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Stock-Based Compensation (Tables)
6 Months Ended
Jun. 30, 2018
Stock-Based Compensation [Abstract]  
Stock-Based Compensation Expense
The following table summarizes stock-based compensation expense for the three and six months ended June 30, 2018 and 2017 (in thousands):

 
 
Three Months Ended
June 30,
  
Six Months Ended
June 30,
 
 
 
2018
  
2017
  
2018
  
2017
 
Research and development expenses
 
$
2,242
  
$
1,919
  
$
4,555
  
$
3,520
 
General and administrative expenses
  
9,884
   
2,023
   
13,954
   
3,602
 
Total
 
$
12,126
  
$
3,942
  
$
18,509
  
$
7,122