Significant Collaboration Transactions - Additional Information (Details)			1 Months Ended						5 Months Ended	12 Months Ended
		Jun. 01, 2019 USD ($) $ / shares shares	Nov. 15, 2021 USD ($)	Jul. 31, 2021 shares	Mar. 31, 2021 USD ($) shares	Dec. 31, 2019 USD ($) shares	Jun. 30, 2019 $ / shares shares	Sep. 30, 2017 USD ($)	Dec. 31, 2017 USD ($)	Dec. 31, 2021 USD ($) Obligations shares	Dec. 31, 2020 USD ($) shares	Dec. 31, 2019 USD ($) shares	Dec. 31, 2018 USD ($) Obligations shares	Jul. 31, 2019 USD ($)	Dec. 01, 2018 USD ($)	Sep. 30, 2018 USD ($)
Business Acquisition [Line Items]
Consideration										$ 8,760,000
Number of additional common units received | shares										7,245,000
Loss on equity method investments										$ (77,284,000)	(396,000)	(27,533,000)
Number Of Allocated Performance Obligations | Obligations										10
Revenue remaining performance obligation amount										$ 21,100,000	20,700,000
Deferred Revenue						$ 147,900,000					128,500,000	147,900,000
Equity Method Investments	[1]									13,194,000	28,924,000
Deferred Revenue, Balance						147,900,000					128,500,000	147,900,000
Amyris Inc | Amyris Collaboration Agreement
Business Acquisition [Line Items]
Due from Related Parties									$ 800,000
Synlogic Inc
Business Acquisition [Line Items]
Revenue remaining performance obligation amount														$ 30,000,000.0
Deferred Revenue, Revenue Recognized										100,000	100,000
Equity method investments fair value		$ 35,800,000
Equity Method Investment Difference Between Aggregated Cost And Underlying Equity														29,800,000
Equity method investment														80,000,000.0
Synlogic Inc | Warrant Agreement [Member]
Business Acquisition [Line Items]
Number of ordinary share called by each warrant | shares		2,548,117
Exercise price of warrants or rights | $ / shares		$ 9.00
Synlogic Inc | Foundry Services Agreement [Member]
Business Acquisition [Line Items]
Exercise price of warrants or rights | $ / shares							$ 8.99
Cash		$ 30,000,000.0
Payments for repurchase of warrants		$ 22,900,000
Warrant Exercisable Interest Rate							19.99%
Maximum [Member] | Synlogic Inc
Business Acquisition [Line Items]
Deferred Revenue						200,000						200,000
Deferred Revenue, Balance						200,000						200,000
Promissory Note | Amyris Inc
Business Acquisition [Line Items]
Upfront payment received			$ 22,800,000
Debt instrument, face amount										$ 12,000,000.0
Maturity date										Oct. 19, 2022
Debt Instrument, Periodic Payment										$ 9,800,000
Promissory Note | Amyris Inc | Amyris Collaboration Agreement
Business Acquisition [Line Items]
Due to Related Parties									12,000,000.0
Promissory Note | Minimum [Member] | Amyris Inc
Business Acquisition [Line Items]
Debt Instrument, Periodic Payment										200,000
Promissory Note | Maximum [Member] | Amyris Inc
Business Acquisition [Line Items]
Debt Instrument, Periodic Payment										300,000
Common Stock [Member] | Synlogic Inc
Business Acquisition [Line Items]
Equity metohd investment ownership percentage		19.90%
Stock issued during period shares | shares							6,340,771
Shares issued price per share | $ / shares		$ 9.00
Issuance of Series E convertible preferred stock, net of issuance costs of $4,830		$ 57,100,000
Warrants, each whole warrant exercisable for one Class A ordinary share, each at an exercise price of $11.50 per share [Member] | Synlogic Inc
Business Acquisition [Line Items]
Equity Method Investments														$ 14,400,000
Joyn Bio, LLC [Member]
Business Acquisition [Line Items]
Deferred Revenue										4,600,000	9,900,000
Deferred Revenue, Revenue Recognized										5,300,000	7,300,000	9,300,000
Equity Method Investments										$ 97,900,000
Equity metohd investment ownership percentage										50.00%
Deferred Revenue, Balance										$ 4,600,000	9,900,000
Joyn Bio, LLC [Member] | Foundry Services Agreement [Member]
Business Acquisition [Line Items]
Upfront amount received						$ 15,000,000.0		$ 20,000,000.0				15,000,000.0
Capital Units Class C [Member] | Joyn Bio, LLC [Member]
Business Acquisition [Line Items]
Payment to acquire equity method investments								20,000,000.0
Capital Units Class C [Member] | Bayer [Member] | Joyn Bio, LLC [Member]
Business Acquisition [Line Items]
Payment to acquire equity method investments									$ 20,000.0
Equity metohd investment ownership percentage									50.00%
Commitment to contribute additional capital									$ 60,000,000.0
Partnership Agreement [Member] | Promissory Note | Amyris Inc
Business Acquisition [Line Items]
Debt Instrument, Term Payment			9,800,000
Promissory Note, Principal Balance			$ 12,000,000.0
Arcaea LLC [Member]
Business Acquisition [Line Items]
Carrying value of the equity method investment					$ 11,900,000
Loss on equity method investments										11,900,000
carrying value outstanding										0
Allocated Upfront Non Cash Consideration										1,200,000
Additional Non Cash Consideration										35,500,000
Revenue remaining performance obligation amount										3,600,000
Deferred Revenue										47,400,000
Deferred Revenue, Revenue Recognized										3,700,000
Deferred Revenue, Balance										47,400,000
Variable Interest Entity, Primary Beneficiary [Member]
Business Acquisition [Line Items]
Equity Method Investments										$ 11,694,000	28,924,000
Arcaea [Member]
Business Acquisition [Line Items]
Consideration					$ 35,500,000
Additional preferred units issued | shares										5,229,900
Number of common units received | shares					1,755,000
Arcaea [Member] | License
Business Acquisition [Line Items]
Common units | shares					9,000,000
Allonnia [Member]
Business Acquisition [Line Items]
Consideration											$ 18,500,000
Number of common units received | shares											3,600,000
Number of additional common units received | shares										1,867,411
Carrying value of the equity method investment										$ 0		24,500,000
Loss on equity method investments										(12,700,000)		(24,500,000)
Allocated Upfront Non Cash Consideration										2,500,000
Additional Non Cash Consideration										$ 12,700,000
Number Of Allocated Performance Obligations | Obligations										10
Deferred Revenue										$ 38,000,000.0	$ 26,100,000
Deferred Revenue, Revenue Recognized										5,100,000	5,000,000.0	$ 0
Fixed non-cash consideration										2,500,000
Deferred Revenue, Balance										38,000,000.0	$ 26,100,000
Series A Preferred Stock
Business Acquisition [Line Items]
Share Sold | shares					1,755,000
Consideration					$ 19,500,000					$ 57,100,000
Additional preferred units issued | shares					7,245,000	5,400,000
Number of additional common units received | shares										1,867,411
Series A Preferred Stock | Arcaea [Member]
Business Acquisition [Line Items]
Business Combination, Issue | shares										5,139,900
Number of additional common units received | shares				5,229,900
Series A Preferred Stock | Allonnia [Member]
Business Acquisition [Line Items]
Share Sold | shares											1,664,911	2,970,000
Consideration						$ 33,000,000.0				$ 200,000
Additional preferred units issued | shares						630,000
Business Combination, Issue | shares										22,500
Share issued In Exchange For The Rights To Certain Intellectual Property | shares											180,000
Additional preferred units issued | shares						5,400,000					1,844,911	5,400,000
Motif FoodWorks, Inc.
Business Acquisition [Line Items]
Additional loss recognized										$ 0	$ 0	$ 0
Equity method investments fair value													$ 65,100,000
Motif FoodWorks, Inc. | Variable Interest Entity, Not Primary Beneficiary | Technical Development Agreement [Member]
Business Acquisition [Line Items]
Deferred Revenue										52,200,000	54,000,000.0
Deferred Revenue, Revenue Recognized										20,200,000	20,800,000	19,000,000.0
Investment owned, balance, shares | shares													9,000,900
Equity Method Investments															$ 65,100,000
Impairment losses on equity method investments													$ 65,100,000
Equity method investments fair value										0
Deferred Revenue, Balance										52,200,000	54,000,000.0
Motif FoodWorks, Inc. | Variable Interest Entity, Not Primary Beneficiary | Technical Development Agreement [Member] | Material Rights Tranche One [Member]
Business Acquisition [Line Items]
Number Of Allocated Performance Obligations | Obligations													10
Revenue remaining performance obligation amount													$ 6,500,000
Motif FoodWorks, Inc. | Series A Preferred Stock
Business Acquisition [Line Items]
Share Sold | shares													8,100,720
Consideration													$ 90,000,000.0
Genomatica [Member] | Variable Interest Entity, Not Primary Beneficiary | Two Thousand And Eighteen Technical Development Agreement [Member]
Business Acquisition [Line Items]
Revenue remaining performance obligation amount																$ 40,000,000.0
Investment owned, at cost													$ 40,000,000.0
Genomatica [Member] | Variable Interest Entity, Not Primary Beneficiary | Foundry Terms Of Service Agreement [Member]
Business Acquisition [Line Items]
Deferred Revenue										17,100,000	30,100,000
Deferred Revenue, Revenue Recognized										12,900,000	9,400,000	$ 6,200,000
Upfront amount received										8,300,000	6,900,000
Deferred Revenue, Balance										17,100,000	30,100,000
Genomatica [Member] | Variable Interest Entity, Not Primary Beneficiary | Foundry Terms Of Service Agreement [Member] | Preferred Stock [Member]
Business Acquisition [Line Items]
Equity securities without readily determinable fair value										$ 55,000,000.0	$ 55,000,000.0
Cooksonia
Business Acquisition [Line Items]
Equity method investment, underlying equity in net assets								8,100,000
Cooksonia | Parent
Business Acquisition [Line Items]
Equity Method Investments								13,100,000
Payment to acquire equity method investments								5,000,000.0
Cooksonia | Joyn Bio, LLC [Member]
Business Acquisition [Line Items]
Equity metohd investment ownership percentage											50.00%
Cooksonia | Capital Unit, Class B [Member]
Business Acquisition [Line Items]
Payment to acquire equity method investments								5,000,000.0
Cooksonia | Capital Unit, Class B [Member] | Related Party Investor [Member]
Business Acquisition [Line Items]
Payment to acquire equity method investments									$ 20,000,000.0
Equity metohd investment ownership percentage									20.00%
Cooksonia | Capital Units Class C [Member] | Joyn Bio, LLC [Member]
Business Acquisition [Line Items]
Equity metohd investment ownership percentage									50.00%
Cooksonia | Variable Interest Entity, Primary Beneficiary [Member] | Capital Unit, Class A [Member]
Business Acquisition [Line Items]
Payment to acquire equity method investments									$ 5,000,000.0
Variable interest entity ownership percentage									70.00%
Cooksonia | Noncontrolling Interest [Member]
Business Acquisition [Line Items]
Equity Method Investments								29,700,000
Equity method investment, underlying equity in net assets								$ 8,100,000

[1] Equity method investments in Platform Ventures with a carrying value of zero as of December 31, 2021 and 2020 were excluded from the table above.