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Cash and Marketable Securities
3 Months Ended
Mar. 31, 2020
Cash and Cash Equivalents [Abstract]  
Cash and Marketable Securities Cash and Marketable Securities
Cash, cash equivalents and restricted cash

A reconciliation of cash, cash equivalents, and restricted cash reported within the Condensed Consolidated Balance Sheets to the amount reported within the Condensed Consolidated Statements of Cash Flows is shown in the table below (in thousands):
March 31,December 31,March 31,December 31,
2020201920192018
Cash and cash equivalents
$207,537  $79,449  $39,661  $77,123  
Restricted cash included within other non-current assets
1,500  1,500  1,500  1,500  
Total cash, cash equivalents, and restricted cash
$209,037  $80,949  $41,161  $78,623  
Marketable securities
All marketable securities were considered available-for-sale at March 31, 2020 and December 31, 2019. On a recurring basis, the Company records its marketable securities at fair value using Level 1 or Level 2 inputs as discussed in Note 2, "Fair Value Measurements". The amortized cost, gross unrealized holding gains or losses, and fair value of the Company’s marketable securities by major security type at each balance sheet date are summarized in the tables below (in thousands):
March 31, 2020
Amortized CostUnrealized Holding GainsUnrealized Holding LossesAggregate Fair Value
Short-term marketable securities:
U.S. government treasuries
$272,289  $1,391  $—  $273,680  
Corporate debt securities(1)
61,920  39  (193) 61,766  
Commercial paper
44,267  —  —  44,267  
Total short-term marketable securities
378,476  1,430  (193) 379,713  
Long-term marketable securities:
U.S. government treasuries
2,525  40  —  2,565  
Corporate debt securities(2)
7,153  —  (34) 7,119  
Total long-term marketable securities
9,678  40  (34) 9,684  
Total
$388,154  $1,470  $(227) $389,397  
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(1)Unrealized holding losses on 10 securities, with an aggregate fair value of $35.7 million.
(2)Unrealized holding losses on 1 security, with an aggregate fair value of $7.1 million.
December 31, 2019
Amortized CostUnrealized Holding GainsUnrealized Holding LossesAggregate Fair Value
Short-term marketable securities:
U.S. government treasuries(1)
$249,478  $594  $(2) $250,070  
U.S. government agency securities
1,999   —  2,000  
Corporate debt securities(2)
53,396  94  (11) 53,479  
Commercial paper
30,358  —  —  30,358  
Total short-term marketable securities
335,231  689  (13) 335,907  
Long-term marketable securities:
U.S. government treasuries(3)
13,865   (2) 13,869  
Corporate debt securities(4)
25,998  34  (15) 26,017  
Total long-term marketable securities
39,863  40  (17) 39,886  
Total
$375,094  $729  $(30) $375,793  
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(1)Unrealized holding losses on 2 securities, with an aggregate fair value of $9.3 million.
(2)Unrealized holding losses on 4 securities, with an aggregate fair value of $25.9 million.
(3)Unrealized holding losses on 1 security, with an aggregate fair value of $10.1 million.
(4)Unrealized holding losses on 3 securities, with an aggregate fair value of $17.2 million.
As of March 31, 2020 and December 31, 2019, some of the Company’s marketable securities were in an unrealized loss position. The Company has not recognized an allowance for credit losses as of March 31, 2020 or December 31, 2019. The Company determined that it had the ability and intent to hold all marketable securities that have been in a continuous loss position until maturity or recovery. Further, these marketable securities were initially, and continue to be, held with investment grade, high credit quality institutions, and the unrealized losses are believed to be the result of the recent widespread market and interest rate impact of the COVID-19 pandemic. All marketable securities with unrealized losses as of each balance sheet date have been in a loss position for less than twelve months or the loss is not material.
The Company recorded unrealized gains on marketable securities as other comprehensive income in the three months ended March 31, 2020. As a result, the Company recorded a tax benefit of $0.1 million for the three months ended March 31, 2020 on the Condensed Consolidated Statements of Operations and Comprehensive Loss and a corresponding tax charge in other comprehensive income. During the three months ended March 31, 2019, no tax benefit was recorded on unrealized gains recorded in other comprehensive income.

All of the Company’s marketable securities have an effective maturity of less than two years.