License and Collaboration Agreements - Summary of Revenues by Collaboration and by Category of Revenue (Details) - USD ($) $ in Thousands	3 Months Ended		6 Months Ended
	Jun. 30, 2022	Jun. 30, 2021	Jun. 30, 2022	Jun. 30, 2021
License And Collaboration Agreements [Line Items]
Total collaboration and license revenues	$ 26,759	$ 9,461	$ 44,764	$ 18,922
License and Development Services for all Gilead Programs
License And Collaboration Agreements [Line Items]
Total collaboration and license revenues	16,693		24,632
Gilead Access Rights
License And Collaboration Agreements [Line Items]
Total collaboration and license revenues	8,316	7,711	16,632	15,422
Taiho Access Rights
License And Collaboration Agreements [Line Items]
Total collaboration and license revenues	$ 1,750	$ 1,750	$ 3,500	$ 3,500