Condensed Consolidated Statements of Operations and Comprehensive Loss - USD ($) $ in Thousands	3 Months Ended		6 Months Ended
	Jun. 30, 2022	Jun. 30, 2021	Jun. 30, 2022	Jun. 30, 2021
Condensed Consolidated Statements of Operations and Comprehensive Loss
Collaboration revenue	$ 712	$ 1,357	$ 1,371	$ 7,858
Operating expenses:
Research and development	12,527	19,505	26,876	41,851
General and administrative	7,552	10,437	15,211	20,181
Total operating expenses	20,079	29,942	42,087	62,032
Operating loss	(19,367)	(28,585)	(40,716)	(54,174)
Other income (expense) :
Interest income	219	48	271	132
Other (expense) income	61	(1,583)	39	2,273
Total other income (expense), net	280	(1,535)	310	2,405
Net loss	(19,087)	(30,120)	(40,406)	(51,769)
Other comprehensive (loss) income
Net unrealized (loss) gain on available-for-sale securities	(141)	(3)	(226)	8
Total other comprehensive (loss) income	(141)	(3)	(226)	8
Comprehensive loss	$ (19,228)	$ (30,123)	$ (40,632)	$ (51,761)
Net loss per share, basic	$ (0.50)	$ (0.80)	$ (1.06)	$ (1.38)
Net loss per share, diluted	$ (0.50)	$ (0.80)	$ (1.06)	$ (1.38)
Weighted-average common shares outstanding, basic	38,298,426	37,581,381	38,183,192	37,543,387
Weighted-average common shares outstanding, diluted	38,298,426	37,581,381	38,183,192	37,543,387