License and Collaboration Agreement - Contract Liability (Details) - Lilly License and Collaboration Agreement - USD ($) $ in Thousands	3 Months Ended		9 Months Ended
	Sep. 30, 2022	Sep. 30, 2021	Sep. 30, 2022	Sep. 30, 2021
License and Collaboration Agreement
Deferred revenues at beginning of period	$ 16,323	$ 26,229	$ 22,367	$ 31,777
Revenues recognized during the period	(4,252)	(1,919)	(10,296)	(7,467)
Deferred revenues at end of period	$ 12,071	$ 24,310	$ 12,071	$ 24,310