X0306 4 2018-05-24 0 0001497253 ORGANOVO HOLDINGS, INC. ONVO 0001651405 Gallant Paul 6275 NANCY RIDGE DRIVE, SUITE 110 SAN DIEGO CA 92121 0 1 0 0 General Manager Common Stock 2018-05-24 4 A 0 66115 0.00 A 239804 D Stock Option (Right to Buy) 1.84 2018-05-24 4 A 0 266667 0.00 A 2028-05-24 Common Stock 266667 266667 D Annual long-term equity award approved by the Compensation Committee of the Issuer's Board of Directors. The shares represent a grant of Restricted Stock Units ("RSUs"). The RSUs represent a right to receive shares of the Issuer's common stock, at no additional conversion or exercise price. The RSUs vest and settle over 16 equal quarterly installments measured from May 15, 2018. This number does not include 586,667 shares of common stock underlying outstanding options held by Mr. Gallant; post-transaction Mr. Gallant beneficially owns or has a right to acquire 826,471 shares of common stock. The option shares vest and become exercisable over 16 equal quarterly installments measured from May 15, 2018. /s/ Paul Gallant 2018-05-29