License and Collaboration Agreement - Contract Liability (Details) - Lilly License and Collaboration Agreement - USD ($) $ in Thousands	3 Months Ended		6 Months Ended
	Jun. 30, 2022	Jun. 30, 2021	Jun. 30, 2022	Jun. 30, 2021
License and Collaboration Agreement
Deferred revenues at beginning of period	$ 19,206	$ 28,859	$ 22,367	$ 31,777
Revenues recognized during the period	(2,883)	(2,630)	(6,044)	(5,548)
Deferred revenues at end of period	$ 16,323	$ 26,229	$ 16,323	$ 26,229