The Company is party as a lessee to various non-cancelable operating leases with varying terms through March 2028 primarily for laboratory and office space and equipment. The Company has options to renew some of these leases after their expirations. On a lease-by-lease basis, the Company considers such options, which may be elected at the Company’s sole discretion, in determining the lease term. The Company also has three finance leases for lab equipment through December 2026, one of which was acquired in a recent business combination. The Company retained acquirees’ classification of its leases. The Company does not have any leases with variable lease payments. The Company’s operating lease agreements do not contain any residual value guarantees, material restrictive covenants, bargain purchase options or asset retirement obligations.

The Company’s headquarters are located in Temple City, California, which is comprised of various corporate offices and a laboratory certified under the Clinical Laboratory Improvement Amendments of 1988, or CLIA, accredited by the College of American Pathologists, or CAP, and licensed by the State of California Department of Public Health. Other CLIA-certified laboratories are located in Houston, Texas; Alpharetta, Georgia; Phoenix, Arizona; New York, New York; Irving, Texas; and Needham, Massachusetts. Additional offices are located in Atlanta, Georgia and are used for certain report generation functions.

The operating and finance lease right-of-use asset, short-term lease liabilities, and long-term lease liabilities as of June 30, 2022 and December 31, 2021 were as follows:


	June 30,		December 31,
	2022		2021
	(in thousands)
Operating lease ROU asset, net	$	18,034	$	7,141
Operating lease liabilities, short term	$	6,216	$	1,842
Operating lease liabilities, long term	$	11,892	$	5,344
Finance lease ROU asset, net	$	2,671	$	1,735
Finance lease liabilities, short term	$	741	$	332
Finance lease liabilities, long term	$	1,938	$	1,398

The following was operating and finance lease expense:


	Three months ended June 30,				Six months ended June 30,
	2022		2021		2022		2021
	(in thousands)
Operating lease cost	$	1,366	$	232	$	1,904	$	301
Finance lease cost:
Amortization of ROU assets		169		—		265		—
Interest on lease liabilities		21		—		35		—
Short-term lease cost		834		61		931		192
Total lease cost	$	2,390	$	293	$	3,135	$	493

Supplemental information related to operating leases and finance lease was the following:


	June 30, 2022
Weighted average remaining lease term - operating leases	3.54 years
Weighted average discount rate - operating leases		3.81	%
Weighted average remaining lease term -finance lease	3.71 years
Weighted average discount rate - finance lease		3.61	%

The following is a maturity analysis of operating and finance lease liabilities using undiscounted cash flows on an annual basis with renewal periods included:


	Operating Leases			Finance Lease
	(in thousands)
Year Ending December 31,
2022 (remaining 6 months)	$	3,451		$	412
2023		6,613			825
2024		4,093			818
2025		2,124			440
2026		1,523			366
2027		1,360			—
Thereafter		217			—
Total lease payments		19,381			2,861
Less imputed interest		(1,273	)		(182	)
Total	$	18,108		$	2,679

The Company leases out space in buildings it owns and leases to third-party tenants under noncancelable operating leases. As of June 30, 2022, the remaining lease terms left range from 6 months to 38 months, including renewal options and may include rent escalation clauses. Lease income primarily represents fixed lease payments from tenants recognized on a straight-line basis over the application lease term. Variable lease income represents tenant payments for real estate taxes, insurance and maintenance.

The lease income was included in interest and other income, net, in the accompanying Condensed Consolidated Statements of Income. Total lease income was as follows:


	Three months ended June 30,				Six months ended June 30,
	2022		2021		2022		2021
	(in thousands)
Lease income	$	84	$	81	$	181	$	237
Variable lease income		11		4		12		5
Total lease income	$	95	$	85	$	193	$	242

Future fixed lease payments from tenants for all noncancelable operating leases as of June 30, 2022 are as follows:


	Lease Payments
	from Tenants
	(in thousands)
Year Ending December 31,
2022 (remaining 6 months)	$	164
2023		264
2024		180
2025		120
Total	$	728