License Agreements - Additional Information (Details) - USD ($) $ in Thousands	1 Months Ended			3 Months Ended
	Jun. 30, 2018	Sep. 30, 2016	Mar. 31, 2016	Mar. 31, 2022
Stanford License Agreements [Member] | Stanford [Member]
License Agreements [Line Items]
Payments for milestone agreement or earned royalties on achievement of milestones			$ 5,000
Payments for achievement of specified development and regulatory milestones	$ 400
Payments for milestone agreement on achievement of milestones				$ 0
Stanford License Agreements [Member] | Stanford [Member] | Maximum
License Agreements [Line Items]
Payments for milestone agreement or earned royalties on achievement of milestones			$ 900
UCSF License And Option Agreements [Member] | Stanford [Member]
License Agreements [Line Items]
Payments for milestone agreement on achievement of milestones				0
Distributed Bio Subscription Agreement [Member]
License Agreements [Line Items]
Payments for achievement of specified development and regulatory milestones		$ 5,900
Payments for milestone agreement on achievement of milestones				$ 0