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    <periodOfReport>2023-07-22</periodOfReport>

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    <issuer>
        <issuerCik>0001849253</issuerCik>
        <issuerName>RYAN SPECIALTY HOLDINGS, INC.</issuerName>
        <issuerTradingSymbol>RYAN</issuerTradingSymbol>
    </issuer>

    <reportingOwner>
        <reportingOwnerId>
            <rptOwnerCik>0001870017</rptOwnerCik>
            <rptOwnerName>PASCHAL-ALCORN LISA JO</rptOwnerName>
        </reportingOwnerId>
        <reportingOwnerAddress>
            <rptOwnerStreet1>TWO PRUDENTIAL PLAZA</rptOwnerStreet1>
            <rptOwnerStreet2>180 N. STETSON AVE.</rptOwnerStreet2>
            <rptOwnerCity>CHICAGO</rptOwnerCity>
            <rptOwnerState>IL</rptOwnerState>
            <rptOwnerZipCode>60601</rptOwnerZipCode>
            <rptOwnerStateDescription></rptOwnerStateDescription>
        </reportingOwnerAddress>
        <reportingOwnerRelationship>
            <isDirector>0</isDirector>
            <isOfficer>1</isOfficer>
            <isTenPercentOwner>0</isTenPercentOwner>
            <isOther>0</isOther>
            <officerTitle>See Remarks</officerTitle>
        </reportingOwnerRelationship>
    </reportingOwner>

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                    <value>3379</value>
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                <transactionPricePerShare>
                    <value>45.91</value>
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                    <value>3379</value>
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                    <value>153938</value>
                </sharesOwnedFollowingTransaction>
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    <footnotes>
        <footnote id="F1">Shares of Class B common stock, par value $0.001 per share, (&quot;Class B Common Stock&quot;) do not represent economic interests in the Issuer. Except as provided in the Issuer's certificate of incorporation or as required by applicable law, holders of Class B Common Stock will be initially entitled to 10 votes per share on all matters to be voted on by the Issuer's stockholders generally. Upon exchange of Common Units (&quot;Common Units&quot;) of New Ryan Specialty, LLC that are held by the Reporting Person and reported in Table II hereof, for an equal number of shares of Class A common stock, par value $0.001 (&quot;Class A Common Stock&quot;) of the Issuer, an equal number of shares of the Issuer's Class B Common Stock will be cancelled for no consideration.</footnote>
        <footnote id="F2">Class B Common Stock is issued on a one for one basis for each Common Unit issued to the Reporting Person upon vesting of the Reporting Person's Restricted LLC Units.</footnote>
        <footnote id="F3">On July 23, 2021, the Reporting Person was Granted 57,657 Restricted LLC Units which vest 20% on each of July 22, 2022 through July 22, 2026.  Such grant was approved by the Board of the Issuer for purposes of Rule 16(b)(3). Each Restricted LLC Unit represents a contingent right to receive one Common Unit. The Common Units are exchangeable on a one-for-one basis for Class A Common Stock, subject to customary adjustments.</footnote>
        <footnote id="F4">Pursuant to the Amended and Restated Limited Liability Company Agreement of New Ryan Specialty, LLC, as amended, the Reporting Person may exchange all or a portion of such person's Common Units (together with the delivery of an equal number of shares of Class B Common Stock) for shares of Class A Common Stock of the Issuer on a one-for-one basis, subject to customary adjustments, or, at the option of the Issuer, cash (based on the then-market value of the Class A Common Stock).  The Common Units do not expire.</footnote>
    </footnotes>

    <remarks>Senior Vice President and Chief Human Resource Officer</remarks>

    <ownerSignature>
        <signatureName>/s/ Mark S. Katz by Power of Attorney</signatureName>
        <signatureDate>2023-07-24</signatureDate>
    </ownerSignature>
</ownershipDocument>
