License and Collaboration Agreements - Summary of Revenues (Details) - Gilead and Taiho - USD ($) $ in Thousands	3 Months Ended
	Mar. 31, 2022	Mar. 31, 2021
Other Collaboration Revenue
License And Collaboration Agreements [Line Items]
Total collaboration and license revenues	$ 10,066	$ 9,461
License and Development Services Revenue
License And Collaboration Agreements [Line Items]
Total collaboration and license revenues	7,939
Collaboration and License
License And Collaboration Agreements [Line Items]
Total collaboration and license revenues	$ 18,005	$ 9,461