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Upfront amount received.

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Amount of obligation to transfer good or service to customer for which consideration has been received or is receivable.

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The percentage of ownership of common stock or equity participation in the investee accounted for under the equity method of accounting.

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This item represents the entity's ownership percentage in the investee multiplied by the investee's total equity.

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This item represents the carrying amount on the entity's balance sheet of its investment in common stock of an equity method investee. This is not an indicator of the fair value of the investment, rather it is the initial cost adjusted for the entity's share of earnings and losses of the investee, adjusted for any distributions (dividends) and other than temporary impairment (OTTI) losses recognized.

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The cash outflow associated with the purchase of or advances to an equity method investments, which are investments in joint ventures and entities in which the entity has an equity ownership interest normally of 20 to 50 percent and exercises significant influence.

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Amount, excluding tax collected from customer, of revenue from satisfaction of performance obligation by transferring promised good or service to customer. Tax collected from customer is tax assessed by governmental authority that is both imposed on and concurrent with specific revenue-producing transaction, including, but not limited to, sales, use, value added and excise.

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Percentage of the Variable Interest Entity's (VIE) voting interest owned by (or beneficial interest in) the reporting entity (directly or indirectly).

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Significant Collaboration Transactions - Joyn Bio LLC Narrative (Details) - USD ($) $ in Thousands	1 Months Ended	12 Months Ended
	Sep. 30, 2017	Dec. 31, 2023	Dec. 31, 2022	Dec. 31, 2021	Dec. 31, 2020
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]
Variable interest entity ownership percentage			100.00%
Equity method investments		$ 0	$ 1,543
Deferred other income			222,600	$ 189,200
Revenue recognized		$ 251,455	477,706	313,837
Joyn Bio
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]
Equity method investment ownership percentage	50.00%
Equity method investments	$ 97,900
Revenue recognized			$ 2,900	$ 5,300
Joyn Bio \| Foundry Services Agreement
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]
Upfront amount received	20,000				$ 15,000
Cooksonia
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]
Equity method investment, underlying equity in net assets	8,100
Cooksonia \| Parent
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]
Payment to acquire equity method investments	5,000
Equity method investments	$ 13,100
Cooksonia \| Joyn Bio
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]
Equity method investment ownership percentage	50.00%
Capital Unit, Class B \| Cooksonia
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]
Payment to acquire equity method investments	$ 5,000
Capital Unit, Class B \| Cooksonia \| Related Party Investor
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]
Payment to acquire equity method investments	$ 20,000
Capital Unit, Class B \| Cooksonia \| Joyn Bio \| Related Party Investor
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]
Equity method investment ownership percentage	20.00%
Capital Units Class C \| Joyn Bio
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]
Payment to acquire equity method investments	$ 20,000
Capital Units Class C \| Joyn Bio \| Bayer
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]
Payment to acquire equity method investments	$ 20,000
Equity method investment ownership percentage	50.00%
Commitment to contribute additional capital	$ 60,000
Capital Units Class C \| Cooksonia \| Joyn Bio
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]
Equity method investment ownership percentage	50.00%
Capital Units Class C \| Cooksonia \| Joyn Bio \| Related Party Investor
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]
Equity method investment ownership percentage	10.00%
Variable Interest Entity, Primary Beneficiary \| Capital Unit, Class A \| Cooksonia
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]
Payment to acquire equity method investments	$ 5,000
Variable interest entity ownership percentage	70.00%
Non- Controlling Interest \| Cooksonia
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]
Equity method investments	$ 29,700
Equity method investment, underlying equity in net assets	$ 8,100