Stockholders' Equity - Additional Information (Details) - USD ($)		1 Months Ended	6 Months Ended
	Aug. 04, 2023	Aug. 31, 2021	Jun. 30, 2024	Jun. 30, 2023	Dec. 31, 2023	Jul. 31, 2020
Class Of Stock [Line Items]
Common stock, shares authorized			250,000,000		250,000,000
Common stock, par value			$ 0.0001		$ 0.0001
Common stock, voting rights			Each share of common stock is entitled to one vote.
Dividends declared			$ 0
Proceeds from issuance of common stock through ATM offering, net of issuance costs				$ 928,000
Common stock issued and sold			97,080,817		95,636,553
Common Stock
Class Of Stock [Line Items]
Warrants to purchase stock			93,518
Warrants to purchase stock			$ 4.28
Common Stock | Astellas | Securities Purchase Agreement
Class Of Stock [Line Items]
Discount for lack of marketability of shares, at fair value on date of issuance	$ 14,400,000
Common Stock | 2018 and 2019 Warrants
Class Of Stock [Line Items]
Warrants to purchase stock			27,604
Warrants to purchase stock			$ 7.25
IPO
Class Of Stock [Line Items]
Common stock, shares authorized						250,000,000
Common stock, par value						$ 0.0001
Undesignated preferred stock, shares authorized						10,000,000
Undesignated preferred stock, par value						$ 0.0001
At the Market Facility | Common Stock | Sales Agreement | Cantor Fitzgerald & Co.
Class Of Stock [Line Items]
Proceeds from issuance of common stock through ATM offering, net of issuance costs				$ 900,000
Aggregate offering price		$ 85,000,000
Weighted average price per share				$ 8.04
Common stock issued and sold			0	119,000
Private Placement | Common Stock | Astellas | Securities Purchase Agreement
Class Of Stock [Line Items]
Common stock, par value	$ 0.0001
Sale of common stock under an underwritten public offering	8,333,333
Sale of stock, price per share	$ 3
Proceeds from issuance of common stock through ATM offering, net of issuance costs	$ 25,000,000