Collaboration and License Agreements - Schedule of Collaboration Revenue (Details) - USD ($) $ in Thousands	3 Months Ended		6 Months Ended
	Jun. 30, 2022	Jun. 30, 2021	Jun. 30, 2022	Jun. 30, 2021
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]
License-related revenue	$ 0	$ 515	$ 30,000	$ 2,141
GSK Agreement | License
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]
License-related revenue	$ 0	$ 515	$ 30,000	$ 2,141