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Stock-Based Awards (Tables)
 3 Months Ended
Mar. 31, 2023
Share-Based Payment Arrangement [Abstract]  
Summary of Stock Option Activity
The following table summarizes the Company’s stock option activity since December 31, 2022:
 
 Number of
Shares		Weighted
Average
Exercise
Price		Weighted
Average
Remaining
Contractual
Term		Aggregate
Intrinsic
Value
   (in years) 
Outstanding as of December 31, 20228,233,330 $5.74 6.68$159 
Granted3,058,400 0.69 
Exercised— — 
Forfeited(100,599)4.76 
Outstanding as of March 31, 202311,191,131 $4.37 7.40$145 
Options exercisable at March 31, 20235,926,959 $5.90 5.74$118 
Vested and expected to vest at March 31, 202311,191,131 $4.37 7.40$145 
The following table summarizes the Company’s stock option under the Inducement Plan activity since December 31, 2022:

Number of
Shares		Weighted
Average
Exercise
Price		Weighted
Average
Remaining
Contractual
Term		Aggregate
Intrinsic
Value
(in years)
Outstanding as of December 31, 2022210,400 $2.61 9.36$— 
Granted— — 
Exercised— — 
Forfeited— — 
Outstanding as of March 31, 2023210,400 $2.61 9.11$— 
Options exercisable at March 31, 202326,541 $3.64 8.92$— 
Vested and expected to vest at March 31, 2023210,400 $2.61 9.11$— 
Summary of Restricted Stock Unit Activity
The table below summarizes the Company’s RSU activity since December 31, 2022:
 
 Number of
Shares		Weighted
Average
Grant-Date
Fair Value
Unvested restricted stock units as of December 31, 2022385,980 $3.64 
Granted— — 
Vested— — 
Forfeited— — 
Unvested restricted stock units as of March 31, 2023385,980 $3.64 
Summary of Stock-Based Compensation Expense Related to Stock Options and Restricted Stock Awards
The Company recorded stock-based compensation expense related to stock options, ESPP and restricted stock unit awards in the following expense categories of its condensed consolidated statements of operations and comprehensive income (loss):
 
 Three months ended March 31,
 20232022
Research and development expenses$646 $584 
General and administrative expenses1,000 1,281 
 $1,646 $1,865