License and Collaboration Agreements - Summary of Revenues (Details) - Gilead and Taiho - USD ($) $ in Thousands	3 Months Ended		6 Months Ended
	Jun. 30, 2022	Jun. 30, 2021	Jun. 30, 2022	Jun. 30, 2021
License and Development Services Revenue
License And Collaboration Agreements [Line Items]
Total collaboration and license revenues	$ 16,693		$ 24,632
Other Collaboration Revenue
License And Collaboration Agreements [Line Items]
Total collaboration and license revenues	10,066	$ 9,461	20,132	$ 18,922
Collaboration and License
License And Collaboration Agreements [Line Items]
Total collaboration and license revenues	$ 26,759	$ 9,461	$ 44,764	$ 18,922