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Stock-based compensation (Tables)
6 Months Ended
Jun. 30, 2022
Stock-based compensation  
Compensation cost recognized for all stock-based compensation awards

Total compensation cost recognized for all stock-based compensation awards in the condensed consolidated statements of operations and comprehensive loss is as follows:

Three Months Ended

Six Months Ended

June 30, 

 

June 30, 

 

    

2022

    

2021

 

2022

    

2021

 

(in thousands)

 

Research and development

$

781

$

1,336

$

1,582

$

2,706

General and administrative

 

1,709

 

2,655

 

3,227

 

4,886

Total stock-based compensation expense

$

2,490

$

3,991

$

4,809

$

7,592

Summary of stock-based compensation expense by type of award

Three Months Ended

Six Months Ended

June 30, 

June 30, 

    

2022

    

2021

2022

    

2021

(in thousands)

Stock options

$

1,618

$

2,480

$

3,095

$

5,002

Restricted stock awards and units

842

1,392

1,634

2,366

Employee stock purchase plan awards

 

30

 

120

81

223

Total stock-based compensation expense

$

2,490

$

3,991

4,809

7,592

Summary of status and changes in unvested restricted stock

A summary of the status of and changes in unvested restricted stock unit activity under the Company’s equity award plans for the six months ended June 30, 2022 was as follows:

    

    

Weighted

Average

Grant Date

Fair Value

    

Units

    

Per Unit

Unvested restricted stock units as of December 31, 2021

 

806,379

$

7.26

Granted

 

739,080

$

3.79

Vested

 

(344,255)

7.45

Forfeited

 

(169,839)

$

5.06

Unvested restricted stock units as of June 30, 2022

 

1,031,365

$

5.08

Summary of stock option activity

The following is a summary of stock option activity for the six months ended June 30, 2022:

    

    

Weighted

    

Remaining

    

Aggregate

Average

Contractual

Intrinsic

Exercise

Life

Value

    

Shares

    

Price

    

(in years)

    

(in thousands)

Outstanding at December 31, 2021

 

5,013,193

$

12.69

5.6

$

1,165

Granted

 

2,965,075

$

4.95

Exercised

 

(74,496)

$

7.90

Cancelled or forfeited

 

(1,274,172)

$

12.42

Outstanding at June 30, 2022

 

6,629,600

$

9.33

 

7.2

$

5,914

Exercisable at June 30, 2022

 

3,378,945

$

12.29

 

5.3

$

1,575