Condensed Consolidated Statements of Operations and Comprehensive Loss - USD ($) $ in Thousands	3 Months Ended
	Mar. 31, 2021	Mar. 31, 2020
Condensed Consolidated Statements of Operations and Comprehensive Loss
Collaboration revenue	$ 6,501	$ 18,067
Operating expenses:
Research and development	22,346	32,294
General and administrative	9,744	10,206
Total operating expenses	32,090	42,500
Operating loss	(25,589)	(24,433)
Other income:
Interest income	19	978
Other income (expense)	3,921	(808)
Total other income	3,940	170
Net loss	(21,649)	(24,263)
Other comprehensive income
Net unrealized gain on available-for-sale-securities	11	525
Total other comprehensive income	11	525
Comprehensive loss	$ (21,638)	$ (23,738)
Net loss per share, basic and diluted	$ (0.58)	$ (0.66)
Weighted-average common shares outstanding, basic and diluted	37,501,065	36,963,255