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Stock-Based Awards (Tables)
 6 Months Ended
Jun. 30, 2023
Share-Based Payment Arrangement [Abstract]  
Summary of Stock Option Activity
The following table summarizes the Company’s stock option activity since December 31, 2022:
 
 Number of
Shares		Weighted
Average
Exercise
Price		Weighted
Average
Remaining
Contractual
Term		Aggregate
Intrinsic
Value
   (in years) 
Outstanding as of December 31, 20228,233,330 $5.74 6.68$159 
Granted3,068,900 0.69 
Exercised(13,401)0.35 
Forfeited(1,189,831)2.65 
Outstanding as of June 30, 202310,098,998 $4.58 6.95$604 
Options exercisable at June 30, 20236,305,828 $5.78 5.66$188 
Vested and expected to vest at June 30, 202310,098,998 $4.58 6.95$604 
The following table summarizes the Company’s stock option under the Inducement Plan activity since December 31, 2022:

Number of
Shares		Weighted
Average
Exercise
Price		Weighted
Average
Remaining
Contractual
Term		Aggregate
Intrinsic
Value
(in years)
Outstanding as of December 31, 2022210,400 $2.61 9.36$— 
Granted— — 
Exercised— — 
Forfeited(9,113)3.12 
Outstanding as of June 30, 2023201,287 $2.59 8.87$— 
Options exercisable at June 30, 202341,457 $3.26 8.72$— 
Vested and expected to vest at June 30, 2023201,287 $2.59 8.87$— 
Summary of Restricted Stock Unit Activity
The table below summarizes the Company’s RSU activity since December 31, 2022:
 
 Number of
Shares		Weighted
Average
Grant-Date
Fair Value
Unvested restricted stock units as of December 31, 2022385,980 $3.64 
Granted— — 
Vested— — 
Forfeited(107,100)3.64 
Unvested restricted stock units as of June 30, 2023278,880 $3.64 
Summary of Stock-Based Compensation Expense Related to Stock Options and Restricted Stock Awards
The Company recorded stock-based compensation expense related to stock options, ESPP and restricted stock unit awards in the following expense categories of its condensed consolidated statements of operations and comprehensive loss:
 
 Three months ended June 30,Six months ended June 30,
 2023202220232022
Research and development expenses$371 $777 $1,017 $1,360 
General and administrative expenses815 1,296 1,815 2,578 
 $1,186 $2,073 $2,832 $3,938