CONDENSED CONSOLIDATED STATEMENTS OF OPERATIONS AND COMPREHENSIVE LOSS (Unaudited) - USD ($) $ in Thousands	3 Months Ended
	Mar. 31, 2022	Mar. 31, 2021
Revenues:
Grant revenue	$ 1,822	$ 7,300
Collaboration revenue	$ 247
Type of Revenue [Extensible List]	us-gaap:LicenseMember	us-gaap:LicenseMember
Total revenues	$ 2,069	$ 7,300
Operating expenses:
Research and development	16,971	18,404
General and administrative	15,305	8,299
Total operating expenses	32,276	26,703
Loss from operations	(30,207)	(19,403)
Other income (expense):
Interest income	72	98
Other income (expense), net	(13)	(118)
Interest expense related to the sale of future royalties	(2,488)
Change in fair value of derivative liability	(193)
Total other income (expense), net	(2,622)	(20)
Net loss	$ (32,829)	$ (19,423)
Net loss per share attributable to common stockholders, basic and diluted	$ (1.01)	$ (0.66)
Weighted average common shares outstanding, basic and diluted:	32,606,715	29,414,148
Comprehensive loss:
Net loss	$ (32,829)	$ (19,423)
Other comprehensive gain (loss):
Unrealized gain (loss) on marketable securities	(4)	4
Net unrealized gains (losses) on securities	(4)	4
Total comprehensive loss	$ (32,833)	$ (19,419)