CONDENSED CONSOLIDATED STATEMENTS OF OPERATIONS (Unaudited) - USD ($) $ in Thousands	3 Months Ended
	Mar. 31, 2022	Mar. 31, 2021
Revenue:
Collaboration revenue	$ 12,704	$ 13,191
Total revenue	12,704	13,191
Operating expenses:
Research and development	14,248	16,550
General and administrative	6,707	6,588
Total operating expenses	20,955	23,138
Loss from operations	(8,251)	(9,947)
Other income (expense):
Other income	819	332
Interest income	4	5
Interest expense	(600)	(698)
Total other income (expense), net	223	(361)
Net loss	$ (8,028)	$ (10,308)
Net loss per share attributable to common stockholders, basic and diluted	$ (0.36)	$ (0.51)
Weighted average common shares outstanding, basic and diluted	22,436,938	20,333,398