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Selected Quarterly Financial Data (Unaudited)
 12 Months Ended
Dec. 31, 2019
Quarterly Financial Information Disclosure [Abstract]  
Selected Quarterly Financial Data (Unaudited) Selected Quarterly Financial Data (Unaudited)
The following tables provide the selected quarterly financial data for the years ended December 31, 2019 and 2018 (in thousands, except per share amounts):
Quarter Ended
December 31, 2019September 30, 2019June 30, 2019March 31, 2019
Collaboration revenue$4,672  $13,604  $4,197  $4,205  
Loss from operations(57,758) (50,155) (62,763) (42,508) 
Net loss(54,025) (46,260) (58,337) (38,992) 
Net loss per share, basic and diluted(1)
$(0.56) $(0.48) $(0.61) $(0.41) 
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(1) Basic and diluted net income per share for the quarter ended December 31, 2019 is calculated using 96,078,950 shares.

Quarter Ended
December 31, 2018September 30, 2018June 30, 2018March 31, 2018
Collaboration revenue$125,676  $1,195  $1,648  $641  
Income (loss) from operations74,722  (37,964) (57,382) (25,748) 
Net income (loss)77,533  (35,371) (54,724) (23,678) 
Net income (loss) per share, basic(1)
$0.82  $(0.38) $(0.59) $(0.26) 
Net income (loss) per share, diluted(1)
$0.79  $(0.38) $(0.59) $(0.26) 
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(1) Basic and diluted net income per share for the quarter ended December 31, 2018 is calculated using 94,299,096 and 97,746,224 shares, respectively, the latter of which is computed by giving effect to all dilutive potential ordinary shares including options.