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License and Collaboration Agreements - Summary of Revenues (Details) - Gilead and Taiho - USD ($)
$ in Thousands
3 Months Ended
Mar. 31, 2021
Mar. 31, 2020
Collaboration    
License And Collaboration Agreements [Line Items]    
License revenue $ 9,461 $ 1,750
Collaboration and License    
License And Collaboration Agreements [Line Items]    
License revenue $ 9,461 $ 1,750