Revenue Recognition - Summary of Revenue Disaggregated by Research Program (Details) - USD ($) $ in Thousands	3 Months Ended		6 Months Ended
	Jun. 30, 2022	Jun. 30, 2021	Jun. 30, 2022	Jun. 30, 2021
Disaggregation Of Revenue [Line Items]
Total collaboration revenue	$ 5,851	$ 8,756	$ 17,210	$ 16,003
MAT2A Program
Disaggregation Of Revenue [Line Items]
Total collaboration revenue	3,740	1,763	11,599	3,230
Pol Theta Program
Disaggregation Of Revenue [Line Items]
Total collaboration revenue	1,224	5,074	3,344	8,609
WRN Program
Disaggregation Of Revenue [Line Items]
Total collaboration revenue	$ 887	$ 1,919	$ 2,267	$ 4,164