Stock-Based Compensation (Tables)

3 Months Ended

Mar. 31, 2020

Share-based Payment Arrangement [Abstract]

Summary of Stock-based Compensation Expense

Stock-based compensation expense included in the Company’s condensed consolidated statements of operations and comprehensive loss is as follows (in thousands):

	Three Months Ended March 31,
	2020		2019
General and administrative	$	889	$	1,062
Research and development	1,066		891
Total stock-based compensation expense	$	1,955	$	1,953

Summary of Stock Option Activity

A summary of the Company’s stock option activity and related information is as follows: 

	Shares	Weighted- Average Exercise Price
Options outstanding at December 31, 2019	5,691,474	$	6.99
Granted	1,358,500	7.02
Exercised	(137,213)	1.65
Canceled	(560,730)	9.27
Options outstanding at March 31, 2020	6,352,031	$	6.91
Exercisable as of March 31, 2020	2,790,620	$	5.26