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Stock-based Compensation (Tables)
 12 Months Ended
Dec. 31, 2022
Share-Based Payment Arrangement [Abstract]  
Schedule of Stock-based Compensation Expense by Classification
Stock-based compensation expense by classification included within the Consolidated Statements of Operations and Comprehensive Loss was as follows (in thousands):
Year Ended December 31,
202220212020
Research and development
$16,721 $15,328 $14,977 
General and administrative
65,203 46,873 18,284 
Total stock-based compensation expense
$81,924 $62,201 $33,261 
Schedule of Incremental Stock-Based Compensation Expense from Modifications
The following table shows the total incremental stock-based compensation expense associated with modifications by the year in which the modification occurred in the years December 31, 2022, 2021 and 2020 (in thousands):
December 31,
202220212020
Previous CEO - Options(1)
$3,741 $— $— 
Previous CEO - Options(2)
— 21,948 15,052 
Other - Options(3)
— 1,019 4,717 
Previous CEO - RSA(4)
— 10,908 20,799 
Other - RSA(5)
— — 9,029 
Total
$3,741 $33,875 $49,597 
__________
(1)The modification for Previous CEO - Options for 2022 is for Ms. Elizabeth Homans, who resigned in December 2022.
(2)The modifications for Previous CEO - Options for 2021 and 2020 is for Dr. Richard Klausner, who resigned as Executive Chairman in October 2021 and is now the Chair of Lyell’s Board.
(3)The modifications for Other - Options are represented by one individual in 2021 and four individuals in 2020.
(4)The modifications for Previous CEO - RSAs for 2021 and 2020 are for Dr. Richard Klausner, who resigned as Executive Chairman in October 2021 and is now the Chair of Lyell’s Board.
(5)The modifications for Other - RSAs are represented by three individuals in 2020.
Schedule of RSA Activity
A summary of the Company’s RSA activity was as follows:
Number of SharesWeighted-Average
Value at Grant Date
Per Share
Unvested shares as of December 31, 2021
2,600,002 $0.0001 
Vested
(2,600,002)$0.0001 
Unvested shares as of December 31, 2022
— $— 
A summary of the Company’s RSU activity was as follows:
Number of SharesWeighted-Average
Value at Grant Date
Per Share
Unvested RSUs as of December 31, 2021
0$— 
RSUs granted1,330,962 $5.98 
RSUs vested(223,200)$5.98 
RSUs forfeited or canceled(235,685)$5.98 
Unvested RSUs as of December 31, 2022
872,077 $5.98 
Schedule of Stock Option Activity
A summary of the Company’s stock option activity was as follows:
Number of
Stock
Options
Weighted-
Average
Exercise
Price
Per Share
Weighted-
Average
Remaining
Contractual
Life
(in years)		Aggregate
Intrinsic
Value (in thousands)
Options outstanding as of December 31, 2021
41,775,179 $5.05 7.84$142,076 
Granted
18,399,530 $4.89 
Exercised
(3,600,478)$2.66 
Canceled or forfeited
(2,725,186)$6.28 
Options outstanding as of December 31, 2022
53,849,045 $5.09 7.84$24,887 
Options exercisable as of December 31, 2022
28,115,416 $4.43 6.78$23,462 
Schedule of Assumptions Used in Black-Scholes Option-Pricing Model for Estimating Fair Value of Stock Options Granted
The fair value of stock options granted to employees, directors and consultants was estimated on the date of grant using the Black-Scholes option pricing model using the following weighted-average assumptions:
Year Ended December 31,
202220212020
Risk-free interest rate2.93 %0.80 %0.79 %
Expected volatility87.3 %78.7 %75.0 %
Expected term (in years)6.036.106.11
Expected dividend yield0 %0 %0 %