Collaboration and License Agreements - Schedule of Financial Information Related to Collaboration and License Agreements (Details) - USD ($) $ in Thousands	Mar. 31, 2022	Dec. 31, 2021
Collaborative Arrangements And Noncollaborative Arrangement Transactions [Line Items]
Accounts Receivable	$ 1,773	$ 5,716
Deferred Revenue, Current	31,266	31,800
Deferred Revenue, Net of Current	19,020	24,368
Deferred Revenue, Total	50,286	56,168
Roche Agreement
Collaborative Arrangements And Noncollaborative Arrangement Transactions [Line Items]
Accounts Receivable	500	1,215
Deferred Revenue, Current	6,021	5,601
Deferred Revenue, Net of Current	16,173	17,215
Deferred Revenue, Total	22,194	22,816
Biogen Agreement
Collaborative Arrangements And Noncollaborative Arrangement Transactions [Line Items]
Accounts Receivable		3,000
Deferred Revenue, Current	23,078	24,032
Deferred Revenue, Net of Current	2,847	6,611
Deferred Revenue, Total	25,925	30,643
Calico Agreement
Collaborative Arrangements And Noncollaborative Arrangement Transactions [Line Items]
Accounts Receivable	1,273	1,501
Deferred Revenue, Current	2,167	2,167
Deferred Revenue, Net of Current		542
Deferred Revenue, Total	$ 2,167	$ 2,709