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Goodwill and Intangible Assets, net
 12 Months Ended
Dec. 31, 2023
Goodwill and Intangible Assets Disclosure [Abstract]  
Goodwill and Intangible Assets, net Goodwill and Intangible Assets, net
All goodwill is allocated to the Cell Engineering reporting unit and segment identified in Note 15. Changes in the carrying amount of goodwill consisted of the following (in thousands):
As of December 31,
20232022
Beginning balance$60,210 $21,312 
Goodwill acquired in acquisitions— 39,712 
Deconsolidation of Zymergen(10,660)— 
Measurement period adjustments (1)
(2,120)(548)
Impact of foreign currency translation1,808 (266)
Ending balance$49,238 $60,210 
(1)For the year ended December 31, 2023, the measurement period adjustment is primarily related to the Zymergen acquisition. See Note 3 for a description.
Intangible assets, net consisted of the following (in thousands):
Gross
Carrying
Value (1)
Accumulated
Amortization (1)
Net
Carrying
Value
Weighted Average
Amortization Period
(in years)
December 31, 2023
Developed technology (2)
$105,279 $(22,663)$82,616 8.8
Customer relationships380 (261)119 0.9
Assembled workforce190 (184)6 0.3
Total intangible assets$105,849 $(23,108)$82,741 
December 31, 2022
Developed technology$115,824 $(8,825)$106,999 9.4
Database3,700 (107)3,593 6.8
Customer relationships380 (71)309 1.6
Assembled workforce190 (50)140 1.0
Total intangible assets$120,094 $(9,053)$111,041 
(1)Gross carrying value and accumulated amortization include the impact of cumulative foreign currency translation adjustments.
(2)During 2023, the Company deconsolidated $13.5 million of developed technology and $3.2 million of database intangible assets related to the deconsolidation of Zymergen (see Note 3).
Amortization expense was $15.7 million, $5.6 million and $1.2 million for the years ended December 31, 2023, 2022 and 2021, respectively. The estimated future amortization expense for intangible assets remaining as of December 31, 2023 is as follows (in thousands):
2024$13,747 
202513,623 
202613,623 
202710,548 
20283,442 
Thereafter27,758 
Total$82,741