Selected Quarterly Financial Data (Unaudited)

12 Months Ended

Dec. 31, 2019

Quarterly Financial Information Disclosure [Abstract]

Selected Quarterly Financial Data (Unaudited)

20. Selected Quarterly Financial Data (Unaudited)

The following table contains quarterly financial information for 2019 and 2018. The Company believes that the following information reflects all normal recurring adjustments necessary for a fair presentation of the information for the periods presented. The operating results for any quarter are not necessarily indicative of results for any future period.

(In thousands, except share and per share amounts)

	2019
	First Quarter			Second Quarter			Third Quarter			Fourth Quarter
Collaboration revenue - related party	$	14,434		$	143		$	344		$	439
Total operating expenses		19,402			18,653			17,900			16,771
Loss from operations		(4,968	)		(18,510	)		(17,556	)		(16,332	)
Net loss		(4,199	)		(17,758	)		(16,878	)		(15,954	)
Net loss per share attributable to common    stockholders—basic and diluted	$	(0.15	)	$	(0.64	)	$	(0.61	)	$	(0.57	)
	2018
	First Quarter			Second Quarter			Third Quarter			Fourth Quarter
Collaboration revenue - related party	$	45,495		$	2,428		$	1,730		$	9,764
Total operating expenses		14,452			19,011			19,760			15,345
Loss from operations		31,043			(16,583	)		(18,030	)		(5,581	)
Net loss		31,212			(15,852	)		(17,222	)		(4,735	)
Net loss per share attributable to common    stockholders—basic	$	1.59		$	(0.73	)	$	(0.62	)	$	(0.17	)
Net loss per share attributable to common    stockholders— diluted	$	1.05		$	(0.73	)	$	(0.62	)	$	(0.17	)