License, Research and Collaboration Agreements - Additional Information (Details) - USD ($) $ in Thousands	3 Months Ended
	Mar. 31, 2022	Mar. 31, 2021
ABL Bio Agreements Member
Collaborative Arrangements And Noncollaborative Arrangement Transactions [Line Items]
Upfront payment received	$ 5,000
Milestone payments	6,000
ABL Bio Agreements Member | Oncology Member
Collaborative Arrangements And Noncollaborative Arrangement Transactions [Line Items]
Research and development milestone payments	110,000
Milestone payment and royalties received	295,000
ABL Bio Agreements Member | Ophthalmology Member
Collaborative Arrangements And Noncollaborative Arrangement Transactions [Line Items]
Milestone payment and royalties received	$ 185,000
Adimab Agreement
Collaborative Arrangements And Noncollaborative Arrangement Transactions [Line Items]
Collaboration agreement date	Oct. 16, 2014
Milestone payments	$ 0
Future milestone payments	2,000
FUJIFILM Diosynth Biotechnologies Agreement
Collaborative Arrangements And Noncollaborative Arrangement Transactions [Line Items]
Future milestone payments	1,200
Prepayments for future work	1,200
Total expense to be recognized	2,400
Cash payment for license agreement	46	$ 200
Research and development	$ 555	$ 1,000