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Marketable Securities
3 Months Ended
Mar. 31, 2024
Investments, Debt and Equity Securities [Abstract]  
Marketable Securities Marketable Securities
All marketable securities were considered available-for-sale at March 31, 2024 and December 31, 2023. On a recurring basis, the Company records its marketable securities at fair value using Level 1 or Level 2 inputs as discussed in Note 2, "Fair Value Measurements". The amortized cost, gross unrealized holding gains or losses, and fair value of the Company’s marketable securities by major security type at each balance sheet date are summarized in the tables below (in thousands):
March 31, 2024
Amortized CostUnrealized Holding GainsUnrealized Holding LossesAggregate Fair Value
Short-term marketable securities:
U.S. government treasuries(1)
$829,902 $51 $(544)$829,409 
Corporate debt securities
1,079 — — 1,079 
Commercial paper
45,807 — — 45,807 
Total short-term marketable securities
876,788 51 (544)876,295 
Long-term marketable securities:
U.S. government treasuries(2)
475,612 — (416)475,196 
Corporate debt securities(3)
15,562 — (35)15,527 
Total long-term marketable securities
491,174 — (451)490,723 
Total
$1,367,962 $51 $(995)$1,367,018 
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(1)Unrealized holding losses on 23 securities with an aggregate fair value of $572.9 million.
(2)Unrealized holding losses on 14 securities with an aggregate fair value of $475.2 million.
(3)Unrealized holding losses on 4 securities with an aggregate fair value of $15.5 million.

December 31, 2023
Amortized CostUnrealized Holding GainsUnrealized Holding LossesAggregate Fair Value
Short-term marketable securities:
U.S. government treasuries
$868,174 $998 $— $869,172 
U.S. government agency securities(1)
7,089 — (3)7,086 
Commercial paper
31,147 — — 31,147 
Total
$906,410 $998 $(3)$907,405 
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(1)Unrealized holding losses on 2 securities with an aggregate fair value of $7.1 million.
As of March 31, 2024 and December 31, 2023, some of the Company's marketable securities were in an unrealized loss position. The Company has not recognized an allowance for credit losses as of March 31, 2024 or December 31, 2023. The Company determined that it had the ability and intent to hold all marketable securities that have been in a continuous loss position until maturity or recovery. Further, a majority of these marketable securities are held in U.S. government securities, and the remainder were initially, and continue to be, held with investment grade, high credit quality institutions. All marketable securities with unrealized losses as of each balance sheet date have been in a loss position for less than twelve months or the loss is not material.

As of March 31, 2024, all of the Company’s marketable securities have an effective maturity of less than two years.