Condensed Consolidated Balance Sheets - USD ($) $ in Thousands	Jun. 30, 2022	Dec. 31, 2021
Current assets:
Cash and cash equivalents	$ 78,329	$ 117,433
Marketable securities, current	69,727	15,106
Related party collaboration receivable	151	732
Prepaid expenses and other current assets	3,421	3,427
Total current assets	151,628	136,698
Property and equipment, net	19,071	21,920
Deposits and other non-current assets	1,515	1,779
Operating lease, right-of-use asset	16,391	33,458
Total assets	188,605	193,855
Current liabilities:
Accounts payable	1,218	574
Accrued expenses	7,185	10,950
Other current liabilities	2,526	5,571
Deferred revenue, current	87,003	33,886
Total current liabilities	97,932	50,981
Deferred revenue, non-current	7,880	8,210
Other non-current liabilities	22,741	39,609
Total liabilities	128,553	98,800
Commitments and contingencies (see note 8)
Stockholders' equity:
Preferred stock $0.001 par value: 5,000,000 shares authorized at June 30, 2022 and December 31, 2021; no shares issued and outstanding at June 30, 2022 and December 31, 2021
Common stock, $0.001 par value: 120,000,000 shares authorized at June 30, 2022 and December 31, 2021; 38,439,251 and 37,918,395 shares issued and outstanding at June 30, 2022 and December 31, 2021, respectively	38	38
Additional paid-in capital	447,888	442,259
Accumulated other comprehensive loss	(364)	(138)
Accumulated deficit	(387,510)	(347,104)
Total stockholders' equity	60,052	95,055
Total liabilities and stockholders' equity	$ 188,605	$ 193,855