMMON 036752103 144,913 608,802 SH   DFND 4 578,017 0 30,785
ANTHEM INC COMMON 036752103 1,993 8,371 SH   DFND 2 8,371 0 0
ANTHEM INC COMMON 036752103 56 235 SH   DFND 19 235 0 0
ANTHEM INC COMMON 036752103 38 160 SH   DFND 1 0 0 160
ANTHEM INC OPTION 036752903 13,472 56,600 SH Call DFND 5 56,600 0 0
ANTHEM INC OPTION 036752953 22,875 96,100 SH Put DFND 5 96,100 0 0
ANTHEM INC BOND 94973VBG1 9,310 2,838,000 PRN   DFND 4 597,000 2,241,000 0
ANTHEM INC BOND 94973VBG1 1,247 380,000 PRN   DFND 2 380,000 0 0
ANTHEM INC BOND 94973VBG1 14,351 4,375,000 PRN   DFND 5 4,375,000 0 0
ANTHEM INC BOND 94973VBG1 1,486 453,000 PRN   DFND 8 374,000 0 79,000
ANWORTH MORTGAGE ASSET CP COMMON 037347101 98 19,780 SH   DFND 2 19,780 0 0
ANWORTH MORTGAGE ASSET CP COMMON 037347101 127 25,506 SH   DFND 5 25,506 0 0
AON PLC COMMON G0408V102 12 85 SH   OTR 1 0 0 85
AON PLC COMMON G0408V102 414 3,020 SH   OTR 2 3,020 0 0
AON PLC COMMON G0408V102 354 2,581 SH   DFND 8 2,581 0 0
AON PLC COMMON G0408V102 319 2,323 SH   DFND 5 300 0 2,023
AON PLC COMMON G0408V102 20,357 148,405 SH   DFND 2 78,381 0 70,024
AON PLC COMMON G0408V102 7,070 51,539 SH   DFND 4 48,583 0 2,956
AON PLC COMMON G0408V102 21 155 SH   DFND 19 155 0 0
AON PLC COMMON G0408V102 2,358 17,193 SH   DFND 2 17,193 0 0
AON PLC COMMON G0408V102 98 715 SH   DFND 1 0 0 715
AON PLC COMMON G0408V102 12,600 91,855 SH   DFND 5 91,855 0 0
AON PLC OPTION G0408V902 2,743 20,000 SH Call DFND 5 20,000 0 0
AON PLC OPTION G0408V952 2,743 20,000 SH Put DFND 5 20,000 0 0
APACHE CORP COMMON 037411105 0 1 SH   OTR 11 0 1 0
APACHE CORP COMMON 037411105 316 6,758 SH   OTR 1 6,603 0 155
APACHE CORP COMMON 037411105 85 1,820 SH   OTR 2 1,820 0 0
APACHE CORP COMMON 037411105 140 2,995 SH   DFND 11 2,995 0 0
APACHE CORP COMMON 037411105 43 920 SH   DFND 2 920 0 0
APACHE CORP COMMON 037411105 13,457 287,846 SH   DFND 2 286,673 0 1,173
APACHE CORP COMMON 037411105 1,465 31,335 SH   DFND 5 85 0 31,250
APACHE CORP COMMON 037411105 9 191 SH   DFND 19 191 0 0
APACHE CORP COMMON 037411105 5,666 121,206 SH   DFND 4 121,206 0 0
APACHE CORP COMMON 037411105 241 5,151 SH   DFND 1 5,151 0 0
APACHE CORP OPTION 037411905 1,819 38,900 SH Call DFND 5 38,900 0 0
APACHE CORP OPTION 037411955 1,720 36,800 SH Put DFND 5 36,800 0 0
APARTMENT INVT & MGMT CO COMMON 03748R101 12 290 SH   OTR 1 0 0 290
APARTMENT INVT & MGMT CO COMMON 03748R101 12,854 303,886 SH   DFND 4 273,591 0 30,295
APARTMENT INVT & MGMT CO COMMON 03748R101 3,342 78,999 SH   DFND 2 72,178 0 6,821
APARTMENT INVT & MGMT CO COMMON 03748R101 4,254 100,558 SH   DFND 5 100,558 0 0
APARTMENT INVT & MGMT CO COMMON 03748R101 322 7,623 SH   DFND 6 7,623 0 0
APARTMENT INVT & MGMT CO COMMON 03748R101 2 37 SH   DFND 5 0 0 37
APARTMENT INVT & MGMT CO COMMON 03748R101 228 5,388 SH   DFND 8 5,388 0 0
APELLIS PHARMACEUTICALS INC COMMON 03753U106 1,065 48,406 SH   DFND 4 48,406 0 0
APELLIS PHARMACEUTICALS INC COMMON 03753U106 161 7,335 SH   DFND 2 7,335 0 0
APERGY CORP COMMON 03755L104 113 2,699 SH   OTR 21 0 2,699 0
APERGY CORP COMMON 03755L104 706 16,913 SH   OTR 2 13,898 1,815 1,200
APERGY CORP COMMON 03755L104 2 53 SH   OTR 4 0 53 0
APERGY CORP COMMON 03755L104 22 518 SH   OTR 1 266 0 252
APERGY CORP COMMON 03755L104 112 2,672 SH   OTR 13 0 2,672 0
APERGY CORP COMMON 03755L104 134 3,210 SH   DFND 16 0 0 3,210
APERGY CORP COMMON 03755L104 6,290 150,647 SH   DFND 2 146,606 0 4,041
APERGY CORP COMMON 03755L104 2 37 SH   DFND 2 37 0 0
APERGY CORP COMMON 03755L104 414 9,925 SH   DFND 19 9,925 0 0
APERGY CORP COMMON 03755L104 44 1,059 SH   DFND 1 799 0 260
APERGY CORP COMMON 03755L104 2,026 48,537 SH   DFND 5 48,537 0 0
APERGY CORP COMMON 03755L104 59 1,406 SH   DFND 5 7 0 1,399
APERGY CORP COMMON 03755L104 62,721 1,502,294 SH   DFND 4 1,459,329 0 42,965
APERGY CORP COMMON 03755L104 444 10,626 SH   DFND 10 10,626 0 0
APERGY CORP COMMON 03755L104 6,786 162,536 SH   DFND 8 162,536 0 0
APOGEE ENTERPRISES INC COMMON 037598109 5 96 SH   DFND 5 0 0 96
APOGEE ENTERPRISES INC COMMON 037598109 579 12,022 SH   DFND 2 5,622 0 6,400
APOGEE ENTERPRISES INC COMMON 037598109 7,578 157,311 SH   DFND 5 157,311 0 0
APOGEE ENTERPRISES INC COMMON 037598109 2 34 SH   DFND 1 0 0 34
APOGEE ENTERPRISES INC COMMON 037598109 288 5,987 SH   DFND 4 5,987 0 0
APOLLO COML REAL EST FIN INC COMMON 03762U105 5 264 SH   OTR 1 0 0 264
APOLLO COML REAL EST FIN INC COMMON 03762U105 132 7,200 SH   DFND 5 0 0 7,200
APOLLO COML REAL EST FIN INC COMMON 03762U105 458 25,052 SH   DFND 2 25,052 0 0
APOLLO COML REAL EST FIN INC COMMON 03762U105 2 127 SH   DFND 1 0 0 127
APOLLO COML REAL EST FIN INC COMMON 03762U105 22,504 1,231,050 SH   DFND 5 1,231,050 0 0
APOLLO COML REAL EST FIN INC BOND 03762UAA3 6,395 6,000,000 PRN   DFND 5 6,000,000 0 0
APOLLO COML REAL EST FIN INC BOND 03762UAB1 1,695 1,695,000 PRN   DFND 5 1,695,000 0 0
APOLLO ENDOSURGERY INC COMMON 03767D108 80 11,432 SH   DFND 5 11,432 0 0
APOLLO GLOBAL MGMT LLC COMMON 037612306 883 27,700 SH   OTR 2 27,700 0 0
APOLLO GLOBAL MGMT LLC COMMON 037612306 14,041 440,571 SH   DFND 5 176,912 0 263,659
APOLLO GLOBAL MGMT LLC COMMON 037612306 144,071 4,520,576 SH   DFND 20 4,520,576 0 0
APOLLO GLOBAL MGMT LLC COMMON 037612306 74,748 2,345,397 SH   DFND 5 2,345,397 0 0
APOLLO GLOBAL MGMT LLC COMMON 037612306 271 8,500 SH   DFND 2 0 0 8,500
APOLLO GLOBAL MGMT LLC COMMON 037612306 288 9,051 SH   DFND 2 9,051 0 0
APOLLO INVT CORP COMMON 03761U106 602 108,000 SH   OTR 2 108,000 0 0
APOLLO INVT CORP COMMON 03761U106 260 46,751 SH   DFND 2 46,751 0 0
APOLLO INVT CORP COMMON 03761U106 141 25,393 SH   DFND 5 25,393 0 0
APOLLO INVT CORP COMMON 03761U106 434 77,870 SH   DFND 5 0 0 77,870
APOLLO MEDICAL HLDGS INC COMMON 03763A207 3 113 SH   DFND 5 113 0 0
APOLLO MEDICAL HLDGS INC COMMON 03763A207 130 5,031 SH   DFND 2 5,031 0 0
APOLLO SR FLOATING RATE FD I FUND 037636107 478 29,295 SH   DFND 5 0 0 29,295
APOLLO SR FLOATING RATE FD I FUND 037636107 197 12,090 SH   DFND 5 12,090 0 0
APOLLO TACTICAL INCOME FD IN FUND 037638103 55 3,500 SH   DFND 5 0 0 3,500
APPFOLIO INC COMMON 03783C100 183 2,997 SH   DFND 2 2,997 0 0
APPIAN CORP COMMON 03782L101 58 1,607 SH   DFND 5 1,607 0 0
APPLE HOSPITALITY REIT INC COMMON 03784Y200 78 4,350 SH   DFND 16 0 0 4,350
APPLE HOSPITALITY REIT INC COMMON 03784Y200 4,867 272,219 SH   DFND 4 271,162 0 1,057
APPLE HOSPITALITY REIT INC COMMON 03784Y200 245 13,680 SH   DFND 5 2,005 0 11,675
APPLE HOSPITALITY REIT INC COMMON 03784Y200 42 2,331 SH   DFND 6 2,331 0 0
APPLE HOSPITALITY REIT INC COMMON 03784Y200 447 25,003 SH   DFND 5 25,003 0 0
APPLE INC COMMON 037833100 10 55 SH   OTR 5 0 55 0
APPLE INC COMMON 037833100 7,417 40,070 SH   OTR 1 31,945 0 8,125
APPLE INC COMMON 037833100 25,408 137,258 SH   OTR 13 0 137,258 0
APPLE INC COMMON 037833100 181,225 979,015 SH   OTR 2 650,721 312,966 15,328
APPLE INC COMMON 037833100 50,493 272,773 SH   OTR 21 0 272,773 0
APPLE INC COMMON 037833100 145 786 SH   OTR 4 0 786 0
APPLE INC COMMON 037833100 706 3,813 SH   OTR 10 3,813 0 0
APPLE INC COMMON 037833100 2,486 13,430 SH   OTR 18 13,430 0 0
APPLE INC COMMON 037833100 241 1,300 SH   OTR 11 0 1,300 0
APPLE INC COMMON 037833100 1,135,853 6,136,100 SH   DFND 2 6,079,251 0 56,849
APPLE INC COMMON 037833100 86,457 467,055 SH   DFND 10 387,876 0 79,179
APPLE INC COMMON 037833100 59,702 322,520 SH   DFND 1 303,765 0 18,755
APPLE INC COMMON 037833100 5,084,307 27,466,407 SH   DFND 4 24,195,484 0 3,270,923
APPLE INC COMMON 037833100 10,351 55,916 SH   DFND 16 2,466 0 53,450
APPLE INC COMMON 037833100 619,255 3,345,335 SH   DFND 8 2,771,831 142,897 430,607
APPLE INC COMMON 037833100 120,361 650,216 SH   DFND 5 650,216 0 0
APPLE INC COMMON 037833100 327 1,764 SH   DFND 2 1,764 0 0
APPLE INC COMMON 037833100 13,944 75,326 SH   DFND 19 75,326 0 0
APPLE INC COMMON 037833100 200,447 1,082,853 SH   DFND 5 191,738 0 891,115
APPLE INC COMMON 037833100 37,965 205,092 SH   DFND 11 205,092 0 0
APPLE INC COMMON 037833100 19,523 105,465 SH   DFND 15 105,465 0 0
APPLE INC OPTION 037833900 517,605 2,796,200 SH Call DFND 5 2,796,200 0 0
APPLE INC OPTION 037833950 822,055 4,440,900 SH Put DFND 5 4,440,900 0 0
APPLIED GENETIC TECHNOL CORP COMMON 03820J100 58 15,570 SH   DFND 2 15,570 0 0
APPLIED INDL TECHNOLOGIES IN COMMON 03820C105 127,251 1,813,982 SH   DFND 4 1,675,425 0 138,557
APPLIED INDL TECHNOLOGIES IN COMMON 03820C105 10,527 150,058 SH   DFND 8 150,058 0 0
APPLIED INDL TECHNOLOGIES IN COMMON 03820C105 114 1,625 SH   DFND 16 0 0 1,625
APPLIED INDL TECHNOLOGIES IN COMMON 03820C105 4,502 64,173 SH   DFND 5 64,173 0 0
APPLIED INDL TECHNOLOGIES IN COMMON 03820C105 6,476 92,322 SH   DFND 2 92,322 0 0
APPLIED MATLS INC COMMON 038222105 38 833 SH   OTR 1 793 0 40
APPLIED MATLS INC COMMON 038222105 263 5,686 SH   OTR 10 5,686 0 0
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APPLIED MATLS INC COMMON 038222105 392 8,484 SH   OTR 18 8,484 0 0
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BARCLAYS BANK PLC PREFERRED 06739H362 211 8,000 SH   DFND 2 8,000 0 0
BARCLAYS BANK PLC PREFERRED 06739H362 992 37,536 SH   DFND 5 6,758 0 30,778
BARCLAYS BK PLC FUND 06738C778 214 8,807 SH   DFND 5 0 0 8,807
BARCLAYS BK PLC FUND 06738C778 1,992 81,815 SH   DFND 5 81,815 0 0
BARCLAYS BK PLC FUND 06742A669 20 168 SH   DFND 5 0 0 168
BARCLAYS BK PLC FUND 06746P563 196 4,400 SH   DFND 5 2,350 0 2,050
BARCLAYS PLC ADR 06738E204 15 1,523 SH   OTR 1 0 0 1,523
BARCLAYS PLC ADR 06738E204 4,509 449,581 SH   DFND 2 192 0 449,389
BARCLAYS PLC ADR 06738E204 216 21,525 SH   DFND 5 13,787 0 7,738
BARCLAYS PLC ADR 06738E204 2 217 SH   DFND 1 0 0 217
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BARNES & NOBLE ED INC COMMON 06777U101 1,900 336,948 SH   DFND 5 336,948 0 0
BARNES & NOBLE INC COMMON 067774109 83 13,108 SH   DFND 2 12,108 0 1,000
BARNES & NOBLE INC COMMON 067774109 28 4,457 SH   DFND 4 4,457 0 0
BARNES GROUP INC COMMON 067806109 2,350 39,899 SH   DFND 2 39,899 0 0
BARNES GROUP INC COMMON 067806109 6,620 112,402 SH   DFND 5 112,402 0 0
BARNES GROUP INC COMMON 067806109 3,282 55,721 SH   DFND 4 47,621 0 8,100
BARNES GROUP INC COMMON 067806109 1,315 22,334 SH   DFND 8 22,334 0 0
BARRETT BUSINESS SERVICES IN COMMON 068463108 9,110 94,339 SH   DFND 2 94,339 0 0
BARRETT BUSINESS SERVICES IN COMMON 068463108 42,803 443,232 SH   DFND 4 378,811 0 64,421
BARRETT BUSINESS SERVICES IN COMMON 068463108 343 3,550 SH   DFND 16 0 0 3,550
BARRETT BUSINESS SERVICES IN COMMON 068463108 684 7,084 SH   DFND 5 7,084 0 0
BARRICK GOLD CORP COMMON 067901108 2,196 167,259 SH   DFND 8 119,247 14,600 33,412
BARRICK GOLD CORP COMMON 067901108 209 15,928 SH   DFND 5 2,815 0 13,113
BARRICK GOLD CORP COMMON 067901108 3,767 286,898 SH   DFND 4 286,898 0 0
BARRICK GOLD CORP COMMON 067901108 139 10,576 SH   DFND 16 0 0 10,576
BARRICK GOLD CORP COMMON 067901108 123 9,404 SH   DFND 10 9,404 0 0
BARRICK GOLD CORP COMMON 067901108 70 5,367 SH   DFND 2 5,367 0 0
BARRICK GOLD CORP COMMON 067901108 29 2,181 SH   DFND 19 2,181 0 0
BARRICK GOLD CORP COMMON 067901108 31,383 2,390,201 SH   DFND 5 2,390,201 0 0
BARRICK GOLD CORP OPTION 067901908 12,874 980,500 SH Call DFND 5 980,500 0 0
BARRICK GOLD CORP OPTION 067901958 37,754 2,875,400 SH Put DFND 5 2,875,400 0 0
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BASSETT FURNITURE INDS INC COMMON 070203104 311 11,295 SH   DFND 5 11,295 0 0
BASSETT FURNITURE INDS INC COMMON 070203104 57 2,075 SH   DFND 2 2,075 0 0
BASSETT FURNITURE INDS INC COMMON 070203104 57 2,053 SH   DFND 4 2,053 0 0
BAXTER INTL INC COMMON 071813109 132 1,791 SH   OTR 10 1,791 0 0
BAXTER INTL INC COMMON 071813109 82 1,117 SH   OTR 18 1,117 0 0
BAXTER INTL INC COMMON 071813109 4,258 57,666 SH   OTR 2 51,044 0 6,622
BAXTER INTL INC COMMON 071813109 13 177 SH   OTR 1 0 0 177
BAXTER INTL INC COMMON 071813109 1,627 22,037 SH   DFND 2 22,037 0 0
BAXTER INTL INC COMMON 071813109 20,922 283,338 SH   DFND 2 282,820 0 518
BAXTER INTL INC COMMON 071813109 9,790 132,586 SH   DFND 5 1,079 0 131,507
BAXTER INTL INC COMMON 071813109 20 275 SH   DFND 19 275 0 0
BAXTER INTL INC COMMON 071813109 5,721 77,483 SH   DFND 8 64,645 12,838 0
BAXTER INTL INC COMMON 071813109 1,579 21,379 SH   DFND 15 21,379 0 0
BAXTER INTL INC COMMON 071813109 87,681 1,187,443 SH   DFND 4 1,161,974 0 25,469
BAXTER INTL INC COMMON 071813109 21,649 293,186 SH   DFND 5 293,186 0 0
BAXTER INTL INC COMMON 071813109 123 1,660 SH   DFND 16 1,660 0 0
BAXTER INTL INC COMMON 071813109 82 1,117 SH   DFND 1 0 0 1,117
BAYCOM CORP COMMON 07272M107 15 599 SH   DFND 5 599 0 0
BAYCOM CORP COMMON 07272M107 51 2,078 SH   DFND 2 0 0 2,078
BAYTEX ENERGY CORP COMMON 07317Q105 2 570 SH   DFND 6 570 0 0
BAYTEX ENERGY CORP COMMON 07317Q105 27,506 8,260,175 SH   DFND 5 8,260,175 0 0
BB&T CORP COMMON 054937107 2,333 46,252 SH   OTR 13 0 46,252 0
BB&T CORP COMMON 054937107 8,340 165,347 SH   OTR 2 55,963 106,384 3,000
BB&T CORP COMMON 054937107 73 1,452 SH   OTR 4 0 1,452 0
BB&T CORP COMMON 054937107 3,035 60,172 SH   OTR 21 0 60,172 0
BB&T CORP COMMON 054937107 490 9,714 SH   OTR 1 5,010 0 4,704
BB&T CORP COMMON 054937107 42,930 851,117 SH   DFND 5 851,117 0 0
BB&T CORP COMMON 054937107 112,272 2,225,845 SH   DFND 2 2,011,125 0 214,720
BB&T CORP COMMON 054937107 59,437 1,178,366 SH   DFND 8 1,094,556 15,800 68,010
BB&T CORP COMMON 054937107 1,931 38,290 SH   DFND 16 0 0 38,290
BB&T CORP COMMON 054937107 4,668 92,547 SH   DFND 10 92,547 0 0
BB&T CORP COMMON 054937107 356,004 7,057,961 SH   DFND 4 6,244,395 0 813,566
BB&T CORP COMMON 054937107 1,236 24,496 SH   DFND 1 24,343 0 153
BB&T CORP COMMON 054937107 427 8,468 SH   DFND 5 3 0 8,465
BB&T CORP COMMON 054937107 1,326 26,290 SH   DFND 15 26,290 0 0
BB&T CORP COMMON 054937107 3,799 75,316 SH   DFND 19 75,316 0 0
BB&T CORP OPTION 054937907 2,522 50,000 SH Call DFND 5 50,000 0 0
BB&T CORP OPTION 054937957 4,035 80,000 SH Put DFND 5 80,000 0 0
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BBX CAP CORP NEW COMMON 05491N104 120 13,329 SH   DFND 2 13,329 0 0
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BCB BANCORP INC COMMON 055298103 133 8,846 SH   DFND 5 8,846 0 0
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BCE INC COMMON 05534B760 875 21,600 SH   OTR 2 21,600 0 0
BCE INC COMMON 05534B760 990 24,442 SH   DFND 8 18,137 1,900 4,405
BCE INC COMMON 05534B760 7,678 189,617 SH   DFND 6 189,617 0 0
BCE INC COMMON 05534B760 202 4,977 SH   DFND 5 0 0 4,977
BCE INC COMMON 05534B760 2,104 51,956 SH   DFND 4 51,956 0 0
BCE INC COMMON 05534B760 58 1,429 SH   DFND 10 1,429 0 0
BCE INC COMMON 05534B760 282 6,975 SH   DFND 1 5,900 0 1,075
BCE INC COMMON 05534B760 363 8,955 SH   DFND 2 0 0 8,955
BCE INC COMMON 05534B760 24 605 SH   DFND 19 605 0 0
BCE INC COMMON 05534B760 57 1,396 SH   DFND 16 0 0 1,396
BCE INC COMMON 05534B760 3 82 SH   DFND 11 82 0 0
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BEACON ROOFING SUPPLY INC COMMON 073685109 1,006 23,598 SH   DFND 2 13,848 0 9,750
BEACON ROOFING SUPPLY INC COMMON 073685109 4,313 101,194 SH   DFND 5 101,194 0 0
BEACON ROOFING SUPPLY INC COMMON 073685109 117 2,744 SH   DFND 4 2,744 0 0
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BEASLEY BROADCAST GROUP INC COMMON 074014101 11 976 SH   DFND 2 976 0 0
BEAZER HOMES USA INC COMMON 07556Q881 8 540 SH   OTR 11 0 540 0
BEAZER HOMES USA INC COMMON 07556Q881 25,954 1,759,604 SH   DFND 4 1,517,679 0 241,925
BEAZER HOMES USA INC COMMON 07556Q881 1,682 114,013 SH   DFND 8 114,013 0 0
BEAZER HOMES USA INC COMMON 07556Q881 98 6,675 SH   DFND 16 0 0 6,675
BEAZER HOMES USA INC COMMON 07556Q881 3,546 240,384 SH   DFND 2 240,384 0 0
BECTON DICKINSON & CO COMMON 075887109 201 838 SH   OTR 1 248 0 590
BECTON DICKINSON & CO COMMON 075887109 298 1,242 SH   OTR 21 0 1,242 0
BECTON DICKINSON & CO COMMON 075887109 11 46 SH   OTR 4 0 46 0
BECTON DICKINSON & CO COMMON 075887109 1,911 7,976 SH   OTR 2 7,466 510 0
BECTON DICKINSON & CO COMMON 075887109 257 1,074 SH   OTR 13 0 1,074 0
BECTON DICKINSON & CO COMMON 075887109 176,880 738,355 SH   DFND 8 581,746 6,110 150,499
BECTON DICKINSON & CO COMMON 075887109 58,805 245,469 SH   DFND 5 245,469 0 0
BECTON DICKINSON & CO COMMON 075887109 6,726 28,076 SH   DFND 5 242 0 27,834
BECTON DICKINSON & CO COMMON 075887109 2,349 9,805 SH   DFND 2 9,805 0 0
BECTON DICKINSON & CO COMMON 075887109 89,016 371,583 SH   DFND 2 371,537 0 46
BECTON DICKINSON & CO COMMON 075887109 927,315 3,870,907 SH   DFND 4 3,466,667 0 404,240
BECTON DICKINSON & CO COMMON 075887109 964 4,025 SH   DFND 1 3,989 0 36
BECTON DICKINSON & CO COMMON 075887109 8,009 33,432 SH   DFND 10 31,341 0 2,091
BECTON DICKINSON & CO COMMON 075887109 2,945 12,295 SH   DFND 15 12,295 0 0
BECTON DICKINSON & CO COMMON 075887109 4,082 17,039 SH   DFND 19 17,039 0 0
BECTON DICKINSON & CO COMMON 075887109 2,264 9,450 SH   DFND 16 0 0 9,450
BECTON DICKINSON & CO OPTION 075887959 335 1,400 SH Put DFND 5 1,400 0 0
BECTON DICKINSON & CO PREF CONV 075887208 27,387 468,711 SH   DFND 4 468,711 0 0
BECTON DICKINSON & CO PREF CONV 075887208 1,110 19,000 SH   DFND 2 19,000 0 0
BECTON DICKINSON & CO PREF CONV 075887208 26,934 460,968 SH   DFND 8 373,774 53,002 34,192
BECTON DICKINSON & CO PREF CONV 075887208 10,740 183,818 SH   DFND 16 133,020 0 50,798
BECTON DICKINSON & CO PREF CONV 075887208 2,598 44,456 SH   DFND 15 44,456 0 0
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BED BATH & BEYOND INC COMMON 075896100 2 81 SH   DFND 1 0 0 81
BED BATH & BEYOND INC COMMON 075896100 97 4,870 SH   DFND 4 4,870 0 0
BED BATH & BEYOND INC COMMON 075896100 1,619 81,235 SH   DFND 2 81,235 0 0
BED BATH & BEYOND INC COMMON 075896100 10 500 SH   DFND 5 0 0 500
BED BATH & BEYOND INC COMMON 075896100 2,142 107,486 SH   DFND 5 107,486 0 0
BED BATH & BEYOND INC OPTION 075896950 777 39,000 SH Put DFND 5 39,000 0 0
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BEIGENE LTD ADR 07725L102 6 40 SH   DFND 2 0 0 40
BEIGENE LTD ADR 07725L102 5,488 35,701 SH   DFND 5 35,701 0 0
BEIGENE LTD ADR 07725L102 5 35 SH   DFND 1 0 0 35
BEIGENE LTD OPTION 07725L902 2,767 18,000 SH Call DFND 5 18,000 0 0
BEIGENE LTD OPTION 07725L952 2,767 18,000 SH Put DFND 5 18,000 0 0
BEL FUSE INC COMMON 077347300 6,871 328,733 SH   DFND 4 297,208 0 31,525
BEL FUSE INC COMMON 077347300 5 250 SH   DFND 16 0 0 250
BEL FUSE INC COMMON 077347300 675 32,308 SH   DFND 2 32,308 0 0
BEL FUSE INC COMMON 077347300 328 15,674 SH   DFND 5 15,674 0 0
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BELDEN INC COMMON 077454106 1,989 32,544 SH   DFND 2 32,342 0 202
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BELLEROPHON THERAPEUTICS INC COMMON 078771102 574 237,369 SH   DFND 5 237,369 0 0
BELLICUM PHARMACEUTICALS INC COMMON 079481107 2,935 397,638 SH   DFND 2 397,638 0 0
BELLICUM PHARMACEUTICALS INC COMMON 079481107 11,945 1,618,514 SH   DFND 4 1,397,197 0 221,317
BELLICUM PHARMACEUTICALS INC COMMON 079481107 11,494 1,557,427 SH   DFND 8 1,302,744 0 254,683
BELMOND LTD COMMON G1154H107 4,699 421,402 SH   DFND 4 416,342 0 5,060
BELMOND LTD COMMON G1154H107 2,942 263,897 SH   DFND 2 263,897 0 0
BEMIS INC COMMON 081437105 295 6,997 SH   OTR 2 0 507 6,490
BEMIS INC COMMON 081437105 20 468 SH   OTR 21 0 468 0
BEMIS INC COMMON 081437105 25 597 SH   OTR 13 0 597 0
BEMIS INC COMMON 081437105 3,302 78,238 SH   DFND 2 78,238 0 0
BEMIS INC COMMON 081437105 1,696 40,175 SH   DFND 2 40,175 0 0
BEMIS INC COMMON 081437105 147 3,487 SH   DFND 5 100 0 3,387
BEMIS INC COMMON 081437105 12,425 294,354 SH   DFND 5 294,354 0 0
BEMIS INC COMMON 081437105 1,394 33,035 SH   DFND 4 33,035 0 0
BENCHMARK ELECTRS INC COMMON 08160H101 5 173 SH   OTR 1 0 0 173
BENCHMARK ELECTRS INC COMMON 08160H101 57,232 1,963,377 SH   DFND 4 1,694,013 0 269,364
BENCHMARK ELECTRS INC COMMON 08160H101 2,356 80,818 SH   DFND 5 80,818 0 0
BENCHMARK ELECTRS INC COMMON 08160H101 7,769 266,535 SH   DFND 2 266,535 0 0
BENCHMARK ELECTRS INC COMMON 08160H101 171 5,850 SH   DFND 16 0 0 5,850
BENEFICIAL BANCORP INC COMMON 08171T102 1,519 93,759 SH   DFND 2 79,899 0 13,860
BENEFICIAL BANCORP INC COMMON 08171T102 36,720 2,266,638 SH   DFND 4 2,053,297 0 213,341
BENEFICIAL BANCORP INC COMMON 08171T102 182 11,250 SH   DFND 16 0 0 11,250
BENEFITFOCUS INC COMMON 08180D106 1,936 57,633 SH   DFND 5 57,633 0 0
BENEFITFOCUS INC COMMON 08180D106 136 4,040 SH   DFND 2 4,040 0 0
BERKSHIRE HATHAWAY INC DEL COMMON 084670108 33,281 118 SH   OTR 2 87 0 31
BERKSHIRE HATHAWAY INC DEL COMMON 084670108 34,691 123 SH   DFND 2 82 0 41
BERKSHIRE HATHAWAY INC DEL COMMON 084670108 3,667 13 SH   DFND 5 0 0 13
BERKSHIRE HATHAWAY INC DEL COMMON 084670108 41,178 146 SH   DFND 4 146 0 0
BERKSHIRE HATHAWAY INC DEL COMMON 084670702 15 79 SH   OTR 4 0 79 0
BERKSHIRE HATHAWAY INC DEL COMMON 084670702 116 624 SH   OTR 13 0 624 0
BERKSHIRE HATHAWAY INC DEL COMMON 084670702 78,048 418,150 SH   OTR 2 380,737 967 36,446
BERKSHIRE HATHAWAY INC DEL COMMON 084670702 1,856 9,943 SH   OTR 1 5,933 0 4,010
BERKSHIRE HATHAWAY INC DEL COMMON 084670702 12 63 SH   OTR 5 0 63 0
BERKSHIRE HATHAWAY INC DEL COMMON 084670702 1,484 7,950 SH   DFND 16 0 0 7,950
BERKSHIRE HATHAWAY INC DEL COMMON 084670702 20,311 108,819 SH   DFND 8 63,333 13,930 31,556
BERKSHIRE HATHAWAY INC DEL COMMON 084670702 27,744 148,644 SH   DFND 1 148,295 0 349
BERKSHIRE HATHAWAY INC DEL COMMON 084670702 1,510 8,088 SH   DFND 2 8,088 0 0
BERKSHIRE HATHAWAY INC DEL COMMON 084670702 258,993 1,387,586 SH   DFND 2 1,243,346 0 144,240
BERKSHIRE HATHAWAY INC DEL COMMON 084670702 634,196 3,397,780 SH   DFND 4 2,794,976 0 602,804
BERKSHIRE HATHAWAY INC DEL COMMON 084670702 97,618 522,998 SH   DFND 5 522,998 0 0
BERKSHIRE HATHAWAY INC DEL COMMON 084670702 171 918 SH   DFND 6 918 0 0
BERKSHIRE HATHAWAY INC DEL COMMON 084670702 7,008 37,545 SH   DFND 15 37,545 0 0
BERKSHIRE HATHAWAY INC DEL COMMON 084670702 136,021 728,749 SH   DFND 5 82,803 0 645,946
BERKSHIRE HATHAWAY INC DEL COMMON 084670702 500 2,680 SH   DFND 19 2,680 0 0
BERKSHIRE HATHAWAY INC DEL COMMON 084670702 5,857 31,378 SH   DFND 10 31,378 0 0
BERKSHIRE HATHAWAY INC DEL OPTION 084670952 28,931 155,000 SH Put DFND 5 155,000 0 0
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BERKSHIRE HILLS BANCORP INC COMMON 084680107 336 8,271 SH   DFND 2 0 0 8,271
BERKSHIRE HILLS BANCORP INC COMMON 084680107 727 17,910 SH   DFND 4 14,710 0 3,200
BERKSHIRE HILLS BANCORP INC COMMON 084680107 1 33 SH   DFND 5 0 0 33
BERRY GLOBAL GROUP INC COMMON 08579W103 1,463 31,848 SH   DFND 5 31,848 0 0
BERRY GLOBAL GROUP INC COMMON 08579W103 204 4,450 SH   DFND 16 0 0 4,450
BERRY GLOBAL GROUP INC COMMON 08579W103 14,177 308,590 SH   DFND 4 291,174 0 17,416
BERRY GLOBAL GROUP INC COMMON 08579W103 100 2,169 SH   DFND 5 454 0 1,715
BERRY GLOBAL GROUP INC COMMON 08579W103 1,833 39,895 SH   DFND 2 39,895 0 0
BEST BUY INC COMMON 086516101 4 54 SH   OTR 2 54 0 0
BEST BUY INC COMMON 086516101 60 800 SH   OTR 11 0 800 0
BEST BUY INC COMMON 086516101 105 1,414 SH   OTR 10 1,414 0 0
BEST BUY INC COMMON 086516101 5 71 SH   OTR 1 0 0 71
BEST BUY INC COMMON 086516101 420 5,629 SH   OTR 18 5,629 0 0
BEST BUY INC COMMON 086516101 15,491 207,704 SH   DFND 8 134,619 42,745 30,340
BEST BUY INC COMMON 086516101 18 243 SH   DFND 5 234 0 9
BEST BUY INC COMMON 086516101 6,125 82,131 SH   DFND 10 82,131 0 0
BEST BUY INC COMMON 086516101 32,434 434,885 SH   DFND 2 434,858 0 27
BEST BUY INC COMMON 086516101 482,732 6,472,670 SH   DFND 4 6,071,660 0 401,010
BEST BUY INC COMMON 086516101 834 11,188 SH   DFND 16 783 0 10,405
BEST BUY INC COMMON 086516101 53,834 721,823 SH   DFND 5 721,823 0 0
BEST BUY INC COMMON 086516101 112 1,500 SH   DFND 6 1,500 0 0
BEST BUY INC COMMON 086516101 78 1,043 SH   DFND 19 1,043 0 0
BEST BUY INC COMMON 086516101 1 8 SH   DFND 1 0 0 8
BEST BUY INC COMMON 086516101 1,512 20,270 SH   DFND 15 20,270 0 0
BEST BUY INC OPTION 086516901 5,221 70,000 SH Call DFND 5 70,000 0 0
BEST BUY INC OPTION 086516951 2,812 37,700 SH Put DFND 5 37,700 0 0
BEST INC ADR 08653C106 1,458 119,285 SH   DFND 5 119,285 0 0
BEST INC ADR 08653C106 64 5,276 SH   DFND 5 2,467 0 2,809
BG STAFFING INC COMMON 05544A109 33 1,401 SH   DFND 2 1,401 0 0
BG STAFFING INC COMMON 05544A109 2 68 SH   DFND 1 0 0 68
BGC PARTNERS INC COMMON 05541T101 198 17,510 SH   OTR 21 0 17,510 0
BGC PARTNERS INC COMMON 05541T101 252 22,301 SH   OTR 13 0 22,301 0
BGC PARTNERS INC COMMON 05541T101 215 18,969 SH   OTR 2 0 18,969 0
BGC PARTNERS INC COMMON 05541T101 41 3,595 SH   OTR 1 0 0 3,595
BGC PARTNERS INC COMMON 05541T101 3,124 275,955 SH   DFND 2 273,536 0 2,419
BGC PARTNERS INC COMMON 05541T101 3,650 322,449 SH   DFND 5 322,449 0 0
BGC PARTNERS INC COMMON 05541T101 7,264 641,668 SH   DFND 4 483,816 0 157,852
BGC PARTNERS INC COMMON 05541T101 2,395 211,543 SH   DFND 8 211,543 0 0
BGC PARTNERS INC COMMON 05541T101 15,393 1,359,843 SH   DFND 5 174,679 0 1,185,164
BGC PARTNERS INC COMMON 05541T101 40 3,537 SH   DFND 1 0 0 3,537
BHP BILLITON LTD ADR 088606108 50 1,000 SH   OTR 2 1,000 0 0
BHP BILLITON LTD ADR 088606108 39 783 SH   OTR 1 0 0 783
BHP BILLITON LTD ADR 088606108 506 10,127 SH   DFND 8 0 10,127 0
BHP BILLITON LTD ADR 088606108 8,623 172,418 SH   DFND 4 172,418 0 0
BHP BILLITON LTD ADR 088606108 284 5,675 SH   DFND 5 0 0 5,675
BHP BILLITON LTD ADR 088606108 4 77 SH   DFND 1 0 0 77
BHP BILLITON LTD ADR 088606108 18,638 372,694 SH   DFND 2 372,694 0 0
BHP BILLITON PLC ADR 05545E209 1 30 SH   OTR 1 0 0 30
BHP BILLITON PLC ADR 05545E209 1 27 SH   DFND 1 0 0 27
BHP BILLITON PLC ADR 05545E209 3,852 85,696 SH   DFND 5 85,696 0 0
BIG 5 SPORTING GOODS CORP COMMON 08915P101 31 4,078 SH   DFND 2 4,078 0 0
BIG LOTS INC COMMON 089302103 1,548 37,051 SH   DFND 2 37,051 0 0
BIG LOTS INC COMMON 089302103 1,475 35,293 SH   DFND 4 35,293 0 0
BIG LOTS INC COMMON 089302103 3 67 SH   DFND 1 0 0 67
BIGLARI HLDGS INC COMMON 08986R309 399 2,175 SH   DFND 5 2,175 0 0
BIGLARI HLDGS INC COMMON 08986R309 37 199 SH   DFND 2 199 0 0
BIGLARI HLDGS INC COMMON 08986R408 12 13 SH   DFND 5 13 0 0
BIGLARI HLDGS INC COMMON 08986R408 87 92 SH   DFND 4 92 0 0
BIGLARI HLDGS INC COMMON 08986R408 19 20 SH   DFND 2 20 0 0
BILIBILI INC ADR 090040106 2,269 162,174 SH   OTR 18 162,174 0 0
BILIBILI INC ADR 090040106 4,852 346,850 SH   DFND 8 233,949 0 112,901
BILIBILI INC ADR 090040106 529 37,803 SH   DFND 16 37,803 0 0
BILIBILI INC ADR 090040106 19,999 1,429,533 SH   DFND 15 1,429,533 0 0
BILIBILI INC ADR 090040106 10,198 728,923 SH   DFND 5 728,923 0 0
BILIBILI INC ADR 090040106 341 24,349 SH   DFND 4 24,349 0 0
BIO RAD LABS INC COMMON 090572207 3,950 13,690 SH   DFND 2 12,440 0 1,250
BIO RAD LABS INC COMMON 090572207 3,306 11,458 SH   DFND 4 10,458 0 1,000
BIO RAD LABS INC COMMON 090572207 3 11 SH   DFND 5 0 0 11
BIO RAD LABS INC COMMON 090572207 1,529 5,300 SH   DFND 5 5,300 0 0
BIO TECHNE CORP COMMON 09073M104 2,354 15,908 SH   DFND 2 14,358 0 1,550
BIO TECHNE CORP COMMON 09073M104 7,231 48,876 SH   DFND 5 48,876 0 0
BIO TECHNE CORP COMMON 09073M104 531 3,592 SH   DFND 4 3,592 0 0
BIOANALYTICAL SYS INC COMMON 09058M103 0 100 SH   DFND 2 0 0 100
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BIOCRYST PHARMACEUTICALS COMMON 09058V103 9 1,519 SH   DFND 1 0 0 1,519
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