Marketable Securities and Fair Value Measurements - Schedule of Financial Assets Measured at Fair Value on a Recurring Basis (Details) - USD ($) $ in Thousands	Dec. 31, 2021	Dec. 31, 2020
Marketable securities:
Total Marketable securities	$ 240,521	$ 425,408
Total	288,321	502,084
U.S. government securities
Marketable securities:
Total Marketable securities	13,084	307,758
Corporate debt securities
Marketable securities:
Total Marketable securities	98,525	38,275
Commercial paper
Marketable securities:
Total Marketable securities	105,390	62,031
Certificates of deposit
Marketable securities:
Total Marketable securities	23,522	17,344
Money market funds
Cash equivalents:
Total Cash equivalents	47,800	46,676
U.S. government securities
Cash equivalents:
Total Cash equivalents	0
Certificates of deposit
Cash equivalents:
Total Cash equivalents		30,000
Level 1
Marketable securities:
Total	0	0
Level 1 | U.S. government securities
Marketable securities:
Total Marketable securities	0	0
Level 1 | Corporate debt securities
Marketable securities:
Total Marketable securities	0	0
Level 1 | Commercial paper
Marketable securities:
Total Marketable securities	0	0
Level 1 | Certificates of deposit
Marketable securities:
Total Marketable securities	0	0
Level 1 | Money market funds
Cash equivalents:
Total Cash equivalents	0	0
Level 1 | U.S. government securities
Cash equivalents:
Total Cash equivalents	0
Level 1 | Certificates of deposit
Cash equivalents:
Total Cash equivalents		0
Level 2
Marketable securities:
Total	288,321	502,084
Level 2 | U.S. government securities
Marketable securities:
Total Marketable securities	13,084	307,758
Level 2 | Corporate debt securities
Marketable securities:
Total Marketable securities	98,525	38,275
Level 2 | Commercial paper
Marketable securities:
Total Marketable securities	105,390	62,031
Level 2 | Certificates of deposit
Marketable securities:
Total Marketable securities	23,522	17,344
Level 2 | Money market funds
Cash equivalents:
Total Cash equivalents	47,800	46,676
Level 2 | U.S. government securities
Cash equivalents:
Total Cash equivalents	0
Level 2 | Certificates of deposit
Cash equivalents:
Total Cash equivalents		30,000
Level 3
Marketable securities:
Total	0	0
Level 3 | U.S. government securities
Marketable securities:
Total Marketable securities	0	0
Level 3 | Corporate debt securities
Marketable securities:
Total Marketable securities	0	0
Level 3 | Commercial paper
Marketable securities:
Total Marketable securities	0	0
Level 3 | Certificates of deposit
Marketable securities:
Total Marketable securities	0	0
Level 3 | Money market funds
Cash equivalents:
Total Cash equivalents	0	0
Level 3 | U.S. government securities
Cash equivalents:
Total Cash equivalents	$ 0
Level 3 | Certificates of deposit
Cash equivalents:
Total Cash equivalents		$ 0