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Basic and Diluted Net Loss Per Share (Tables)
 6 Months Ended
Jun. 30, 2018
Basic and Diluted Net Loss Per Share [Abstract]  
Basic Loss Per Share
The following table summarizes the distributable losses for the three and six months ended June 30, 2018 and 2017 (in thousands):

  
Three Months Ended
June 30,
  
Six Months Ended
June 30,
 
 
 
2018
  
2017
  
2018
  
2017
 
Net loss
 
$
(62,046
)
 
$
(20,359
)
 
$
(91,673
)
 
$
(84,214
)
Distribution to Ionis
  
(7,792
)
  
—
   
(7,792
)
  
—
 
Distributable losses
 
$
(69,838
)
 
$
(20,359
)
 
$
(99,465
)
 
$
(84,214
)

The following table summarizes the reconciliation of weighted-average shares outstanding used in the calculation of basic loss per share for the three and six months ended June 30, 2018 and 2017:

  
Three Months Ended
June 30,
  
Six Months Ended
June 30,
 
Determination of shares:
 
2018
  
2017
  
2018
  
2017
 
Weighted-average preferred shares outstanding
  
—
   
28,884,540
   
—
   
28,884,540
 
Weighted-average common shares outstanding owned by Ionis
  
60,832,494
   
—
   
53,182,685
   
—
 
Weighted-average common shares outstanding owned by others
  
21,492,157
   
—
   
21,332,650
   
—
 
Total weighted-average shares outstanding
  
82,324,651
   
28,884,540
   
74,515,335
   
28,884,540
 

The following table summarizes the calculation of basic loss per share for the three and six months ended June 30, 2018 and 2017 (in thousands, except per share amounts):
 
  
Three Months Ended
June 30,
  
Six Months Ended
June 30,
 
 
 
2018
  
2017
  
2018
  
2017
 
Losses allocated to preferred shares
 
$
—
  
$
(20,359
)
 
$
—
  
$
(84,214
)
Weighted-average preferred shares outstanding
  
—
   
28,884,540
   
—
   
28,884,540
 
Basic loss per preferred share
 
$
—
  
$
(0.70
)
 
$
—
  
$
(2.92
)
                 
Losses allocated to Ionis
 
$
(51,606
)
 
$
—
  
$
(70,990
)
 
$
—
 
Plus: Distribution to Ionis
  
7,792
   
—
   
7,792
   
—
 
Losses available to Ionis
 
$
(43,814
)
  
—
   
(63,198
)
  
—
 
Weighted-average common shares outstanding owned by Ionis
  
60,832,494
   
—
   
53,182,685
   
—
 
Basic loss per common share owned by Ionis
 
$
(0.72
)
 
$
—
  
$
(1.19
)
 
$
—
 
                 
Losses allocated to common shares owned by others
 
$
(18,232
)
 
$
—
  
$
(28,475
)
 
$
—
 
Weighted-average common shares outstanding owned by others
  
21,492,157
   
—
   
21,332,650
   
—
 
Basic loss per common share owned by others
 
$
(0.85
)
 
$
—
  
$
(1.33
)
 
$
—