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Accrued Liabilities and Other Long-Term Liabilities
 3 Months Ended
Mar. 31, 2022
Accrued Liabilities and Other Liabilities [Abstract]  
Accrued Liabilities and Other Long-Term Liabilities Accrued Liabilities and Other Long-Term Liabilities
Accrued Liabilities
Accrued liabilities consisted of the following (in thousands):
As of
March 31,
2022
As of
December 31,
2021
Accrued compensation and benefits$2,634 $5,216 
Accrued research and development expenses7,846 5,868 
Other accrued liabilities1,146 907 
Total accrued liabilities$11,626 $11,991 
Accrued compensation and benefits consist primarily of accrued bonuses and accrued vacation.